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http://www.plosbiology.org/article/info%3Adoi%2F10.1371%2Fjournal.pbio.1001114http://www.plosbiology.org/article/info%3Adoi%2F10.1371%2Fjournal.pbio.1001114http://www.sciencedirect.com/science/article/pii/S002219101000171Xhttp://dx.doi.org/10.1371/journal.pbio.1001115