atlas of trichoptera of the sw pacific — australian region ||

ATLAS OF TRICHOPTERA OF THE SW PACIFIC - AUSTRALIAN REGION

SERIES ENTOMOLOGICA

EDITOR

K.A. SPENCER

VOLUME 37

Atlas of Trichoptera of the

SW Pacific - Australian Region

By

Arturs Neboiss

1986 DR W. JUNK PUBLISHERS ~. a member of the KLUWER ACADEMIC PUBLISHERS GROUP II. DORDRECHT / BOSTON / LANCASTER .'liliiii

Distributors

jor the United States and Canada: Kluwer Academic Publishers, 190 Old Derby Street, Hingham, MA 02043, USA jor the UK and Ireland: Kluwer Academic Publishers, MTP Press Limited, Falcon House, Queen Square, Lancaster LAI lRN, UK jor all other countries: Kluwer Academic Publishers Group, Distribution Center, P.O. Box 322, 3300 AH Dordrecht, The Netherlands

Library of Congress Cataloging in Publication Data

Neboiss, Arturs. Atlas of Trichoptera of the SW Pacific-Australian

region.

(Series entomologica ; vol. 37) Bibliography: p. Includes indexes. I. Caddis-flies--Oceania. 2. Caddis-flies-

Australasia. 3. Insects--Oceania. 4. Insects-Australasia. I. Title. II. Series: Series entomologica ; v. 37. QL517.8.eGN42 1986 595.7'45'099 85-23755

ISBN-13 :978-94-01 0-8634-9 DOl: 10.1007/978-94-009-4814-3

Copyright

e-ISBN -13 : 978-94-009-4814-3

© 1986 by Dr W. Junk Publishers, Dordrecht. Softcover reprint ofthe hardcover 1st edition 1986

All rights reserved. No part of this publication may be reproduced, stored in a retrieval system, or transmitted in any form or by any means, mechanical, photocopying, recording, or otherwise, without the prior written permission of the publishers, Dr W. Junk Publishers, P.O. Box 163, 3300 AD Dordrecht, The Netherlands.

v

CONTENTS

Introduction VII

Acknowledgments VIII

Explanations and abbreviations 2

Caddis-fly structures 3

Distribution - abbreviations used 4

Map of the SW Pacific - Australian region 5

Key to families 6

Superfamily Rhyacophiloidea

Family Hydrobiosidae 10

Glossosomatidae 54

Hydroptilidae 59

Superfamily Hydropsychoidea

Family Philopotamidae

Stenopsychidae

Hydropsychidae

Polycentropodidae

Ecnomidae

Psychomyiidae

Superfamily Limnephiloidea

Family Chathamiidae

Tasimiidae

Limnephilidae

Goeridae

Lepidostomatidae

Oeconesidae

Kokiriidae

Plectrotarsidae

Conoesucidae

Antipodoeciidae

Calocidae

Helicophidae

Odontoceridae

Atriplectididae

Philorheithridae

Helicopsychidae

Calamoceratidae

Leptoceridae

Literature cited

Index of generic names

Index of species names

97

llO

ll4

137

149

157

159

161

164

166

168

170

176

180

183

195

197

205

209

2ll

213

219

223

230

274

279

280

VI

Macrostemum wallacei (McLachlan, 1866) ~

McLachlan 1866 [44] pI. XVII, fig.4. IDe. New Guinea, coil. A.R.Wallace

INTRODUCTION

The caddis-fly species included in this atlas are those either described or recorded from the SW Pacific - Australian region. For the purposes of this work the SW Pacific - Australian region is defined as the area within the boundaries of Weber's line, north to the Equator,east to the International dateline,south to the small islands off the South Island of New Zealand and west to coastal Western Australia (see Map page 5). The latgest landmass of the region is the Australian continent.

The history of caddis-fly studies in the SW Pacific region begins with the description of the Australian species Plectrotarsus gravenhorsti by Kolenati in 1848. Four years later, several New Zealand and Australian species were described by Francis Walker (1852) from the British Museum collection. The earliest New Guinea descriptions date from 1863, when McLachlan described three Anisocentropus species. These were collected by Mr Allen, a companion of Alfred Russel Wallace during his travels through the Malay Archipelago. The first Fijian caddis-flies (three species) were described by Brauer (1867) from the Godeffroy Museum material.

Compared to other groups of insects, particularly butterflies and beetles, caddis-flies were rarely collected and their numbers in collections were proportionally very low. As a result, knowledge of this region's fauna has increased slowly. In 1894 a total of 22 species were known from New Zealand, 10 from Australia and only 8 from New Guinea.

At the start of this century, three separate expeditions added considerable amounts of new material. These were the Hamburg - South West Australia expedition, Dr Mjoberg's Swedish Scientific expedition to North Queensland and the German Kaiserin-AugustaFluss expedition to the present day Sepik River area of New Guinea. The material was examined and new species were described by Ulmer (1908, 1915, 1916).

New material from other sources was investigated and described by several authors during a period of two decades,Banks (1913, 1920), Tillyard (1921,1922, 1924,1925), Navas (1923-1933); however, the number of known species still remained low. In the following years additional collections were made by travelling naturalists and the specimens were deposited in various European and American Museums where they remained for many yealS. These collections included Fijian material assembled by W.M.Mann in 1915, described by Banks (1924, 1936), Tasmanian specimens assembled by Tonnoir 1922-23,described by Jacquemart 1965, and material from New Guinea collected by Miss Cheesman 1933-38, described by Kimmins (1962).

Preliminary studies of the region's fauna were published by Mosely (1924-1951), Banks (1939) and McFarlane (1939, 1951); however, it was not until 1953 (Mosely and Kimmins) that a comprehensive study of the Australian and New Zealand fauna was compiled. This brought together information of the fauna from both countries, described numerous new species and discussed family arrangements. Unfortunately, it did not include information on the fauna of New Guinea or the SW Pacific Islands.

The last three decades have seen a growing concern and awareness of freshwater quality and pollution problems. Surveys established to deal with this field have provided detailed information on aquatic

VII

environments using insects as a biological measure of water quality. Such surveys have amassed considerable amounts of material and initiated extensive taxonomic work to enable comparative ecological studies.

The family classification of Trichoptera was analysed by Ross (1956, 1967), Riek (1968, 1977),Schmid (1970) and Neboiss (1977). In New Zealand, McFarlane continued studies of the local fauna and described numerous species in a series of papers (1956 - 1981) which were further added to by Leader (1972), Cowley (1976) and Henderson (1983).

Studies of the Australian fauna identified numerous new species and this has enabled the reevaluation of generic and family attributes as detailed by Neboiss(1962-1984 )and Ivells(1979-1984). The SW Pacific island fauna still remains largely unexplored. Some recently collected material has been described by Malicky (1978, 1981), Kumanski (1979) and Neboiss (1984) .

The above historical resume clearly demonstrates the paucity of information concerning the Trichoptera of the SW Pacific region. This is further compounded by the scattered depositories of early described material making comparative studies difficult.

Given the archipelago nature of the SW PacificAustralian region, it is possible to divide it into five areas, each with a high degree of endemism and its own specific composition of species. Of particular interest is the separation of Tasmania from the Australian mainland as a distinct area.

Australia 335 species Tasmania 165 species New Zealand 160 species New Guinea 152 species Pacific islands 48 species

Twenty seven families are recognised for the above species, and their arrangement follows that used by Wiggins (1982). The genera and species are grouped on visual similarities, not always adhering to accepted taxonomic arrangements.

A key with some basic illustrations is provided as a general guide to the families. It should be used with discretion due to atypical forms,incomplete information and unknown characters of the opposite sex. Each family is preceded by a brief diagnosis of its characters and synopsis of its genera. Each generic name is followed by a number indicating the number of species included. Throughout the atlas, species distribution is indicated in an abbreviated form except when restricted to individual islands (name given in full) or for extra-limital records.

For each species a set of illustrations is given which follows a basic format. Leading from left to right are views of male genitalia (lateral, dorsal and ventral) and, where available, views of female genitalia. The above format may be incomplete if illustrations are not available. Sources of the illustrations are listed in the species index. An index of generic and species names (including synonymies) has also been compiled.

This atlas is the first attempt to bring together information on all species described or recorded from this large region and is envisaged as a precursor to future studies.

ACKNOWLEDGMENTS

I am indebted to many people who offered invaluable assistance and co-operation in preparation of this atlas.

My sincere thanks are due to the colleagues in New Zealand, Alex McFarlane, Keith Wise and Ian Henderson, who sent their original drawings and generously supplied information on the New Zealand caddis-flies. Illustrations of almost all species in the family Hydroptilidae have been prepared from the original negatives made available by Mrs Alice Wells of Adelaide University. John Morse of Clemson University, South Carolina, enthusiastically supported the idea at the early stage of development and provided original illustrations for the Tripleccides group of species.

Permission to reproduce illustrations from their papers, and some originals were received from Ferdinand Schmid, Ottawa, Hans Malicky, Lunz, and Edgar Riek, Canberra. New illustrations have been prepared from a number of type specimens received on loan from P.C.Barnard, British Museum (Natural History), London; R.Poggi, Museo Civico di Storia Naturale, Giacomo Doria, Genova; F.Newton, Museum of Comparative Zoology, Harvard University, Cambridge , Massachusetts, J.D.Unzicker, Illinois Natural History Survey, Urbana, Illinois and Miss Margaret Schneider, Department of Entomology, University of Queensland.

Valuable comments and advice was received from L.Botosaneanu, Zoological Museum, Amsterdam, T. New, La Trobe University, Bondoora, J. Dean and D. Cartwright, Melbourne and Metropolitan Board of Works, Melbourne. Many technical problems were solved in discussions with my colleagues at the Museum of Victoria, particularly K. Walker and G. Poore. I thank also my wife Austra for her constant encouragement and for the many sacrifices she has made during the preparation of this volume.

Through the generosity of the editors and publishers I was able to use many published illustrations. Those reproduced are individually acknowledged in the species index, showing their bibliography reference and figure number. I am grateful to the following:

the Managing Editor of the Australian Journal of Zoology, Melbourne;

the British Museum (Natural History), Department of Entomology, London, for the numerous illustrations from the articles by ;Zimmins in the Bulletin of the

VIII

British Museum, Natural History (Entomology) 1957-1980, and from 'The Trichoptera (Caddis-flies) of Australia and New Zealand' by Mosely and Kimmins (1953);

the Editor of the 'Pacific Insects', B.P. Bishop Museum, Entomology Department, Honolulu;

the Editor, Bishop Museum Press, for the permission to reproduce three figures from the article by Ross (1951) in the Occasional Papers No 20;

Swets Publishing Service, Lisse, for theillustrations from the article by Kumanski in the Aquatic Insec ts (1979);

the Director, Canterbury Museum, illustrations by McFarlane in the Records of the Canterbury Museum, Christchurch;

the Editor of the Records of the Auckland Institute and Museum, Auckland;

the Chief Editor of the Journal of the Australian Entomological Society, Brisbane;

to Taylor and Francis Ltd. London, for figures by Mosely in the Annales and Magazine of Natural History;

the Registrar of the Royal Entomological Society London, for articles by Mosely in the Transactions of the Royal Society of Entomology, and by Leader in the Journal of Entomology, series B;

the Secretary of the Schweizerbart'sche Verlagsbuchhandlung, Stuttgart, for the extracts from Ulmer's publication in the Archiv fur Hydrobiologie (1951);

the figures from Ross' 'Evolution and classification of the mountain caddisflies' (1965) have been reproduced by the permission of the Board of Trustees of the University of Illinois and the publisher, University of Illinois Press, Urbana;

the Managing Editor, Entomological Society of Canada, Ottawa, figures by Schmid in the Canadian Entomologist;

the Editor, Science information publishing centre, DSIR, Wellington, figures by Cowley and Henderson, in the New Zealand Journal of Zoology;

the Editor, Journal of the Royal Society of New Zealand, Wellington, for the figures by Mosely, Tillyard and ~cFarlane;

the Director, Museum d'Histoire naturelle,Geneve, for the illustrations by Malicky in Revue Suisse de Zoologie;

the General Director, Rijksmusea van Geologie & Mineralogie en Natuurlijke Historie, Leiden, figures by Ulmer in the Notes from the Leiden Museum;

the Council of the Royal Society of South Australia, Adelaide for the illustrations in the article by Wells published in the Society's Transactions.

To my wife Austra

~--te9

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Trichoptera structural ch aracters

3

EXPLANATIONS

Adult structures - abbreviations Wing structures

an antenna sct scutellum A anal vein ats apical tibial spur sg abdominal segment arc arculus cl tarsal claw sn tibial and tarsal spines C costa co coxite st sternite Cu cubitus cx coxa sw setal wart dc discoidal cell e compound eye sua superior appendage jl jugal lobe fe femur te tergite M media ha harpago teg tegula mc median cell ina inferior appendage ti tibia ny nygma la lateral lobe of segment 9 tr tarsus R radius lbp labial palp tro trochanter Rs radial sector If lateral filament. sternite 5 vag vaginal structure Sc subcosta ms meso scutum vp ventral process tc thyridial cell mt meta thorax Other abbreviations Cross-veins mxp maxillary palp c-sc costa-subcosta oc ocellus D dorsal cu-a cubitus-analis par paramere L lateral hc humeral ph phallus or phallic complex V ventral m-cu media-cubitus pr pronotum r radius pts preapical tibial spur ~ character to aid r-m radius-media

identification

HYDROBIOSIDAE: ULMEROCHOREMA 1:4:4+

stigma (ULMER. 1916)G) m: 7 -9 mm; f: 7.5 - 9.5 mm AUne

AC

AN

AU

AU c

AU n

AU ne

AU nw

AU se

AU sw

BO

C

CH

FI

GI

GU

JA

KR

LH

NB

NC

NG

NH

NO

NZ

NZ n

NZ s

SA

SL

SM

SN

SO

ST

TA

4

DISTRIBUTION - Abbreviations used

Auckland Islands 50°40 S 166°33 E (not on the map)

Antipodes Islands 49°40 S 178°47 E (not on the map)

Australia collectively (excluding Tasmania)

Australia central

Australia northern

Australia north-eastern, Cape York Penninsula

Australia north-western, Kimberleys

Australia south-eastern

Australia south-western

Borneo

Campbell Island 52°30 S 169°05 E (not on the map)

Chatham Islands 44°00 S 176°30 W

Fiji Islands 18°00 S 178°00 E

Gilolo Island now Halmahera

Guadalcanal Island 9°32 S 160°12 E

Java

Kermadec Islands 29°16 S 177°55 W

Lord Howe Island 31°33 S 159°05 E

New Britain 6°00 S 150°00 E

New Caledonia 21°30 S 165°30 E

New Guinea collectively

New Hebrides now Vanuatu 16°00 S 167°00 E

Norfolk Island 29°02 S ]67°57 E

New Zealand collectively

New Zealand north Island

New Zealand south Island

Sarawak (not on the map)

Sulawesi (Celebes)

Sumatra (not on the map)

The Snares Islands 48°07 S 166°38 E (not on the map)

Solomon Islands collectively

Stewart Island 47°00 S 167°50 E

Tasmania

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6

KEY TO FAMILIES

Insects small,usually 5 mm or less in length; an

tennae always shorter than forewing; maxillary

palpi 5-segmented in both sexes;scutellum flat,

triangular, pointed posteriorly, margins verti

cal (Fig.2); mesoscutum without setal warts;

wings narrow, slender, pointed apically (Fig.1>,

fringes very long, those of hindwing usually

longer than width of the wing ••• HYDROPTILIDAE

Insects usually more than 5 mm long; antennae

variable in length, in some genera two or three

times the length of forewing; maxillary palpi

1 - 6 segmented in males, 5 or 6-segmented in

females; scutellum broad posteriorly, convex,

without vertical margins (Fig.3);mesoscutum fre

quently with setal warts; wings proportionally

broad, fringes usually short, less than the width

of the wing ••••••••••••••••••••••••••••••••• 2

2 Ocelli present (Fig.4) •••••••••••••••••••••••• 3

Ocelli absent (Fig.5) ••••••••••••••••••••••••• ?

3 Maxillary palpi with segment 5 flexible, distinct

ly longer than segment 4 (Fig.6) ••••••••••••••

PHILOPOTAMIDAE

Maxillary palpi 3 or 5-segmented, apical segment

not flexible or annulate,approximately the same

length or shorter than preceding segment •••• 4

4 Maxillary palpi 5-segmented in both sexes, seg

ments 1 and 2 short, stout; without setal warts

on mesoscutum ••••••••••••••••••••••••••••••• 5

Maxillary palpi 3-segmented in male, 5-segmented

in female; segment 1 short, cylindrical; seg

ment 2 usually longer than segment 1, slender;

setal warts present on mesoscutum ••••••••••• 6

5 Segment 2 of maxillary palpi with globular, me

sally directed, lateral projection (Fig.?) ••••

••••••••••••••••••••••••••••••• GLOSSOSOMATIDAE

Segment 2 of maxillary palpi cylindrical, without

lateral projection (Fig.B) •••••• HYDROBIOSIDAE

Fig. 2 Fig. 3

Fig. 4 Fig.5

Fig.6

Fig. 8

7

6 Mesoscutum with one pair of elongate setal warts

(Austral ian species only) (Fig.9) • LIMNEPHILIDAE

Mesoscutum with two pairs of indistinct warts, or

with two groups of setate punctures, the poste

rior pair very small, lateral to scutellum

(Fig.10) •••••••••••••••••••••• PLECTROTARSIDAE

7 Maxillary palpi S-segmented in both sexes, apical

segment fLexible, annulate, at least twice as

Long as segment 4 (Fig.11) •••••••••••••••••• 8

MaxiLLary paLpi 1 to S-segmented in males, aLways

S-segmented in femaLes; the S-segmented palpi

with Last segment short, firm, sometimes the

Last, or the two Last segments with granuLar

texture, but not annuLate (Fig.12) ••••••••• 12

8 Mesoscutum without setaL warts (Fig.13) •••••••••

•••••••••••••••••••••••••••••••• HYDROPSYCHIDAE

Mesoscutum with setaL warts (Fig.16) •••••••••• 9

9 Forewing vein R1 usually forked at apex (sometimes

difficuLt to see) (Fig.14) •••••••••• ECNOMIDAE

Forewing vein R1 not forked at apex •••••••••• 10

10 Mesoscutal warts separated by anterior extention

of scuteLlum (Fig.1S) •••••••••• STENOPSYCHIDAE

Mesoscutal warts not separated by anterior scu

telLar extention, scuteLlum rounded anteriorly

(Fig.16) ••••••••••••••••••••••••••••••••••• 11

11 Spurs 3:4:4; foretibia with preapicaL spur •••••

••••••••••••••••••••••••••••• POLYCENTROPODIDAE

Spurs 2:4:4 or 1:2:4; foretibia without pre

apicaL spur ••••••••••••••••••••• PSYCHOMYIIDAE

12 Midtibia with preapicaL spurs •••••••••••••••• 13

Midtibia without preapicaL spurs ••••••••••••• 21

13 MaxilLary paLpi segment 1 short, expanded apico

mesaLLy and bearing a tuft of setae (Fig.17) ••

•••••••••••••••••••••••••••••• PHILORHEITHRIDAE

Maxillary palpi segment 1 elongate,without apico-

mesaL expansion •••••••••••••••••••••••••••• 14

Fig. 10

Fig. 13

Fig. 15 Fig. 16

8

14 Mesoscutum with one pair of setal warts •••••• 15

Mesoscutum without setal warts, but with two lon

gitudinal bands or groups of setiferous punc-

tures •••••••••••••••••••••••••••••••••••••• 19

15 In forewing and hindwing Rl joins R2 before wing

margin (Fig.18) ••••••••••••••••••• OECONESIDAE

In forewing and hindwing Rl reaches wing margin

separately from R2 ••••••••••••••••••••••••• 16

16 Head with mouth parts slightly to considerably

extended; forewing vein Rl joins Sc basad of

pterostigma by short cross-vein or sharp bend

(Fig.19) KOKIRIIDAE

Head with mouthparts normal, not extended; fore

wing vein Rl without sharp bend, sometimes

joined to Sc by cross-vein ••••••••••••••••• 17

17 Preapical spurs of mid-tibia arising close to mid

point of tibia (Fig.20) •••••• LEPIDOSTOMATIDAE

Preapical spurs of mid-tibia arising well distal

of mid-point.of tibia (Fig.21) ••••••••••••• 18

18 Forewing with discoidal cell closed; maxillary

palpi of male 3-segmented •••••••••••• GOERIDAE

Forewing discoidal cell open; maxillary palpi of

male 4-segmented ••••••••••••••••••• TASIMIIDAE

19 Forewing median cell present, closed; maxillary

palpi 6-segmented in both sexes; spurs 2:4:3

(genus Anisocentropus) •••••••• CALAMOCERATIDAE

Forewing median cell either open or absent;maxil

lary palpi 5-segmented in both sexes; spurs

2:4:4 •••••••••••••••••••••••••••••••••••••• 20

20 Forewing veins Rl and R2 join shortly before wing

margin (Fig.22) ••••••••••••••••• ODONTOCERIDAE

Forewing vein Rl runs separately from R2 to wing

margin (Fig.23) ••••••••••••••• ATRIPLECTIDIDAE

21 Mesoscutum with one pair of setal warts,sometimes

indistinct •••••••••••••••••••••••••••••••.• 22

Mesoscutum without setal warts ••••••••••••••• 23

Fig. 19

Fig. 20 A~ ~ ~---~ ~

Fig. 21 .4 "

Fig. 22

Fig. 23

9

22 Forewing rounded apically, discoidal cell closed;

scutellum with pair of rounded setal warts ••••

(Fig.24) •••••••••••••••••••••• HELICOPSYCHIDAE

Forewing acute apically, discoidal cell open;

scutellum with one, mesal, dome shaped wart

(Fig.25) •••••••••••••••••••••• ANTIPODOECIIDAE

23 Mesoscutum without bands of setate punctures; if

setae present, they are minute and dispersed

over most of mesoscutum •••••••••••••••••••• 24

Mesoscutum with pair of irregular bands of set ate

punctures, extending for part or entire length

of mesoscutum •••••••••••••••••••••••••••••• 26

24 Wi ng coup ling by hamu lion basa l sect i on of costa l

margin of hindwing, distal hamuli more closely

grouped; jugal lobe reduced; hindwing with large

vein-free area discally (Fig.26) • HELICOPHIDAE

Wing coupling by enlarged hairs along costal mar

gin of hindwing, at most only slightly curved,

jugal lobe well developed; hindwing discal area

without vein-free area (Fig.27) •••••••••••• 25

25 Pronotum with two pairs of setal warts, mesal pair

small, rounded, lateral pair elongate (Fig.28)

(except males of Caenota, where only one elongate

pair is present,but then hindwing vein R, joins

Sc for short distance before ending separately

at wing margin), forewing without hyaline spot

at arculus •••••••••••••••••••••••••• CALOCIDAE

Pronotum with pair of elongate setal warts (Fig.

29); forewing usually with hyaline spot at arcu

lus (Fig.3D); hindwing vein R, usually runs se

parately to wing margin, rarely Sc joins R, near

wing margin •••••••••••••••••••••• CONOESUCIDAE

26 Antennae stout, about as long as forewing (longer

in brachypterous species); scutellum with pair

of indistinctly fused warts; maxillary palpi

stout, 5-segmented in both sexes; segment 2

lengthened, prolonged beyond insertion of seg

ment 3 in males (Fig.3') ••••••••• CHATHAMIIDAE

Antennae filiform, slender, 2 to 3 times as long

as forewing; scutellum without warts; maxillary

palpi slender, 5-segmented and similar in both

sexes (Fig.32) ••••••••••••••••••• LEPTOCERIDAE

hindwing

Fig. 24 Fig. 25

Fig. 27

~~~ m) ~

----------- Fig. 29

Fig. 31

Fig. 32

10

HYDROBIOSIDAE

Members of this famiLy are medium sized, wings often

motte Led, dark brown to bLack, in most genera with up

right, thickened hairs; maLes often with smaLL areas of

modified setae; wing span between 10 and 35 mm.

OceLLi present. MaxiLLary paLpi 5-segmented in both

sexes;segments 1 and 2 cyLindricaL,shorter than others,

terminaL segment simpLe;antennae as Long as or sLightLy

shorter than forewing.

Mesoscutum and scuteLLum without setaL warts. Wings

eLongate, ovaL, venation compLete, often differinn in

sexes; forewings with hyaLine Line aLong M and hyaLine

areas around cross-veins r-m, m-cu and at arcuLus; ves

titure in some genera evenLy pubescent, in others with

tufts of thickened upright hairs on veins; discoidaL

ceLL either cLosed or open in forewing, open in hind

wing. LateraL fi Lament to sternite 5 in maLes often pre-

sent; prominent ventromesaL process usuaLLy present

on segments 6, 7 or 8. FemaLe abdomen in some genera

terminates in sLender, rigid ovipositor.

TibiaL spurs 1:4:4 or 2:4:4, maLes of some genera

0:4:4.

The famiLy occurs predominantLy in the AustraLian

and NeotropicaL regions, aLthough a few species are

found in the OrientaL, Nearctic and PaLearctic regions.

UntiL recentLy [99] this group was regarded as a

subfamiLy within the famiLy RhyacophiLidae. There are

about 40 genera with approximateLy 150 species, which

are pLaced in two subfamiLies [77]: ApsiLochoreminae

Neboiss and Hydrobiosinae ULmer. Both subfamiLies, with

a totaL of 26 genera and 138 species, are recorded from

the SW Pacific - AustraLian region.

Synopsis of genera

(showing number of species in each genus; recessed names are synonyms)

APSILOCHOREMINAE

Allochorema MoseLy, 1953

Apsil ochorema U L me r, 1907

Achorema MoseLy, 1941

Bachorema MoseLy, 1953

HYDROBIOSINAE

(2)

(11)

Allobiosis MoseLy, 1953 (1 )

Atrachorema McFarLane, 1964 (2)

Austrochorema MoseLy, 1953 (11)

Costachorema McFarLane, 1939 (7)

Edpercivalia McFarLane, 1964 (9)

Percivalia McFarLane, 1939

Ethochorema Neboiss, 1977 (7)

Hydrobiosis McLachLan, 1868 (19)

Hydrochorema TiLLyard, 1924 (2)

Ipsebiosis Neboiss, 1977 (1)

Koetonga Neboiss, 1962 (1)

Megogata Neboiss, 1962

Moruya Neboiss, 1962

(1)

(3)

Neurochorema TiLLyard, 1924 (4)

Psilochorema McLachLan, 1866 (13)

Psyllobetina Banks, 1939 (5)

Ptychobiosis Neboiss, 1977 (2)

Synchorema TiLLyard, 1924 (2)

Tanjilana Neboiss, 1962 (2)

Tanorus Neboiss, 1984 (4)

Ornatus Korboot, 1964

Taschorema MoseLy, 1936 (9)

NoUobiosis Banks, 1939

Tiphobiosis Ti LLyard, 1924 (9)

Traillochorema Mc Fa r Lane, 1981

Ulmerochorema MoseLy, 1953 (9)

Anachorema MoseLy, 1953

Xanthochorema Kimmins, 1953 (1)

(1)

11

HYDROBIOSIDAE: APSILOCHOREMA 2:4:4+

gisbum (MOSELY, 1953) m: 7 - 8.5 mm; f: 8.5 - 10 mm AU ne,se, TA

ob/iquum (MOSELY, 1953) m: 7 - 8.5 mm; f: 8 - 9.5 mm AU ne,se, TA

~~ ina V

urdalum NEBOISS, 1962 m: 6 - 8 mm; f: 6.5 - 9 mm

rossi KIMMINS, 1957 m:4.5mm GU

12

HYDROBIOSI DAE: APSILOCHOREMA 2:4:4+

oxypages NEBOISS, 1984 m:8mm NG

burgersi ULMER, 1938 m:5.5mm; NG

~o 0 o:::-:f. '-/ . '.

"':,:

mose/yella KIMMINS, 1955 m:6mm SA

zimmermani ROSS, 1951 m: 6.5 mm FI

13

HYDROBIOSIDAE: A PSI LOCHOREMA 2:4:4+

mose/y; ROSS, 1951 m: 5 -7 mm; f: 7 -8 mm FI

- .. ~ ... _.".- - --~ :'~ i

cf

banks; (MOSELY, 1941) m: 6 mm; f: 6 mm FI

cheesmanae KIMMINS, 1958 m: 5.5 mm; f: 6 mm NH

ina V

~

14

HYDROBIOSIDAE: ALLOCHOREMA 2:4:4+

tasmanicum MOSELY, 1953 (j) m: 6 - 7 mm; f: 5.5 - 6.5 mm TA

. "","

reclivatum NEBOISS, 1962 m:6mm;f:6mm AUse

v

HYDROBIOSIDAE: XANTHOCHOREMA 0:4:4+

caledon KIMMINS, 1953 (j) m:9.5mm NC

15

HYDROBIOSIDAE: AUSTROCHOREMA 2:4:4+

pegidion NEBOISS, 1962 m: 6.5 mm - 7.5 mm; f: 7 - 7.5 mm TA

alpinum NEBOISS, 1962 m: 7 - 9 mm; f: 8 - 10 mm AUse

wentum MOSELY, 1953(i) m:6mm;f6mm AUse

16

HYDROBIOSIDAE: AUSTROCHOREMA 2:4:4+

nama NEBOISS,1962 m: 7 - 8 mm; f: 7 - 8 mm AUse

patulum NEBOISS, 1962 m:8mm AUse

concubium NEBOISS, 1962 m:8mm AUse

spinosum NEBOISS, 1962 m:7mm AUse

17

HYDROBIOSIDAE: AUSTROCHOREMA

crinitum NEBOISS, 1977

~",:.

evansi (MOSELY, 1953) m: 5 - 5.5 mm; f: 6 mm TA

-: .1

lepnevae JACQUEMART, 1965 m:5.5mm TA

complexa JACQUEMART, 1965 m:6mm TA

~ ... " .. .' .....

. ' .. . ..

:.,~y:

I 4'

I I

I \ I \ ~

2:4:4+

18

HYDROBIOSIDAE: IPSEBIOSIS 2:4:4+

spicula NEBOISS, 1977(j) m: 8.5 - 9 mm; f: 9 - 9.5 mm TA

D

HYDROBIOSIDAE: KOETONGA 2:4:4+

clivicola NEBOISS, 1962 (f) m: 11 - 12 mm; f: 12 -14 mm AU se, TA

19


stigma (ULMER, 1916) (j') m: 7 - 9 mm; f: 7.5 -9.5 mm AUne

tasmanicum (MOSELY, 1953) m: 6 - 8 mm; f: 6.5 - 8.5 mm TA

membrum NEBOISS, 1962 m:8-10 mm;f: 9-10 mm AU se

HYDROBIOSIDAE:

len tum NEBOISS, 1962

rubiconum NEBOISS, 1962

luxaturum NEBOISS, 1962

20

ULMEROCHOREMA

m: 8 mm; f: 9 - 11 mm AUse, TA

m:7-9mm;f:8-10mm AUse,TA

~ ~

f: 6.5 mm AUse

~~6 // , st~~

v

v " ,

I

I

,(

I I

1:4:4+

,

21


breve (MOSELY, 1953) m: 7 mm;f: 7-7.5 mm TA

v

seona (MOSELY, 1953) m: 5 - 7.5 mm; f: 6 - 8 mm

onychion NEBOISS, 1977 m: 6 -7.5 mm;f: 6.5-8 mm AUse, TA

22

HYDROBIOSIDAE: MEGOGATA 2:4:4+

necopina NEBOISS, 1962 CD m: 7mm;f: 7-7.5 mm AUse

HYDROBIOSIDAE: TANJILANA 2:4:4+

akroreia NEBOISS, 1962(!) m: 9.5 - 10 mm; f: 10 -11 mm AUse

zothecula NEBOISS, 1962 m: 8 - 8.5 mm; f: 8.5 - 9.5 mm AUse

23

HYDROBIOSIDAE: TASCHOREMA 2:4:4+

asmanum MOSELY, 1936 (i) m: 10 -12 mm; f: 11 - 12.5 mm TA

apobamum NEBOISS, 1977 m: 10-12.5mm;f: 11 -14mm TA

ferulum NEBOISS, 1977 m: 10- 12 mm; f: 11 - 13 mm TA

-.... "'-";-

:_,i;-/~- -~::~~: .~:>.

24


pallescens (BANKS, 1939) m: 7 -10.5 mm; f: 8 -10.5 mm AUsw

rugulum NEBOISS, 1962 m:7.5mm

sl6

kimminsi NEBOISS, 1962 m:9.5mm AUse

sl6

25


evansi MOSELY, 1953 m:8-10mm;f:8-11 mm AU ne,se, TA

pedunculatum JACQUEMART, 1965 m: 6 -7 mm TA

viridarium NEBOISS, 1962 m: 6 -7.5 mm; f: 6 - 8 mm TA

26

HYDROBIOSIDAE: ETHOCHOREMA 2:4:4+

nesydr;on (NEBOISS. 1962) (j) m: 8 - 9.5 mm; f: 8 -10 mm TA

~ '\

~~

£'/Il~'\ . , \ " . ., I \ I '_'. ::" I ,

', .... ' .. ~ ~ {: ...... :>.: .. 1\\ .. ,.. . ..... 'Ii. ":

secutum NEBOISS.1977 m:8.S-9mm TA

ochraceum (MOSELY. 1953) m: 8 - 9 mm; f: 9 -10 mm AU ne,se

~ ... ,

\... "-

ph L

27

HYDROBIOSI DAE: ETHOCHOREMA 2:4:4+

turbidum (NEBOISS, 1962) m: 10- 14 mm; f: 11 - 15 mm AUse

D

brunneum (MOSELY, 1953) m: 8 -13.5 mm; f: 9 -13.5 mm AU ne,se

kelion NEBOISS, 1977 m: 8.5 - 10.5 mm; f: 11 mm TA

28

HYDROBIOSIDAE: ETHOCHOREMA 2:4:4+

hesperium (NEBOISS, 1962) m:7.Smm AUse

HYDROBIOSIDAE: PTYCHOBIOSIS 2:4:4+

nigrita (BANKS, 1939) (j) m: 9 -12mm; f: 10-13 mm AU ne,se, TA

rieki (NEBOISS, 1962) m:10-11mm AUse

~ ~::=vp===::~

29

HYDROBIOSIDAE: COSTACHOREMA 2:4:4+

psaroptera McFARLANE, 1939 (i') m: 12 - 13.5 mm; f: 13 - 16 mm NZ

callistum McFARLANE,1939 m: 9.5 - 11 mm; f: 9.5 - 13 mm NZ

notoptera WISE, 1972 m: 10.5 - 11.2 mm; f: 12.5 - 13 mm AC

30

HYDROBIOSIDAE: COSTACHOREMA 2:4:4+

xanthoptera McFARLANE, 1939 m: 11 -13 mm;f: 14-18mm NZ

brachyptera McFARLANE,1939 m: 9 -14 mm; f: 10 -14 mm NZs

hebdomon McFARLANE,1981 m: 15 mm; f: 15 mm

hecton McFARLANE,1981 m: 11 -13.5mm;f: 11-13mm NZ

31

HYDROBIOSIDAE: A TRACHOREMA 2:4:4+

mangu McFARLANE, 1964 (i) m: 6 mm;f: 7 mm NZn

tuarua McFARLANE, 1966 m: 5.5 - 7.5 mm; f: 7 mm NZs

HYDROBIOSI DAE: TRAILLOCHOREMA 2:4:4+

rakiura McFARLANE, 1981 (i) m:9mm ST

32

HYDROBIOSIDAE: HYDROB/OSIS 2:4:4+

frater McLACHLAN, 1868 <!> m: 7 - 8 mm; f: 10- 11 mm NZ

ph

kiddi McFARLANE,1951 m:9-9.5mm NZs

silvico/a McFARLANE,1951 m: 8 - 8.5 mm; f: 8.5 - 10 mm NZs

o

33

HYDROBIOSIDAE: HYDROB/OS/S 2:4:4+

budgei McFARLANE, 1960 m: 10 mm; f: 11 mm NZn

;

.... I::~···· . '" ... ,,-

ph D

johnsi McFARLANE, 1981 m:11-12mm;f:12mm NZs,ST

"'---==--:::'-{--

cha/codes McFARLANE, 1981 m: 12 mm; f: 14 mm NZs

34


umbripennis McLACHLAN,1868 m: 8 -10.5 mm; f: 10 -12.5 mm NZ

\\\II/~/JI

parumbripennis McFARLANE,1951 m:9-12mm;f:11-14.5mm NZ,ST

lindsayi TILL YARD, 1925 m: 9.5 - 10 mm CH

gol/anis MOSELY, 1953 m: 7.5 - 9 mm; f: 8 - 9.5 mm NZ

35


spatulata McFARLANE, 1951 m: 7.5 - 9.5 mm; f: 7.5 - 8 mm NZ

styx McFARLANE,1951 m: 8.5 -10.5 mm NZs

harpidiosa McFARLANE,1951 m:9-10mm;f:12mm NZ

36

HYDROBIOSIDAE: HYDR08/0S/S 2:4:4+

soror MOSELY, 1953 m: 7 - 8 mm; f: 8 - 9 mm NZ

c/avigera McFARLANE,1951 m: 7.5-9 mm; f: 8.5 -10 mm NZ

v

charadraea McFARLANE,1951 m: 9 - 10 mm; f: 9.5 - 11 mm NZ

37

HYDROBIOSI DAE: HYDR08/0S/S

styracine McFARLANE,1960 m: 9 - 11.5 mm; f: 11 - 12.5 mm

/.1 ............ .

/ /" \ , :

" .... ".

'"

copis McFARLANE,1960

fa/cis WISE, 1958

m: 9 - 10 mm; f: 10 - 12 mm NZ

m: 10 -12 mm NZn

ph

2:4:4+

NZn

38

HYDROBIOSIDAE: TANORUS 2:4:4+

densus (KORBOOT,1964)(j) m:8.5mm

empheres NEBOISS, 1984 m: 12mm NG

giluweana NEBOISS, 1984 m: 10.5mm NG

papuana (KIMMINS, 1962) m:10mm;f:11 mm NG

39

HYDROBIOSI DAE: EDPERCIVALIA 2:4:4+

maxima (McFARLANE,1939)(f) m: 11.5 -14 mm; f: 14 -15 mm NZs

harrisoni WISE, 1982

banksiensis (McFARLANE, 1939) m: 11 mm NZs

fusca (McFARLANE,1939) m: 10 -12 mm; f: 9.5 -10.5 mm NZs

40

HYDROBIOSIDAE: EDPERCIVALIA 2:4:4+

shandi (McFARLANE,1951) m:9-10.5mm;f:10-11 mm NZs, ST

thomasoni (McFARLANE, 1960) m: 9.5 - 10.5 mm; f: 9 mm NZn

borealis (McFARLANE, 1951) m: 8.5-9 mm NZn

spaini McFARLANE, 1973 m: 11 -12.5mm;f: 11 mm NZs

41

HYDROBIOSIDAE: EDPERCIVALIA 2:4:4+

cassico/a (McFARLANE,1939) m:10.5-12mm;f:11-11.5mm NZ

st2

st3

ph D-L

HYDROBIOSIDAE: HYDROCHOREMA 2:4:4+

crassicaudatum TILL YARD, 1924 G) m: 6 - 7 mm; f: 6 - 7.5 mm NZ,ST

ph

tenuicaudatum TI LL YARD, 1924 m:8.5-10.5mm;f:10-11 mm NZ

42

HYDROBIOSIDAE: NEUROCHOREMA 2:4:4+

confusum (McLACHLAN, 1868) (j) m: 6.5 - 8 mm; f: 7 - 8.5 mm NZ

armstrongi McFARLANE,1951 m: 8 - 9 mm; f: 7.5 - 8.5 mm NZ

pilosum McFARLANE,1964 m:6mm NZs

forsteri McFARLANE,1964 m: 8.8 mm; f: 9.8 mm NZs

43

HYDROBIOSIDAE: PSILOCHOREMA (d 0:4:4+) 2:4:4+

mimicum McLACHLAN,1866(f) m: 8 - 9.5 mm; f: 8 -10 mm NZ

tautoru McFARLANE, 1964 m: 8.5 - 9.5 mm; f: 9 mm NZs

acheir McFARLANE, 1981 (size not given) NZs, ST

v

44

HYDROBIOSI DAE: PSI L OCHOREMA (cf 0:4:4+) 2:4:4+

vomerharpax McFARLANE,1964 m: 7 - 8 mm; f: 7.5 - 8.5 mm NZs

leptoharpax McFARLANE,1951 m: 7 - 8 mm; f: 7.5 - 9 mm NZ

cheirodes McFARLANE,1981 m: 7 mm; f: 7.5 - 8 mm NZs, ST

45

HYDROBIOSIDAE: PSILOCHOREMA (c:f 0: 4: 4+) 2:4:4+

nemorale McFARLANE,1951 m: 8 - 9.5 mm; f: 8 - 9 mm NZ

embersoni WISE, 1982 m: 11.Smm NZs

folioharpax McFARLANE, 1956 m: 6.3 - 7 mm; f 9 mm NZs

mataura McFARLANE,1956 m:8mm NZs, ST

46

HYDROBIOSIDAE: PSILOCHOREMA (cf 0:4:4+) 2:4:4+

donaldson; McFARLANE, 1960 m: 7.5 mm; f: 7.5 - 9 mm NZ n

bidens McFARLANE, 1951 m: 7.5 - 8.5 mm; f: 8 - 9 mm NZ

macroharpax McFARLANE, 1951 m: 7 - 8 mm; f: 8 - 9 mm NZ

47

HYDROBIOSIDAE: TlPHOBIOSIS 2:4:4+

montana TILLYARD, 1924(j) m: 4.5 - 5.5 mm; f: 4.5 mm NZs

trifurca McFARLANE, 198-1 m: 6.4 mm; f: 6.2 mm NZ s

intermedia MOSELY, 1953 m: 5 - 5.5 mm; f: 4 - 5 mm NZ s

~h

48

HYDROBIOSIDAE: TIPHOBIOSIS 2:4:4+

salmoni McFARLANE, 1981 m: 5 - 7.5 mm; f: 7.6 mm NZ 5

veniflex McFARLANE,1960 m: 8 mm; f: 8.5 mm NZ

t'; ,

, Iii

kuscheli WISE, 1972 m:3.5mm AC

~ i.l \1' I. \ ;1 ~V1

49

HYDROBIOSI DAE: TIPHOBIOSIS 2:4:4+

plicosta McFARLANE,1960 m: 5:8 mm NZn

child; McFARLANE, 1981 m: 4.8 - 6.4 mm; f: 7.6 - 8.7 mm NZ

fulva TILLYARD,1924 f: 8 mm (species not recognisable) NZs

50

HYDROBIOSIDAE: SYNCHOREMA 2:4:4+

zygoneura TlLLYARD,1924(D m:6-7mm;f:6mm NZs

til/yardi McFARLANE, 1964 m: 6 - 7 mm; f: 7 - 8 mm NZ

51

HYDROBIOSIDAE: MORUYA 2:4:4+

opora NEBOISS, 1962 m: 7.5 - 9 mm; f: 8 - 9.5 mm TA

<? v

charadra NEBOISS, 1962(i) m: 5 - 6.5 mm; f: 6 - 7 mm TA

<? v D

tasmanica (JACQUE MART, 1965) m:9-10mm TA

52

HYDROBIOSIDAE: PS YL LOBETINA 2:4:4+

locula NEBOISS, 1962 m: 7 - 9.5 mm; f: 8 - 10 mm AUse

ina 0 v

plutonis BANKS, 1939 CD m: 7 - 9 mm; f: 8 - 9 mm AU ne,se

ina o

perkinsi NEBOISS, 1962 m: 7 - 8 mm; f: 7 - 8.5 mm AUne

ina

o

53

HYDROBIOSI DAE: PS YL LOBETINA

attunga NEBOISS, 1962 m: 7 - 9 mm; f: 8 - 9 mm AUse

ina D

cumberlandica NEBOISS, 1962 m: 7 - 8.5 mm; f: 7 - 9 mm

ina D

HYDROBIOSIDAE: ALLOBIOSIS

erratica MOSELY, 1953 (j) m: 6 mm; f: 6 mm AUse

\_ .. -

AUse

2:4:4+

( ,~'

~ V

2:4:4+

54

GLOSSOSOMATIDAE

AduLts of this famiLy are generaLLy smaLL, duLL,

greyish to bLack insects,with wing span of 8 to 12 mm;

aLL rather uniform in appearance.

OceLLi present. MaxiLLary paLpi 5-segmented in both

sexes, the first two segments short, subequaL, second

with gLobuLar mesoLateraL projection,third segment the

Longest. Antennae stout, shorter than forewing.

Mesoscutum and scuteLLum each with pair of setaL

warts. Wings eLongate, ovate apicaLLy, discoidaL ceLL of

forewing cLosed. A characteristic, more or Less dis

tinct, bLister-Like protuberance near the dorsaL mar

gin of abdominaL sternite 5 in both sexes (species of

SW Pacific - AustraLian region); ventraL process on

sternite 6 usuaLLy present. In femaLes the midtibia and

tarsus more or Less diLated, abdominaL segments 8 and

9 each with Long internaL apodeme on either side.

TibiaL spurs 2:4:4.

The famiLy, which was regarded as a subfamiLy with

in the famiLy RhyacophiLidae, occurs in aLL faunaL re

gions and there are about 20 genera with more than 400

species. ALL species from the SW Pacific - AustraLian

region are here referred to the genus Agapetus. It is

aLso noted that the famiLy is weLL represented in Aus

traLia and New Guinea and on other SW Pacific isLands,

but it does not occur in New ZeaLand.

Synopsis of genera

(OnLy one genus in the region)

Agapetus Curtis, 1834 (17)

~ midLeg

55

GLOSSOSOMATIDAE: AGAPETUS 2:4:4+

tasmanicus (MOSELY. 1953) m: 3.5 - 5 mm; f: 4 - 4.7 mm TA

A r---_

kimminsi ROSS. 1956 m:5.5mm AUse

ina

v

jafiwi ROSS. 1951 m:4mm NG

productus (KIMMINS. 1962) m:3mm NG

~FT :: "'- ph L ',. • ... i'" ~-....,~ ~ ....... , ,.'

56


dayi ROSS, 1956 m: 5.5 mm AU se

latosus ROSS, 1951 m:3.5mm NG

ina

monticolus BANKS, 1939 m: 3.5 - 5 mm AUse

diacanthus NEBOISS, 1986 m: 4 - 5 mm

/ ----------_ ..

apalapsili (MALICKY, 1978) m: 3.5 - 4.5 mm NG

G LOSSOSOMAT I DAE: AGAPETUS

eralus (MOSELY, 1953) m: 4 - 5 mm; f: 4.5 - 5.5 mm

pontona (MOSELY, 1953)

laparus NEBOISS, 1977

ulmeri ROSS, 1951

m:4.5mm

V

m:4-4.5mm

m:3.5mm NG

AUse

TA

57

2:4:4+

TA

<;> mid leg

58


nokowoula NEBOISS, 1986 m:4.2mm NH

ph L

salomonis (KIMMINS, 1957) m: 3.5 mm GU

evansi ROSS, 1956 m:5.5mm AUse

ablusus NEBOISS, 1986 m: 4 - 5 mm; f: 3.8 - 4.8 mm AUse

59

HYDROPTILIDAE

This is a Large famiLy,commonLy known as the micro

cadises; they are the smaLLest of aLL Trichoptera with

wing span of 4 to 12 mm.

OceLLi either present or absent according to genus.

MaxiLLary paLpi 5-segmented in both sexes, the first

two segments very short, the apicaL segment simpLe.

The antennae are shorter than the forewing; shorter in

femaLes than in the maLes; basaL segment shorter than

the head.

Mesoscutum without setal warts,scutellum flat, tri

angular, pointed posteriorLy, margins verticaL. Wings

elongate, sLender, usualLy acute apically, densely co

vered with fine pubescence, hindwing fringe with very

Long hairs,longer than the width of the wing; venation

reduced, interpretation of veins often very difficult.

TibiaL spur3 variable 0-1:2-3:3-4.

The fa~ily is known from alL faunaL regions and

there are about 80 genera with 650 species. In the

SW Pacific - Australian region 14 genera,with 132 spe

cies,are recorded.

Synopsis of genera

Acanthotrichia Wells, 1982 (1)

Acritoptila Wells, 1982 (3)

Austratrichia WeLls, 1982 (1)

Caledonotrichia Sykora, 1967 (2)

Hellyethira Neboiss, 1977 (18)

Hydroptila Da lman, 1819 (20)

Maydenoptila Neboi ss, 1977 (8)

Mulgravia WeLls, 1982 (2)

Orphninotrichia MoseLy, 1934 (9)

Orthotrichia Eaton, 1873 (51)

Targatrichia Neboiss, 1977

Oxyethira Eaton, 1873

subgenus Trichoglene Neboiss, 1977 (6)

subgenus Dampfitrichia Mosely, 1937 (4)

Gnathotrichia Ulmer, 1951

Stenoxyethira Kimmins, 1951

Paroxyethira MoseLy, 1924 (5)

Tricholeiochiton Kloet & Hincks, 1944 (5)

Xuthotrichia MoseLy, 1934 (1)

60

HYD ROPTI LI DAE: HYDROPTI LA 0:2:4-

robusta WELLS, 1979 m: 2.3 mm; f: 2.5 mm AU sa

~ ms

tasmanica MOSELY, 1934 m: 3 mm TA

losida MOSELY, 1953 m· 3 mm; f: 3.2 mm AU na,sa

61

HYDROPTI LlDAE: HYDROPTILA 0:2:4-

explicata WELLS, 1984 m: 2.4 mm NG furcula WELLS, 1984 m: 2.3 - 2.5 mm NG

quadrifida WELLS, 1984 m: 1.9 - 2.3 mm NG

bispina KIMMINS, 1962 m: 2.3 mm; f: 2.5 mm NG

1 v

~ \ 0°

incertula MOSELY, 1934 m:2.5mm NG, AU ne,se

e:?§ 25>-~

62

HYD ROPTI LI DAE: HYDROPTI LA 0:2:4-

tri/obata KIMMINS, 1957 m: 2 mm NG, GU

bugata WELLS, 1984 m: 1 .7 - 2.1 mm NG

setigera WELLS, 1984 m: 2.5 - 2.7 mm NG,NB

,1 ; ~I :. :,'~ . :.

. :> ... ~ i· sinuosa WELLS, 1979 m:2.2mm AU ne

obscura WELLS, 1979 m: 2 mm; f: 2.3 mm AUne

63

HYDROPTILIDAE: HYDROPTILA 0:2:4-

ca/cara WELLS,1979 m: 2.4 mm; f: 2.4 mm

A~rY=··~ d'

~

acinacis WELLS, 1979 m: 2.4 mm; f: 2.5 mm AUse, TA

scamandra NEBOISS, 1977 m: 2.5 mm; f: 2.7 mm AU ne,se

64

HYDROPTI LlDAE: HYDROPTILA 0:2:4-

koropa WELLS,1984 m: 2 - 2.2 mm NG

vittata WELLS, 1984 m: 2 - 2.7 mm; f: 2.1 - 2.9 mm NG

caperata WELLS, 1984 m: 2.1 - 2.3 mm NG

warisa WELLS, 1984 m: 1.7 - 2.2 mm; f: 1.7 - 2.4 mm NG

65

HYDROPTI LlDAE: ORPHNINOTRICHIA 0:2:4+

maculata MOSELY, 1934(i) m: 2.5 mm; f: 3.4 mm AU ne,se ,TA

J\

benambrica WELLS, 1983 m: 2.4 - 3 mm; f: 2.7 - 3.2 mm AUse

regia WELLS, 1980 m: 2 mm; f: 2.4 mm AUse

66

HYD ROPTI LI DAE: ORPHNINOTRICHIA 0:2:4+

media WELLS, 1980 m: 1.9 mm AUse subu/at,9. WELLS, 1983 m: 1.6 mm AU se

acta NEBOISS, 1977 m: 3 - 3.5 mm; f: 3.5 mm TA

·V

silicis WELLS, 1980 m:2.5mm AUne

\

67

HYDROPTI LI DAE: ORPHNINOTRICHIA 0:2:4+

papillata WELLS, 1980 m: 2.2 mm; f: 2.4 mm AU se

~ ~f ::4 .............. r ~! .. '-" n / ..•..... , '-~. :' '"" .'

i \ ' (r ......... i -~,.\ ...•. i.

justini WELLS, 1983 m: 2 - 2.3 mm; f: 2 - 2.5 mm AUse

68

HYDROPTI LI DAE: AUSTRATRICI-tIA 0:3:4+

neboissi WELLS, 1982(j) m: 2.7 - 2.8 mm; f: 2.6 mm AUse

HYDROPTI LI DAE: MULGRA VIA 0:3:4+

coronata WELLS, 1982(j) m: 2.7 mm AU ne

carteri WELLS, 1983 m: 2.5 - 2.8 mm; f: 2.6 - 3.1 mm AUse

69

HYDROPTI LlDAE: ACRITOPTILA 0:3:4+

g/obosa WELLS, 1982 (i) m: 1.9 - 2.3 mm; f: 1.6 - 2 mm AUsw

hamatus WELLS, 1982 m:2.4mm

margaretae WELLS, 1982 m: 2.7 mm; f: 2.2 - 2.4 mm AUsw

70

HYDROPTI LlDAE: CALEDONOTRICHIA 0:3:4+

iIIiesi SYKORA, 1967 (!) m:3mm

minor SYKORA, 1967 m:2.Smm

71

HYDROPTILIDAE: MAYDENOPTILA 0:3:4+

cuneola NEBOISS, 1977(j) m:3.5-4mm;f:4mm AUse, TA

antennifera WELLS, 1983 m: 3.5 - 3.7 mm AUse

'" . \.

r-----~Y1., ... ~ ~ .. \ \

kurandica WELLS, 1980 m:3mm AU ne

72

HYDROPTI LlDAE: MAYDENOPTILA 0:3:4+

baynes; WELLS, 1983 m: 2.1 - 2.8 mm; f: 2.6 mm AU sw

rup;na NEBOISS, 1977 m: 2.1 - 3 mm; f: 2.2 - 3.1 mm AU se, TA

exp/;cata WELLS,1980 m: 2.9 - 3.5 mm TA

~)v. " v.,

rg , :X

m: 2.6 mm AU se

HYDROPTILIDAE: MAYDENOPTILA

pseudorupina WELLS, 1980 m: 3 mm;f: 3 mm

HYDROPTI LlDAE: XUTHOTRICHIA

ochracea MOSELY, 1934 (i) m: 3 mm AU ne,se

HYDROPTI LI DAE: ACANTHOTRICHIA

bilamina WELLS, 1982(i) m:2.3mm AU ne,se

73

AU ne,se

v

./ ::.

• 0,

o 00

.'

1 :3:4+

0:3:4+

0:3:4+

74

HYDROPTI LI DAE: TRICHOLEIOCHITON 0:3:4+

fidelis WELLS, 1982 m: 2 - 2.5 mm; f: 3 - 3.3 mm AU nW,ne

jabirel/a WELLS, 1985 m: 2.2-2.5 mm; f: 2.5 mm AU n

~j ~ .n09.f{~~YN !\\J' ···':r:··ll .. Yf\ *~r~·:t~, ~'7j~l.~\\

~~ ~'i'--:4 ( , ... 1\'.11' •. ~

------...:.... .----"",'

edmondsi WELLS, 1982 m: 2.2 mm; f: 2.3 - 2.5 mm AUnw

r '. ", .. :;"

75

HYDROPTI LI DAE: TRICHOLEIOCHITON 0:3:4+

tridens WELLS, 1982 m: 1.7 - 2.5 mm; f: 2 - 2.5 mm AU nW,n

0 0 0 00 0: 0° 0 0

o 0 00 0 0 0

v

bifurca WELLS, 1982 m: 1.9 - 2.4 mm; f: 2 - 3.2 mm AU nW,n

76

HYDROPTI LlDAE: HELLYETHIRA 0:3:4+

simp/ex (MOSELY, 1934) CD m: 3.8 mm; f: 3.5 mm AU ne,se, TA

cubitans WELLS, 1979 m: 2 mm AU nW,ne

basi/obata WELLS, 1979 m: 2.8 mm AU se, TA

pu/vina WELLS, 1979 m:2mm AUnw

/oripes WELLS, 1979 m: 1.9mm AUnw

77

HYDROPTI LlDAE: HELL YETHIRA 0:3:4+

exserta WELLS. 1979 m: 2.1 mm; f: 2 mm AU se, TA

multi/obata WELLS, 1979 m: 2.9 mm; f: 2.5 mm AU se, TA

malleoforma WELLS, 1979 m: 2.7 mm; f: 2.8 mm AU ne,se, TA

litua WELLS, 1979 m: 3.1 mm AUsw

78

HYDROPTI LI DAE: HELL YETHIRA 0:3:4+

fimbriata (MOSELY,1934) m: 2.5 mm AU se

dentata WELLS,1979 m: 1.8 mm AU nw

allynensis WELLS, 1979 m: 2.1 mm AUse

corn uta WELLS, 1979 m: 2.4 mm; f: 2.2 mm AU ne

ramosa WELLS, 1983 m: 1.5 - 2.2 mm; f: 2.3 - 2.4 mm AU n

v

79

HYDROPTI LlDAE: HELL YETHIRA 0:3:4+

vernoni WELLS, 1983 m: 2.1 - 2.5 mm; f: 2 - 2.3 mm AUne

v

veruta WELLS, 1985 m: 1.5-1.7 mm; f: 1.6-2.1 mm AU n

.. " .. ' " .. ,; v eskensis (MOSELY, 1934) m: 3 mm; f: 2.4 mm AU ne

v

sentisa WELLS, 1979 m: 2.5 mm; f: 2.4 mm AUnw

80

HYDROPTI LlDAE: PAROXYETHIRA 0:3:4+

hendersoni MOSELY, 1924 (f) m: 3 mm; f: 3 - 4.5 mm NZ,ST

eatoni MOSELY, 1924 m:3mm;f:3mm NZs

hintoni LEADER,1972 m: 2.5 mm; f: 2.6 mm NZ

'::;-::'::;::===::~,;!':''''.::..'''::==:::::::::::==:::::::::::

C-~-

81

HYDROPTI LlDAE: PAROXYETHIRA 0:3:4+

kimminsi LEADER, 1972 m: 3 mm; f: 3 mm NZ n

til/yardi MOSELY, 1924 m: 3.5 mm; f: 3.5 mm NZ

82

HYDROPTI LI DAE: OX YETHIRA (DAMPFITRICHIA) 0:2:4+

artuvillosus (WELLS, 1981) AUnw

.... -~./

plumosa (WELLS, 1981) m:3.2mm AUne

warramunga WELLS, 1985 m :1.1-1.5 mm; f: 1.4-2 mm AU n

v

incana ULMER,1906 m: 2.2 - 2.8 mm; f: 2.5 - 3 mm JA, SM, NG, AU n 0:3:4+

83

HYDROPTILIDAE: OXYETHIRA (TRICHOGLENE)

columba (NEBOISS, 1977) ® m: 1.7 - 2.8 mm; f: 1.7 - 2.5 mm

triangulata WELLS, 1981 m: 1.7 - 2.1 mm; f: 1.6 - 2 mm

' ......

mienica WELLS, 1981 m: 2.3 mm; f: 2.3 mm TA

(

AU ne,se, TA

v

AU ne

v

0:3:4+

.(tfj) a~ ril ..•.. " .. ~

84

HYDROPTI LlDAE: OX YETHIRA (TRICHOGLENE) 0:3:4+

brevis WELLS, 1981 m: 2.3 mm; f: 2.9 mm AUsw

v

retracta WELLS, 1981 m: 2.2 mm; f: 2.2 mm AUsw

v

a/biceps (McLACHLAN, 1862) m: 2.4 - 3 mm; f: 3 - 3.5 mm NZ, ST, CH, C, AC Antipodes; Snares

v

v D

85

HYDROPTI LlDAE: OR THO TRICHIA

adornata WELLS, 1979

zonata (NEBOISS, 1977) m: 3.5 - 3.7 mm; f: 3.5 - 4 mm TA

atraseta WELLS, 1979 m: 2.9 mm; f: 3.1 mm AUse

v

0:3:4-

~ 0 .. ~. : .. .

~.;.( .... ~':' ,.'.". . . .

l,' ...... .

~t,,~~~~ __ ' I

I:

1\ \,'~::<:-.

86

HYDROPTI LlDAE: ORTHOTRICHIA 0:3:4-

rostrata WELLS, 1979 m: 2.3 mm; f: 2.8 mm AUse

bishopi WELLS, 1979 m: 2.4 mm; f: 2.6 mm AUse

armata WELLS, 1979 m:2.4mm AUse

di/gri WELLS, 1983 m: 3.2 mm; f: 2.9 mm AUse

87

HYDROPTI LlDAE: ORTHOTRICHIA

scutata WELLS, 1979 m: 2.3 mm; f: 2.6 mm AU nW,n

turrita WELLS, 1979 m: 1.9mm AUnw

orbostensis WELLS, 1979 m:2.8mm AUse

, "

0:3:4-

v

~

aculeata WELLS, 1979 m: 1.7 mm AUnw

(~ ~==------

88

HYDROPTI LlDAE: ORTHOTRICHIA

velata WELLS, 1983 m: 1.9 - 2.2 mm; f: 2.2 - 2.4 mm AU nW,ne

-":':\ ," J \:- :'--".'1. ... ,.~."'-.'.".,

~ v

inornata WELLS, 1979 m:1.7mm AUnw

~

eurhinata WELLS, 1985

':".- ... ,

suteri WELLS, 1979

~S~

m: 1.7 mm AU n

m: 1.4 mm; f: 1.3 mm

,.' 0 "0 0 o c

o 0

o 0 o

I D. '\

bullata WELLS, 1979

0:3:4-

m: 1.9 mm AU nW,ne

89


kokodana KIMMINS, 1962 m:2.7mm NG obscura KIMMINS, 1962 m:2.7mm NG

~ .. "'., .: ",

..... ''i ... )

\( \ v

v

fragilis WELLS, 1984 m: 1.3 -1.5 mm NG

cucullata WELLS, 1984 m:3.3mm NG

tumoris WELLS, 1984 m:2.6mm NG

advena WELLS, 1984 m:2.3mm NG

90


glebula WELLS, 1984 m: 2.6 - 2.8 mm NG disparalis WELLS,1984 m: 2.4 mm NG

annulata WELLS, 1984 m:3.1 mm NG lentigo WELLS, 1984 m: 2.7 - 3.4 mm NG

litotes WELLS, 1984 m: 1.3 mm NG

talea WELLS, 1984 m:2.2mm NG

cinctigera WELLS, 1984 m:1.S-1.7mm NG

91


morula WELLS, 1979 m: 2.4 mm; f: 2.2 mm AUne

alveata WELLS, 1979 m: 2.6 mm AU ne

digitata WELLS, 1984 m: 2.1 - 2.5 mm NG

92

HYDROPTI LI DAE: ORTHOTRICHIA 0:3:4-

gracilis WELLS, 1979 m: 1.B mm; f: 1.9 mm AU ne,se

paranga WELLS, 1979 m: 1.B mm AUnw

kh%ensis WELLS, 1979 m: 1.6 mm AUne

exigua WELLS, 1979 m: 1.7mm AU nW,ne

stipa WELLS, 1979 m: 1.7 mm AUnw

93


yabbaca WELLS, 1983 m: 2,5 mm; f: 2 - 2.6 mm AU ne

urimica WELLS, 1984 m: 1.2 - 1.6 mm; f: 1.5 mm NG

o f}

attenuata WELLS, 1983 m: 1.7 - 2 mm; f: 2.2 - 2.4 mm AUse

v

94


tyleri WELLS, 1979 m: 1.8 mm AUnw bellicosa WELLS, 1979 m: 1.Bmm AUnw

tortuosa WELLS, 1979 m: 2.5 mm; f: 2.9 mm AU na,sa

capi/lata WELLS, 1979 m:2.2mm AU na,sa

ensiformis WELLS, 1984 m: 1.2 - 1.5 mm NG,NB

95

HYD ROPTI LI DAE: ORTHOTRICHIA 0:3:4-

aberrans WELLS, 1979 m:3.Smm AUse divaricata WELLS, 1983 m:4.9mm AUne

( \

conferta WELLS, 1983 m: 4 mm; f: 3.8 mm AUse

v

muscari WELLS, 1983 m:3.5mm AUne

96


flabella WELLS, 1983 m:4mm

m: 3.5 - 3.8 mm AU se

97

PHI LOPOTAMIDAE

Adults of this family are small to medium sized in

sects with wing span from 12 to 20 mm, often dark, blac

kish or mottled greyish-brown.

Ocelli present. Maxillary palpi 5-segmented in both

sexes, the second segment has meso-distal brush of se

tae; segment 5 long, flexible.

Mesoscutum without scutal warts, although sometimes

with small groups of setae; scutellum usually with a

pair or with one fused setal wart. Wings elongate,

ovoid, venation complete, discoidal cell present and

closed in both wings, median cell closed only in fore

wing; small hyaline areas at cross-veins and at arcu

lus in the forewing.

Tibial spurs 1:4:4 or 2:4:4.

The family occurs in all faunal regions. Approxi

mately 15 genera with 400 species are known, of which

5 genera with 53 species are recorded from the SW Pa

cific - Australian region.

Synopsis of genera

Chimarra Stephens, 1829 (26)

Chimarrha Burmeister, 1839

Cryptobiosella Henderson, 1983 (4)

Hydrobiosella Tillyard, 1924 (19)

Zelobiosella Mosely, 1953

Neobiosella Wise, 1958 (1)

Xenobiosella Henderson, 1983 (1)

98

PHI LOPOTAMIDAE: CRYPTOBIOSELLA 2:4:4+

furcata HENDERSON,1983 m:7mm NZn

tridens HENDERSON, 1983 CD m: 6.7 - 8.9 mm NZs

hastata HENDERSON,1983 m:8.8mm NZn

spinosa HENDERSON,1983 m:6.6mm NZn

99

PHILOPOTAMIDAE: HYDROB/OSELLA 2:4:4+

amblyopia NEBOISS, 1982 m: 5.5 -7 mm; f: 6 -7 mm AU sw

michaelseni (ULMER, 1908) m: 6.5 - 7 mm; f: 7 - 7.5 mm AU sw

waddama MOSELY, 1953 m: 6.5 - 8 mm; f: 7 - 9 mm AU se, TA

PHILOPOTAMIDAE: HYDROBIOSELLA

stenocerca TILLYARD,1924(i) m:7-9 mm

mixta (COWLEY, 1976) m:8-10mm NZ

\

uncinata KIMMINS, 1953 m:3.7mm NC

I I I I I

100

NZ

,. If '

Ig10 \

~

~ tonela (MOSELY, 1953) m: 4.5 - 5.5 mm NZ

aorere HENDERSON,1983 m: 4.9 - 5.6 mm NZs

2:2:4+

ph L

~-J ph L

101

PHILOPOTAMIDAE: HYDROBIOSELLA 2:4:4+

armata JACQUEMART, 1965 m: 5.5 - 7 mm; f: 6 - 7 mm TA

tasmanica MOSELY,1953 m:7.5-9.5mm;f:8-9.5mm TA

cognata KIMMINS, 1953

/

,./

I

.~

m: 7.5 - 8.5 mm; f: 8 - 8.5 mm

:::::::."::::::::.~ =~ -";;::::::'::::"~~I ..-

}/ I

. ~. .', .. ' : .. ':--_-.. ",','.

sagitta NEBOISS, 1977 m: 7.5 - 8.5 mm; f: 7.7 mm

letti KORBOOT, 1964 m:4.5mm AUse

TA

TA

102

PH I LOPOTAM I DAE: HYDROBIOSELLA

orba NEBOISS, 1977

corinna NEBOISS, 1977

anasina NEBOISS, 1977

cerula NEBOISS, 1977

, I I

bispina KIMMINS, 1953

m:6mm TA

m: 7.5 - 8.5 mm; f: 7.5 - 9 mm

m: 7 - 9 mm; f: 7 - 8.5 mm TA

m: 7-8 mm TA

m:8mm AUse

2:4:4+

TA

103

PHILOPOTAMIDAE: HYDROBIOSELLA 2:4:4+

arcuata KIMMINS, 1953 m:8mm AUne

PHI LOPOTAMIDAE: NEOBIOSELLA 2:4:4+

jrrorata WISE, 1958 (j) f:6mm NZn

PHI LOPOTAMI DAE: XENOBIOSELLA 2:4:4+

motueka HENDERSON, 1983 (j) m: 5.3 - 5.9 mm

PHILOPOTAMIDAE:

manni (BANKS, 1924)

indigota MOSELY, 1941 Syn. (?) of manni (Banks)

obscurel/a (BANKS, 1924)

I I

:' ! , , ,

,---/

104

CHIMARRA 1:4:4+

m:6.Smm FI

m:6.Smm FI

m:4.6mm FI

ina caudal

anoac/ana (MALICKY, 1978) m:4mm NG

105

PHILOPOTAMIDAE: CHIMARRA 1:4:4+

sinuosa KIMMINS, 1962 m:4.7mm

ina V

falcata KIMMINS, 1962 m:5.4mm

aiyura KORBOOT, 1965 m:5.5mm NG

austra/ica (ULMER, 1916) m:5mm AU ne

monticola KIMMINS, 1953 m: 5.5 mm AU se

106

PHILOPOTAMIDAE: CHIMARRA 1:4:4+

ulmer; KIMMINS, 1962 m: 4.7 mm NG

sedlacek; SYKORA, 1967

,,------ ......... t:-----··,\ ......... _-

\ I I I

goroca SYKORA, 1967

uranka MOSELY, 1953

aureofusca KIMMINS, 1957

m:5.5mm NG

m:5.5mm NG

m:5mm AU ne

m:4.2mm

ina V

ph o

L

ph

o

GU

107

PHI LOPOTAMI DAE: CHIMARRA 1:4:4+

papuana KIMMINS, 1962 m: 4.1 mm NG

sabrona KIMMINS, 1962 m:3.2mm NG

schmidi KIMMINS, 1962 m:4.5mm NG

cheesmanae KIMMINS, 1962 m:4mm NG

cyclopica KIMMINS, 1962 m:4.2mm NG

L

108

PHILOPOTAMIDAE: CHIMARRA 1 :4:4+

/oriana (NAVAS, 1933) m:4.3mm NG

" .... ;,..---- ----..

, ,

1 ~;

biramosa KIMMINS, 1957 m: 4.5 mm; f: 4.7 mm GU

signata (BANKS, 1936) m: 4 - 4.3 mm FI

xenillion NEBOISS, 1985 m:3.7mm NG

109

PHILOPOTAMIDAE: CHIMARRA

hienghene MALlCKY,1981 m: 6 mm NC

~:S) C--

ph L

gressitti SYKORA, 1967 m:4mm NG

ph L

kokodana KIMMINS, 1962

australis (NAVAs, 1923)

nervosa (BRAUER, 1867)

m:3.8mm

f: 7.8mm

f: 6mm

NG

AU (no figures available)

FI (no figures available)

1:4:4+

ph V

ina V

110

STENOPSYCHIDAE

All species of this family are medium to moderately

large with wing span of 18 to 35 mm,often with colour

ful black and golden wing pattern.

Ocelli absent, although position of lateral ocelli

often indicated by small tubercle (Australian species

only). Maxillary palpi 5-segmented in both sexes, seg

ment 2 with only a few coarse bristles, segment 5 as

long or longer than all other segments combined, mes

ally at the base of maxillary palpi arise elongate pi

lifers; mouth parts slightly to distinctly extended.

Antennae rather robust, about as long or slightly lon

ger than forewing, basal segment short.

Mesoscutum with a pair of rounded setal warts, se

parated by an anteriorly produced mesal section of the

scutellum; there is a single mesal wart on scutellum.

Wing venation complete, regular and similar in both

sexes; discoidal and median cells present and closed

in both wings.

Tibial spurs 3:4:4.

A total of 3 genera with about 70 species are known

from Ethiopian, Asian Palearctic, Oriental and Austra

lian regions. There is only one genus found in the

SW Pacific - Australian region, and that is endemic to

Australia. The genus Stenopsychodes was previously

placed in the family Polycentropodidae [63, 114].

Synopsis of genera

(Only one genus in the region)

Stenopsychodes Ulmer, 1916 (9)

111

STENOPSYCHIDAE: STENOPSYCHODES 3:4:4-

til/yardi BANKS, 1939 m: 12 mm; f: 15 mm AU ne

mjobergi ULMER, 1916 (i) m: 13 mm AU ne

hiemalis TI LL YARD, 1922 m: 17 mm AU se

112

STENOPSYCH I DAE: STENOPSYCHODES 3:4:4-

melanochrysa TILLYARD,1922 m: 16 mm; f: 16 -17 mm AUse

aureonigra SCHMID,1969 m:12-14mm AUse

syrdena NEBOISS, 1974 m: 12 -13 mm; f: 15 mm AUse

113

STE NOPSYCH I DAE: STENOPSYCHODES 3:4:4-

lineata NEBOISS, 1977 m: 9 -10 mm; f: 10 -11 mm TA

montana TILL YARD, 1922 m:11 mm;f:13mm AUse

opsia NEBOISS, 1974 m:11 mm;f:13-14mm AUse

114

HYDROPSYCHIDAE

A Large famiLy of caddis fLies found throughout the

worLd in great variety of habitats; there are some

smaLL species, but the majority are medium to rather

Large in size with wing span ranging from 8 to 40 mm.

OceLLi absent. MaxiLLary paLpi 5-segmented in both

sexes, but reduced or absent in certain genera of

Macronematinae; segment 5 as Long as or Longer than

the first four segments together (except in genus Her

bertorossia, in which segment 5 is about as Long as

segment 4). Antennae sLender, usuaLLy as Long as or

Longer than forewing, except in the subfamiLy Macro

nematinae, in which they are much Longer.

Mesoscutum without setaL warts; scuteLLum with one

median setaL wart. Wings with venation compLete; dis

coidaL and median ceLLs present and cLosed in forewing;

hindwing much wider than the forewing.

TibiaL spurs variabLe 0-2:2-4:2-4.

The famiLy is known from aLL faunaL regions and

contains more than 900 species in 60 genera, of which

17 genera with 76 species are known to occur in the

SW Pacific - AustraLian region.

Synopsis of genera

HYDROPSYCHINAE

Abacaria MoseLy, 1941 (6)

Aoteapsyche McFarLane, 1976 (6)

Asmicridea MoseLy, 1953 (2)

Caledopsyche Kimmins, 1953 (1)

Cheumatopsyche WaLLengren, 1891

Herbertorossia ULmer, 1957 (7)

Hydropsyche Pictet, 1834 (6)

Hydropsychodes ULmer, 1905 (2)

Orthopsyche McFarlane, 1976 (2)

Smicrophylax Neboiss, 1977 (5)

DIPLECTRONINAE

Austropsyche Banks, 1939 (3)

Diplectrona Westwood, 1840 ( 16)

MACRONEMATlNAE

Aethaloptera Brauer, 1875 (1)

Baliomorpha Neboiss, 1984 (9)

Leptopsyche McLachLan, 1866 (1)

Macrostemum KoLenati, 1859 (5)

Oestropsyche Brauer, 1868 (1)

(3)

\

'/ \J ~

mxp. Abacaria

mxp. Herbertorossia

115

HYDROPSYCHIDAE: HYDROPSYCHE 2:4:4-

burgersi ULMER,1915 m:,16-17mm NG

tapena KIMMINS, 1957 m:9mm GU

flynni KORBOOT, 1964 m: 13 -14 mm; f: 17 -18 mm NG

116

HYDROPSYCHIDAE: HYDROPSYCHE 2:4:4-

bryanti BANKS, 1939 m: 8 - 10 mm; f: 9 - 11 mm JA, SM, (NG?)

ph D

mose/yi KIMMINS, 1962 m: 7.5 - 8 mm; f: 7 - 9 mm NG

hobbyi MOSELY, 1951 m: 9 mm; f: 10 mm SA,NG

ph D

testacea (NAVAS, 1933) f: 19.5 mm NG

117

HYD ROPSYCH I DAE: CHEUMA TOPSYCHE 2:4:4-

modica (McLACHLAN, 1871) m: 7.5 - 8.5 mm; f: 8 - 10 mm AU,TA

amiena SYKORA, 1967 m: 11 mm NC

ina V

beroni KUMANSKI,1979 m:8.2mm NG

118

HYDROPSYCHIDAE: HERBER TOROSSIA 2:4:4-

ungulata (ULMER, 1906) (j) m: 8 - 9 mm NG

striata KIMMINS,1962 m: 8 -10 mm NG

kimminsi KUMANSKI,1979 m: 10 -10.5 mm NG

119

HYDROPSYCHIDAE: HERBER TOROSSIA 2:4:4-

orakaivai KIMMINS, 1962 m: 6 -7 mm; f: 7 - 8 mm NG

rapsoni KORBOOT, 1964 m: 6 -7 mm; f: 7 - 8 mm NG Syn. (?) of orakaivai Kimmins

)

sabronensis KIMMINS, 1962 m: 8.5 mm; f: 9 mm NG

excavata KIMMINS, 1962 m: 7 - 9 mm; f: 8 -10 mm NG

120

HYDROPSYCHIDAE: ABA CARlA 2:4:4-

ruficeps (BRAUER, 1867) m: 7 mm; f: 8 mm FI

fijiana (MOSELY, 1934)G') m: 5 mm; f: 7 mm FI

subfusca KIMMINS, 1962 m:5mm;f:5-6mm NG

barretti KORBOOT, 1964 m: 6 mm; f: 6.5 mm NG Syn. (7) of subfusca Kimmins

121

HYDROPSYCHIDAE: ABACARIA 2:4:4-

orkeni ILLlES,1969 m: 6.5 -7 mm; f: 7.5 - 8 mm NG

picea (BRAUER, 1867) m: 5 mm (no figures available) FI

HYDROPSYCHIDAE: CALEDOPSYCHE 2:4:4-

cheesmanae KIMMINS, 1953 (i) m: 4.6 mm; f: 5 mm NC

122

HYDROPSYCHIDAE: HYDROPSYCHODES 2:4:4-

cheesmanae KIMMINS, 1962 m: 7 -7.5 mm; f: 6 mm NG

expeditionis ULMER, 1938 m: 7 mm; f: 6.2 mm NG

123

HYDROPSYCHIDAE: AOTEAPSYCHE 2:4:4-

c%nica (McLACHLAN, 1871) m: 8.5 - 9.5 mm; f: 11 - 14 mm NZ

philpotti (TILLYARD,1924) m: 7 - 8 mm; f: 9 - 10 mm NZs

ha 0

tepoka (MOSELY,1953) m:10-12.5mm;f:13.5-18mm NZ

L

raruraru (McFARLANE, 1973) (!) m: 7 - 8.5 mm; f: 9.5 - 12 mm NZ

ha 0

124

HYDROPSYCHIDAE: AOTEAPSYCHE 2:4:4-

tipua (McFARLANE, 1964) m:11.Smm NZs

ph -....;.,:jiIIL

catherinae (McFARLANE, 1960) m: 7 - 8.5 mm; f: 10- 11.4 mm NZ

125

HYDROPSYCHIDAE: ORTHOPSYCHE 2:4:4-

fimbriata (McLACHLAN, 1862) (i) m:10.5-12mm;f:11-13mm NZn

ha 0

thomasi (WISE, 1962) m:9-10mm NZn

,

,.// .................. _ ......... .

ha 0

126

HYDROPSYCHI DAE: SMICROPHYLAX

creektona NEBOISS,1977G') m: 6 -7 mm; f: 6.5 -7.5 mm

australis (ULMER, 1908)

parvula (MOSELY, 1953)

simplex (JACQUEMART, 1965)

~--------...

...

~ ..... :,';... If .. '

m: 4 - 5 mm; f: 4.5 - 5.5 mm

~:lj!"2Jfi.,,,,.,q:m If

m: 5.5 - 6 mm; f: 6.5 -7 mm

m: 4.5 - 6 mm; f: 6 - 8 mm

TA

AUse

TA

ulmeri (BANKS, 1939) m: 4.5 - 5 mm; f: 5 - 5.5 mm AU ne

2:4:4-

... ~""'·····If······ ~"

127

HYDROPSYCHIDAE: ASMICRIDEA 2:4:4-

grisea (MOSELY, 1933) ® m: 8 -10 mm; f: 9 - 11 mm TA

ph D

edwardsi (McLACHLAN, 1866) m: 8 - 10 mm; f: 9 - 11.5 mm AU se, TA

128

HYDROPSYCHIDAE: DIPLECTRONA 2:4:4-

triangulata SYKORA, 1967 m:9mm

subtriangulata KUMANSKI,1979 m:9.5mm NG

-.. --~--------', - .. , . 'w .. __ .. ", .... -'

zea/andensis MOSELY, 1953 m: 7 - 8 mm; f: 8.5 - 9 mm NZ

mafulua KIMMINS, 1962 m:6mm NG

129

HYDROPSYCHIDAE: DIPLECTRONA 2:4:4-

papuana KUMANSKI,1979 m: 7 mm NG

bulla WISE, 1958 m: 7 mm; f: 7.5 mm NZn

tasmanica JACQUEMART, 1965 m: 6 - 6.5 mm TA

angusta BANKS, 1939 m: 10 mm AUse

130

HYDROPSYCHI DAE: DIPLECTRONA 2:4:4-

cognata KIMMINS, 1953 m:7mm AUse

ph L

spinata (BANKS, 1939) m:6-7mm AU ne

castanea KIMMINS, 1953 m: 10 mm TA

/yel/a NEBOISS, 1977 m: 9.5 -10 mm TA

131

HYDROPSYCHIDAE: DIPLECTRONA

satana MOSELY, 1953

rossi KIMMINS, 1953

hystricosa NEBOISS,1978

inermis (BANKS, 1939)

m:7mm AU ne

m:7mm AU ne

m:7-9mm;f:8-10mm

( \ \"'~/

D

m:6.5mm

, , \ ... /

, , ,

ina V

AUse

2:4:4-

AU ne

132

HYDROPSYCHIDAE: AUSTROPSYCHE

victoriana BANKS, 1939 (j) m:, 11 - 12 mm; f: 11.5 - 14.5 mm

bispinosa (JACQUEMART, 1965) Syn. (7) of victoriana Banks

/ I

/./'

I /' \/

--

bifurcata (KIMMINS, 1953)

m: 12 mm AUse

m: 11 mm AUse

2:4:4-

AUse

133

HYDROPSYCH I DAE: BALIOMORPHA 1 :4:4-

pulchripenne (TILL YARD, 1922) (j) m: 11 - 12.5 mm; f: 11 - 12.5 mm AU se

i//------

I

\ \~----

m: 9.5 - 11.5 mm; f: 7.5 - 9 mm

dubia (ULMER,1905) m: 17 - 21 mm; f: 19 -20 mm AUse

banksi (MOSELY, 1953) m: 9.5 - 11.5 mm; f: 7.5 - 8.5 mm AU ne

134

HYDROPSYCHIDAE: BALIOMORPHA 1:4:4-

caudicea NEBOISS, 1984 m: 12 mm NG

echinata NEBOISS, 1984 m: 10.7 mm NG

pezidion NEBOISS, 1984 m: 7.5 mm; f: 7.5 mm NG

chiloma NEBOISS, 1984 m:8.5mm NG

(oriai (NAVAS, 1933) f: 8.8 mm NG

135

HYDROPSYCHIDAE: MACROSTEMUM

saundersi (McLACHLAN, 1866) m: 9.5 - 11 mm; f: 10 - 11 mm

r={'<;(~ \, ! \ .... ~-_/

loriai (NAVAS, 1930) m: 12 -13 mm; f: 12 -13 mm

auriferum NEBOISS, 1984 m:9.5-11 mm;f:10-11 mm

wallacei (McLACHLAN. 1866) f: 11 mm dulce (McLACHLAN, 1866) f: 8.5 mm

NG (p. VI)

NG (Mysoil

ITypes lost, no figures available)

NG

2:4:4-

NG, AU ne

NG

HYDROPSYCHIDAE: LEPTOPSYCHE 2:4:4-

gracilis McLACHLAN,18660) m: 13 mm NG

136

HYDROPSYCHI DAE: AETHALOPTERA

sexpunctata (KOLENATI.1859) m: 8 -11 mm;f: 8-11 mm

, , I

I I ,

,,' ,

HYDROPSYCHIDAE: OESTROPSYCHE

vitrina (HAGEN. 1859) m: 11 -17 mm; f: 11 -16 mm

cf

NG

0:3:2-

NG, AU ne

st 8

1:2:2-

137

POL YCENTROPODIDAE

SmaLL to medium size, aduLts rather stockiLy buiLt,

wing span of 8 to 25 mm, forewings usuaLLy dark brown

mottLed with whitish to cream or goLden spots; found

in vide variety of habitats.


sexes, first two segments short, each shorter than the

third; the third segment arises before the apex of the

second, apicaL segment Long, annuLated, fLexibLe. An

tennae stout, about as Long as, or sLightLy shorter

than forewing, individuaL segments short.

Mesoscutum with a pair of rounded setaL warts,

sometimes setiferous punctures aLso present; scuteLLum

with a singLe rounded mesaL wart. Forewings moderateLy

broad, denseLy pubescent, discoidaL and median ceLLs

cLosed; R1 not forked apicaLLy. AbdominaL sternite 5

with LateraL fi Laments.

The subfamiLy HyaLopsychinae (regarded as famiLy by

some authors) is characterised by Large eyes, particu

LarLy in maLes, Less so in femaLes; maxiLLary paLpi

Less deveLoped, apicaL segment short, not annuLate;

LabiaL paLpi absent.

TibiaL spurs 3:4:4, in femaLes the mid-tibia usu

aLLy flattened.

The famiLy is known from aLL faunaL areas with 25

genera and approximateLy 300 species; in the SW Paci

fic - AustraLian region 9 genera with 41 species are

recorded.

Synopsis of genera

POLYCENTROPODINAE

Adectophylax Neboiss, 1982 (1)

Neureclipsis McLachLan, 1864 (1)

Nyctiophylax Brauer, 1865 (3)

Plectrocnemia Stephens, 1836 (7)

Austrecnomina Korboot, 1964

Polycentropus Curtis, 1835 (13)

Polyplectropus ULmer, 1905 (12)

Tasmanoplegas Neboiss, 1977 (1)

HYALOPSYCHINAE

Hyalopsyche ULmer, 1904 (2)

Hyalopsychella ULmer, 1930 (1)

ms

POL YCENTROPODI DAE: ADECTOPHYLAX

volutus NEBOISS, 1982 (i) m: 4.5 - 5 mm; f: 5.5 - 6 mm

POL YCENTROPODI DAE: NYCTIOPHYLAX

, , ,

parvus MOSELY, 1953

" , ,

, ,

.... _----

repandus NEBOISS, 1977

f1avus ULMER, 1915

m:3.5mm AU n

m:4mm NG

138

3:4:4-

3:4:4-

\ , ~ ................... \J

139

POL YCENTROPODI DAE: PLECTROCNEMIA 3:4:4-

maclachlani MOSELY, 1953 m: 5.5 -7.5 mm; f: 7.5 -10 mm NZ

caudata NEBOISS, 1977 m: 8 - 8.5 mm TA

~~ ((~'-- ~ ~;=lJ.!hf,i'flIi,t;:tiJIii\P" \~~- -

eximia NEBOISS, 1982 m: 9 - 9.5 mm; f: 9 - 9.5 mm AU sw

v

manicata NEBOISS, 1977 m: 8 - 10 mm; f: 8 - 10.5 mm TA

.. /

'" I I /'"

\ I I! I·lj.(

/ \! \ / \

/ \ I \

\ / "./ « V

140

POLYCENTROPODIDAE: PLECTROCNEMIA 3:4:4-

australica BANKS, 1939 m: 7-8mm AUse

,\,', 'I "

I / \~----_/

altera NEBOISS, 1977 m: 8 - 11 mm; f: 9 - 11 mm TA

v

lacuna NEBOISS, 1977 m: 8 - 9 mm; f: 9 mm TA

141

POLYCENTROPODIDAE: TASMANOPLEGAS 3:4:4-

spi/ota NEBOISS, 1977 (i) m: 5.5 - 6 mm; f: 5.5 - 6 mm TA

9 v

POL YCENTROPODI DAE: NEURECLIPS/S 3:4:4-

napaea NEBOISS, 1986 m: 5.5 - 6 mm; 6.5 - 7 mm AU se

142

POLYCENTROPODIDAE: POLYCENTROPUS 3:4:4-

grandis KIMMINS, 1962 m: 10 -11 mm; f: 13 -14 mm NG

mose/yi KIMMINS, 1962 m: 10 mm; f: 11 mm NG

mounthageni KUMANSKI,1979 m: 6.2 mm NG

piceus KIMMINS, 1962 m: 6 mm; f: 6.5 mm NG

143

POL YCENTROPODI DAE: POL YCENTROPUS 3:4:4-

australis ULMER, 191~) m: 7 - 8 mm; f: 8 mm NG

simi/is KIMMINS, 1962 m: 7 - 7.5 mm; f: 7 mm NG Syn. (?) of australis Ulmer

rosse/inus NAVAS, 1924 m: 7 mm; f: 8.5 mm NG

auricollis KIMMINS, 1962 m: 5 - 6 mm; f: 5 - 7 mm NG

144

POL YCENTROPODI DAE: POL YCENTROPUS

sinuosus KIMMINS, 1962

kenampi (KORBOOT, 1964)

drummondi ILLlES, 1969 Syn. (7) of kenampi (Korboot)

e/egans KUMANSKI,1979

m: 7 mm; f: 7 - 8 mm NG

m: 5.5 mm; f: 5.5 mm NG

m: 6.5 mm; f: 6.5 mm NG

m:6.1 mm NG

3:4:4-

145

POL YCENTROPODI DAE: POL YPLECTROPUS

greenwoodi MOSELY, 1941 m:8mm

fijianus BANKS, 1936 m:5mm FI

manni BANKS, 1936 m:4.6mm FI

r·h~J--:':-~::---==::;;·

//-------':~::::::··":~h·,·-~--t--~·~~~~~~~ I , , I

\

chapmani KUMANSKI,1979 m: 5.1 mm

FI

NG

3:4:4-

ph D

~:.-.:::-. : ina L

146

POL YCENTROPODI DAE: POL YPLECTROPUS

puerilis (McLACHLAN, 1868) m: 7.5 -10 mm; f: 10 -11.5 mm

a/tera McFARLANE,1981

aurifusca McFARLANE, 1956

waitakerensis WISE, 1962 Syn. (7) of aurifusca McFarlane

m: 8 -13 mm; f: 10 -14 mm NZn

m: 9 mm; f: 10 mm NZs

m:7-10.5mm;f:8.5-11 mm

3:4:4-

NZs, ST

9 I~:~ - - --'- . ---

NZ n (S?)

la

147

POLYCENTROPODIDAE: POL YPLECTROPUS 3:4:4-

impluvii WISE, 1962 m: 7 - 7.5 mm; f: 8.5 - 9 mm NZn

ina V ina D

puhia McFARLANE, 1956 m:9.5 mm NZs

' . ..,.. .. .:. "-";' .: .......

lacusalbinae KIMMINS, 1953 m:8mm AUse

ina V

bradleyi KIMMINS, 1957 m: 4 mm; f: 4.5 mm GU

148

POL YCENTROPODI DAE: HYALOPSYCHE 3:4:4-

d;sjuncta NEBOISS, 1980 rn: 6 mm NG,AU ne

~P'

r;valis (BETIEN,1909) f: 9 - 9.5 mm India, NG?

POL YCENTROPODI DAE: HYALOPSYCHELLA 3:4:4-

w;nkler; ULMER,1916 m:4mm BO, SM, (NG?)

149

ECNOMIDAE

Mostly small to medium sized caddis-fly species with

wing span between 6 and 18 mm, duLL greyish-brown mot

tled wi ngs.


sexes; segment 2 Longer than segment 1 and about as

Long as segment 3, apicaL segment fLexibLe. Antennae

at most as Long as the forewing.

Mesoscutum and scuteLLum each with a pair of roun

ded setaL warts. Wings narrow, rounded apicaLLy; fore

wing R1 usuaLLy forked at apex; discoidaL and median

ceLLs present and cLosed.

AbdominaL sternite 5 without LateraL fiLaments; fe

maLe abdomen terminates either bLuntLy or with eLon

gate ovipositor.

TibiaL spurs 2:4:4 or 3:4:4.

The famiLy, regarded by some authors as a subfamiLy

of Psychomiidae [63], occurs in aLL faunaL regions,

except the Nearctic. A totaL of 7 genera with more than

100 species are known, of which, 2 genera with 26 spe

cies are recorded from the SW Pacific - AustraLian re

gion.

Synopsis of genera

Ecnomina Kimmins, 1953 (17)

Ecnomus McLachLan, 1864 (9)

femaLes of Ecnomus

femaLes of Ecnomina

150

ECNOMIDAE: ECNOMUS 3:4:4-

russellius NEBOISS, 1977 m: 7.5 - 8.5 mm; f: 8 - 9 mm TA

continentalis ULMER,1916 m: 5 -7 mm; f: 5.5 - 8 mm AU ne,se

pansus NEBOISS, 1982 m: 4.5 - 6 mm; f: 5.5 -7 mm AU se,sw

cygnitus NEBOISS, 1982 m: 5 - 6.5 mm; f: 6 -7 mm AU ne,se

151

ECNOMIDAE: ECNOMUS 3:4:4-

til/yardi MOSELY, 1953 m: 6 - 7 mm; f: 6.5 - 8.5 mm AUse, TA

papuanus ULMER, 1938 m:4mm NG

cyclopicus KIMMINS, 1962 m:5mm NG

turgidus NEBOISS, 1982

m: 5 - 6 mm; f: 5.5 - 6.5 mm AUsw

atratus MOSELY, 1932 m: 3.5 - 4 mm NH

ECNOMI DAE: ECNOMINA

spinosa KIMMINS, 1953(j)

/egu/a NEBOISS, 1977

/ /

( "-- ---- --.....

vega NEBOISS, 1977

m:507mm AUse

m: 505 - 6 mm; f: 6 - 7 mm

m:505mm TA

152

3:4:4-

TA

153

ECNOMI DAE: ECNOMINA 3:4:4-

viatica NEBOISS, 1982 m: 3 - 4 mm; f: 3 - 4 mm AUsw

v

baty/e NEBOISS, 1977 m: 4- 4.5 mm;f: 4- 5 mm TA

sentosa NEBOISS, 1982 m: 4 - 4.5 mm; f: 4 - 4.5 mm AUsw

v

154


krokale NEBOISS, 1978 m:3mm AUne

chorisis NEBOISS, 1978 m:2.7mm AU ne

cohibilis NEBOISS, 1982 m: 3.5 - 4 mm; f: 3.5 - 4 mm AUsw

thinotes NEBOISS, 1978 m: 3.5 - 4 mm; f: 3.5 - 4 mm AU ne v

155


scindens NEBOISS, 1982 m: 4.5 - 5 mm; f: 4.5 - 5 mm AUsw

\/ .... J, ...... / v

irrorata KIMMINS, 1953 m: 4.5 - 5 mm; f: 5 - 6 mm TA

v

merga NEBOISS, 1982 m:4mm AUsw

156

ECNOMI DAE: ECNOMINA

trifurcata KIMMINS, 1953

!---".-~-""". : ..... :--

..•..........•.

trulla NEBOISS, 1982

/

/' I ,

_--------c:::-.....

....... -......... ,"' ",

mesembria NEBOISS, 1982

zealandica WISE, 1958

m:3,5mm AUse

m:4mm AUsw

m: 4 - 4.5 mm; f: 4 - 4.5 mm

f: 3.5 mm NZn

3:4:4-

AUsw

v

157

PSYCHOMYIIDAE

Ocelli absent. Maxillary palpi 5-segmented in both

sexes, segment 1 short, segment 2 usually longer than

segment 1 and about as long as segment 3; segment 5

long, annulate, flexible.

Mesoscutum with two rounded setal warts; scutellum

with a pair of separate or a single median setal wart.

Wings sLender, discoidaL ceLL reLativeLy Long, veins

R2 and R3 fused, in hindwing venation reduced. Female

abdomen often terminates in slender rigid ovipositor.


The family, with a total of approximately 15 genera

and 150 species, is represented in all faunal regions

except the Neotropical.The distribution in the Austra

lian region is restricted to New Guinea and New Zea

land with a single species in each.

Synopsis of genera

Tinodes Curtis, 1834 (1)

Zelandoptila Ti llyard, 1924 (1)

Zelomyia McFarlane, 1956

PSYCHOMYII DAE: TlNODES

aberrans KIMMINS, 1962 m; 3.5 mm; f; 3 - 4 mm

PSYCHOMYII DAE: ZELANDOPTILA

158

NG

I

I I

\ __ r

mose/yi TILLYARD, 1924 CD m: 3.6 - 4.5 mm; f: 4.4 - 4.8 mm

2:4:4-

1 :2:4-

NZ

159

CHATHAMIIDAE


sexes; in maLes the segment 2 is Long; segment 3 arises

before the apex of the second. Antennae stout, basaL

segment shorter than the head.

Mesoscutum mostLy with fine setate punctures,

arranged in two irreguLar bands; scuteLLum rounded an

teriorLy with a pair of indistinctLy fused setal warts.

Wing coupLing by enLarged, curved macrotrichia on fore

margin of hindwing, wing span about 15 to 22 mm; mid

tibia distinctLy Longer than femur. FemaLe abdomen

terminates with a Long, rigid ovipositor.

TibiaL spurs 2:2:4.

The Larvae inhabit intertidaL zone rock pooLs, fee

ding mainLy on caLcareous marine aLgae.

The famiLy is restricted to the AustraLian - New

ZeaLand region, where it is known by 2 genera and 4

species.

Synopsis of genera

Chathamia TiLLyard, 1925 (2)

Philanisus WaLker, 1852 (2)

Anomalostoma Brauer, 1865

160

CHATHAMIIDAE: CHATHAM/A 2:2:4-

brevipennis TILL YARD, 1925 (j') m & f: brachypterous CH

integripennis RIEK, 1977 m: 5 mm; f: 5.5 mm NZ n

CHATHAMIIDAE: PH/LAN/SUS 2:2:4-

p/ebeius WALKER, 1852 (j') m: 7 - 8 mm; f: 9 -10.5 mm NZ, AU se

/,.,. ------.......... .

fasciatus RIEK, 1977 m: 7 mm; f: 9 mm KR

......... " -........ .

161

TASIMIIDAE

The species in this famiLy are moderateLy smaLL to

medium size,dark, inconspicuous caddis-fLies with wing

span of 12 to 18 mm; their generaL appearance simiLar

to those of the famiLy Conoesucidae.

OceLLi absent. MaxiLLary paLpi 4-segmented in maLes

5-segmented in femaLes. Antennae sLightLy Longer than

forewing, basaL segment Longer than the head. Eyes co

vered with short setae.

Mesoscutum with a pair of rounded setaL warts, se

parated from the median Line; scuteLLum with a pair of

eLongate warts, which sometimes are fused. Wings rather

broad, rounded apicaLLy, forewing with anaL veins and

jugaL Lobe reduced.

TibiaL spurs 2:4:4.

The famiLy is known onLy from the AustraLian and

NeotropicaL regions with two genera in each, there are

onLy 9 described species, of those 7 are in the Austra

L ian region.

Synopsis of genera

Tasiagma Neboi ss, 1977 (2)

Tasimia MoseLy, 1936 (5)

Molanniella Banks, 1939

TASIMIIDAE: TASIMIA

pa/pata MOSELY, 1936 (!)

~.!":< "',--/ .,:'-.

. i ph D

162

m: 6.5 - 7.5 mm; f: 7.5 - 8.5 TA

denticu/ata JACQUEMART, 1965 m: 6 -7 mm; f: 7 -7.5 mm

~ ph

D

ph L

drepana NEBOISS, 1977 m:6mm TA

atra (BANKS, 1939) m: 5.5 - 6.5 mm AUse

2:4:4-

TA

163

TASIMIIDAE: TASIMIA 2:4:4-

natasia MOSELY, 1953 m:5mm AUse

TASIMIIDAE: TASIAGMA 2:4:4-

ciliata NEBOISS, 1977 (i) m: 7 - 7.5 mm; f: 7.5 - 8 mm AU se, TA

/. .

/' "8\,.0;;:',:' I "

'-----, , / , /

eremica NEBOISS, 1986 m: 5.8 - 6.2 mm LH

164

LIMNEPHILIDAE

To this famiLy beLongs some of the Largest Austra

Lian caddis-fLy species; the wing span is from 25 to

40 mm, the wings are broad, apicaLLy rounded, usuaLLy

yeLLowish-brown to brown without distinct coLour

pattern.

OceLLi present. MaxiLLary paLpi 3-segmented in maLe

5-segmented in femaLe, sLightLy pubescent; LabiaL paL

pi smaLL. Antennae as Long as, or sLightLy shorter

than forewing, basaL segment enLarged.

Mesoscutum with a pair of smaLL setaL warts (Aus

traLian species); scuteLLum with either a pair of se

parate or one continuous mesaL wart. Forewing with

discoidaL ceLL cLosed, median ceLL absent, anaL veins

fused; hindwing usuaLLy broader than forewing.

TibiaL spurs 1:2:2 (AustraLian species);tibiae and

tarsi usuaLLy armed with severaL rows of strong dark

spines.

Large famiLy wideLy distributed in the cooLer regi

ons of PaLearctic and Nearctic from which about 30 ge

nera with more than 1000 species are recorded. In the

AustraLian region onLy one genus with 3 species is

known.

Synopsis of genera


Archaeophylax Kimmins, 1953 (3)

165

L1MNEPHI L1DAE: ARCHAEOPHYLAX 1 :2:2+

ochreus MOSELY,1953(j) m: 15 - 18 mm; f: 16 -19 mm AU se, TA

v

vernalis NEBOISS, 1977 m: 14 mm TA

~ ph

canarus NEBOISS, 1962 m:11-12mm AUse

166

GOERIDAE

OceLLi absent. MaxiLLary paLpi 2 or 3-segmented in

maLes, sometimes with thickened or erectiLe Lobes heLd

in front of the face; in femaLes the maxiLLary paLpi

are 5-segmented, segments reguLar. The antennae with

basaL segment enLarged, usuaLLy Longer than the head.

Mesoscutum with or without a pair of smaLL ovaL or

eLongate setaL warts; scuteLLum with a Large median

wart.The forewing in maLes of some species with foLds

or grooves,discoidaL ceLL cLosed,cross-vein sc-r pre

sent; hindwing vein R1 reduced in some species; dis

coidaL ceLL open.

TibiaL spurs 2:4:4.

The famiLy has been recorded from aLL faunaL regi

ons, except the NeotropicaL and contains approximateLy

10 genera with 80 species. OnLy one genus with two

species are known from the SW Pacific.

Synopsis of genera

(OnLy one genus i~ the region)

Goera Stephens, 1829 (2)

167

GOERIDAE: GOERA 2:4:4-

fijiana BANKS, 1924 m: 6 mm; f: 6.5 mm FI

aneityuma NEBOISS, 1986 m: 6 - 6.5 mm; f: 7.5 - 8 mm NH (Aneityum)

/\( )) \

168

LEPIDOSTOMATIDAE

Ocelli absent. Maxillary palpi 1 to 3-segmented in

males, segments often swollen, covered with modified

hairs or scales; 5-segmented in females, segments

simple. Antennae with basal segment and sometimes also

the second segment much longer than the head, covered

with erect hairs or scales.

Mesoscutum and scutellum each with one pai r of seta l

warts. Wings elongate oval, densely pubescent, in males

often with folds or grooves lined with scales; in fe

males wing venation regular.

Tibial spurs 2:4:4.

The family is represented by about 30 genera and

250 species from almost all faunal regions, it is absent

from South America. In the SW Pacific - Australian re

gion it is restricted to western New Guinea and a few

nearby islands.

Synopsis of genera

Dinarthropsis Ulmer, 1913 (1)

Goerodes Ulmer, 1907 (1)

Neolepidostoma Ulmer, 1910 (1)

wing venation - generalised

169

LEPI DOSTOMATI DAE: GOERODES 2:4:4-

japenensis KIMMINS, 1962 m: B mm; f: B mm NG

LEPIDOSTOMATIDAE: DINA R THROPSIS 2:4:4-

picea ULMER,1913G) m:B.5-9mm JA, NG

LEPI DOSTOMA TI DAE: NEO L EPI DOS TOMA 2:4:4-

daabanum ULMER,1951 f:7mm NG (Kei I)

170

OECONESIDAE

The species of this famiLy are moderateLy Large to

Large with broad, brown mottLed wings and wing span of

30 - 40 mm. The wings in resting position are foLded

fLat over the body.

OceLLi absent. MaxiLLary paLpi 1 or 2-segmented in

maLes; 5-segmented in femaLes. Antennae about as Long

or sLightLy Longer than the forewing, basaL segment

shorter than the head.

Mesoscutum with a pair of eLongate setaL warts,scu

teLLum with a singLe mesaL wart. Wings are broad,

rounded apicaLLy, venation differing in sexes, often

reduced or modified in maLes; vein R1 joins R2 cLose

to wing margin; discoidaL ceLL cLosed.

TibiaL spurs 2:4:4.

The famiLy is restricted to Tasmania and New Zea

Land with the totaL of 6 genera and 16 described spe

cies; it has not been recorded from the AustraLian

mainLand.

OriginaLLy described by TiLLyard (1921) [101J as a

tribe of the famiLy Sericostomatidae it was raised to

famiLy by Neboiss (1975) [76J.

Synopsis of genera

Oeconesus McLachLan, 1862 (4)

Pseudoeconesus McLachLan, 1894 (7)

Tarapsyche McFarLane, 1960 (1)

Tascuna Neboiss, 1975 (1)

Zelandopsyche TiLLyard, 1921 (2)

Zepsyche McFarLane, 1960 (1)

171

OECONESIDAE: OECONESUS

maori McLACHLAN,1862(I)

cf

incisus MOSELY, 1953 Syn. (?) of maori McLachlan

lobatus WISE, 1958 Syn. (7) of maori McLachlan

similis MOSELY, 1953

m: 11 -14 mm; f: 13.5 -15 mm

m: 16 mm

m: 13 mm NZn

m: 14 -16 mm; f: 16 -18 mm

ph V

2:4:4-

NZ

1\ '\ ,'-'.".-~--~ . .'

D

NZ

172

OECONESIDAE: PSEUDOECONESUS 2:4:4-

stramineus McLACHLAN, 1894 CD m: 13 mm; f: 15.5 mm NZ

bistirpis WISE, 1958 m: 10 mm; f: 15 mm NZn

tristirpis WISE, 1958 m: 11 mm NZ n

squamosus MOSELY, 1953 m: 12.5 -16 mm

~tf. ~ NZ mxp

hudsoni MOSELY, 1953 m:12-14mm NZn

OECONESIDAE: PSEUDOECONESUS

mimus McLACHLAN,1894

karoriensis MOSELY, 1953 Syn. (7) of mimus Mclachlan

m:13-14mm;f:14mm

m: 14 mm NZn

173

2:4:4-

NZn

174

OECONESIDAE: TASCUNA 2:4:4-

ignota NEBOISS, 1975 (i) m: 16 -17 mm; f: 17.5 mm TA

I I ( I I \ '--

OECONESIDAE: TARAPSYCHE 2:4:4-

olis McFARLANE, 1960 (i) m:9-11.5mm;f:10-11 mm NZ

175

OECONESI DAE: ZELANDOPSYCHE 2:4:4-

ingens TILLYARD,1921(f) m: 19 mm NZs

maclellani McFARLANE,1981 m:18-19mm NZs

OECONESIDAE: ZEPSYCHE 2:4:4-

acinaces McFARLANE, 1960 (f) m:9-12mm;f:9-11 mm NZ

176

KOKIRIIDAE

Adults of this family are stocky, medium sized in

sects with wing span of 14 to 24 mm, dark brown or

blackish in colour.

Ocelli absent. Maxillary palpi 3-segmented in male;

5-segmented in female. Antennae stout, about as long

as forewing. Mouthparts more or less elongate, some

times in form of proboscis.

Mesoscutum and scutellum each with a pair of setal

warts. Forewing vein R1 joined to Sc basad of ptero

stigma by a short cross-vein or a sharp bend;R2 and R3

frequently fused; discoidal cell closed; in hindwing

the vein R1 incomplete, terminating before reaching

wing margin; discoidal cell open.


The family is restricted to the Australian and Neo

tropical regions with the total of 6 genera and 8 spe

cies, of which 5 genera and 7 species are from the

Australian region.

Synopsis of genera

Kokiria McFarlane, 1964 (1)

Mecynostomella Kimmins, 1953 (1)

Tanjistomella Neboiss, 1974 (1)

Taskiria Neboiss, 1977 (3)

Taskiropsyche Neboiss, 1977 (1)

177

KOKI RIIDAE: KOKIRIA 2:4:4-

miharo McFARLANE,1964(j) m: 7 - 8 mm; f: 7.5 mm

KOKIRIIDAE: MECYNOSTOMELLA 2:4:4-

fusca KIMMINS, 1953 (j) m:11 mm;f:13mm NC

178

KOKI RIIDAE: TASKIRIA 2:2:4-

austera NEBOISS, 1977 (j') m: 7 - 8 mm; 7.5 - 8.5 mm TA

mccubbini NEBOISS, 1977 m:9-9.5mm TA

otwayensis NEBOISS, 1984 m:11mm AUse

179

KOKI RIIDAE: TANJISTOMELLA 2:4:4-

verna NEBOISS, 1974(f) m: 9 mm; f: 9 - 9.5 mm AUse

.-"".

:'\ \1_,//'

KOKIRIIDAE: TASKIROPSYCHE 1:4:4-

lacustris NEBOISS, 1977 (f) m: 8.5 - 9.5 mm TA

180

PLECTROT ARSI DAE

AduLts of this famiLy are smaLL to medium size, the

wing span ranges from about 10 to 22 mm.They are found

flying during daytime in bright sunshine in a moth-L ike

manner around fLowering tea-tree (Leptospermum) and

other shrubs.

OceLLi present.MaxiLLary paLpi 3-segmented in maLe;

5-segmented in femaLe. Antennae stout, about as Long

as forewing. Mouth parts either normaL or forming an

eLongate proboscis in some species.

Mesoscutum with two pairs of indistinct setaL warts

or with two groups of set ate punctures, the posterior

pair very smaLL, situated LateraL to scuteLLum; setaL

warts on scuteLLum partiaLLy or fuLLy connected me

saLLy.

Wings broad, rounded apically and denseLy pubescent;

discoidaL ceLL cLosed, smaLL; at the base of hindwing

there are severaL strong frenuLar bristLes; vein R1

joins R2 near the wing margin.

TibiaL spurs variabLe 1-2:2-4:4.

The famiLy is restricted to AustraLia and Tasmania,

and contains 3 genera with 5 species.

Synopsis of genera

Liapota Neboiss, 1959 (1)

Tasmania Jacquemart, 1965

Nanoplectrus Neboiss, 1977 (1)

Plectrotarsus KoLenati, 1848 (3)

181

PLECTROTARSIDAE: PLECTROTARSUS 1:4:4+

gravenhorsti KOLENATI, 1848 G) m: 6.5 - 8.5 mm; f: 7.5 - 10 mm AU se, T A

tasmanicus MOSELY, 1936 m: 8 - 9.5 mm; f: 8 - 11 mm TA

minor MOSELY,1953 m:6-7mm;f:7mm AUsw

~yr';; ~,_\~::: ..

...... .: ........... :

182

PLECTROTARSIDAE: LlAPOTA 2:2:4+

lavara NEBOISS, 1959 (j) m: 5.5 - 6 mm; f: 5 - 6 mm TA

PLECTROTARSIDAE: NA NOPL ECTRUS 1 :2:4+

truchanasi NEBOISS, 1977 G) m:4mm TA

A~······"·· . . . , .'. . . - . ··.n

183

CONOESUCIDAE

AduLts of this fami Ly are smaLL to medium sized with

wing span of 10 to 25 mm, bLackish-brown to bLack,

usuaLLy found amongst vegetation on the banks of fast

fLowing smalL to medium size streams.

OceLLi absent. MaxiLLary paLpi 1 to 3-segmented in

maLes, often partLy or entireLy membranous, heLd uprigth in front of the face; in femaLes the maxiLLary

paLpi 5-segmented. Antennae stout, about as Long as,

or sLightLy shorter than the forewing, basaL segment

Long.

Pronotum with a pair of transverse, eLongate setaL

warts; mesoscutum without setaL warts, but with deep

anteromesaL fissure; scuteLLum with pair of setaL

warts. Both wings about equaL in width, eLongate ovoid;

forewing usuaLLy with hyaLine spot at arcuLus, jugaL

Lobe weLL deveLoped; discoidaL ceLL present.

TibiaL spurs 2:2:2-4.

This famiLy, previousLy part of the famiLy Sericos

tomatidae [77J, is restricted to the AustraL ian region,

where it is widespread in New ZeaLand, south-eastern

AustraL ia and Tasmania. There are 13 genera with 37 spe

cies.

Synopsis of genera

Beraeoptera MoseLy, 1953 (1)

Coenoria MoseLy, 1953 (1)

Confluens Wise, 1962 (2)

Conoesucus MoseLy, 1936 (6)

Conuxia McFarLane, 1966 (1)

Conia McFarLane, 1956

Costora MoseLy, 1936 (8)

Hampa MoseLy, 1953 (1 )

Lingora MoseLy, 1936 (4)

Matasia MoseLy, 1936 (1 )

Olinga McLachLan, 1894 (3)

Periwinkla McFarLane, 1973 (1)

Pycnocentria McLachLan, 1866 (5)

Pycnocentrodes TiLLyard, 1924 (3)

184

CONOESUCI DAE: COS TORA 2:2:4-

iena MOSELY, 1936 (i) m: 11 mm; f: 12 mm TA

krene NEBOISS,1977 m: 5.5 mm TA

luxata NEBOISS, 1977

seposita NEBOISS, 1977

,.I

/--- -------/\" I \

I \ ' ..... __ ./_-:-- _____ 1 __ _

m:5.5mm TA

m:7mm TA

ramosa JACQUEMART, 1965 m: 6-7 mm TA

185

CONOESUCI DAE: COS TO RA 2:2:4-

rotosca MOSELY, 1953 m: 7 - 8 mm; f: 7 - 8.5 mm TA

delara MOSELY, 1953 m: 7 - 7.5 mm; f: 8 - 9 mm AU 5e, TA

ebenina NEBOISS, 1977 m: 6 - 7.5 mm; f: 7 - 9 mm AU 5e, TA

- - - --'.~- ~ -./;;- ---, v

186

CONOESUCI DAE: CONOESUCUS 2:2:4-

norelus MOSELY,1953 m:5-6.5mm;f:6-7mm TA

semiauratus MOSELY, 1953 m:7mm AUse

fromus MOSELY, 1936 (j) m: 6 - 7.5 mm; f: 7 - 8 mm TA

, .',

" " "

v "'-" __ /'8',,.,,. ~" "

187

CONOESUCI DAE: CONOESUCUS 2:2:4-

digitiferus JACQUEMART, 1965 m: 7 - 7.5 mm; f: 8 - 9.5 mm TA

v

nepotulus NEBOISS, 1977 m: 5 - 6.5 mm; f: 6 - 7 mm TA

brontensis NEBOISS, 1977 m: 7 - 8 mm; f: 8.5 -10 mm TA

S'. ;,; .

,:<;:".y,.-~.~".,,,,,,,,, ~

188

CONOESUCI DAE: HAMPA 2:2:4-

patona MOSELY,1953(i) m:5-6'mm;f:6-7mm AU 5e, TA

CONOESUCI DAE: MATASIA 2:2:4-

satana MOSELY, 1936 (i) m: 5 - 6 mm; f: 5.5 - 7 mm TA

v

189

CONOESUCI DAE: LlNGORA 2:2:4-

aurata MOSELY,1936(i) m: 5 - 6.5 mm; f: 5.5 -7 mm TA

.~ •. =~ •............. . '.:::::';<:;" ......... : .. ,".:-:.::.:

". . . - ... -.... -- .

~

~~ v

vesca NEBOISS, 1977 m:5.5mm TA

,~ ~

::'.'."

coomata MOSELY, 1953 m:5.5mm AUse

plicata BANKS, 1939 m:4.5mm AUse

190

CONOESUCI DAE: PYCNOCENTRODES

aureola (McLACHLAN, 1868) (f) m: 6.5" 8 mm; f: 6.5" 10 mm

-modesta COWLEY, 1976 Syn. (n of aureola (McLachlan)

m: 6.3 mm; f: 6.5 mm

aeris WISE, 1958 m: 6.5" 10 mm; f: 6.5" 10 mm

ph

CONOESUCI DAE: COENORIA

boera MOSELY, 1953 (f) m: 4 mm AUse

NZn

NZ

2:2:4-

NZ

2:2:2-

191

CONOESUCIDAE: CONFLUENS 2:2:4-

hamiltoni (TILL YARD, 1924) (j) m: 9 - 9.5 mm; f: 11 mm NZ n

olingoides (TILL YARD, 1924) m: 8 - 9.5 mm; f: 9.5 - 10 mm NZ s

CONOESUCI DAE: BERAEOPTERA 2:2:4-

roria MOSELY, 1953 (j) m: 4.5 - 5 mm; f: 5.5 - 6.2 mm NZ

192

CONOESUCI DAE: PYCNOCENTRIA 2:2:4-

evecta McLACHLAN, 1868 m: 7 - 8 mm; f: 7.5 - 8.5 mm NZ

D

forcipata MOSELY, 1953 m: 5 mm; f: 5.5 mm NZ

sylvestris McFARLANE, 1973 m: 5.5 - 6 mm; f: 7 mm NZs

funerea McLACHLAN, 1866 (j) m: 6 - 7 mm; f: 6.5 - 7 mm NZ

193

CONOESUCI DAE: PYCNOCENTRIA

hawdonia McFARLANE, 1956 m: 4.5 - 5.5 mm; f: 4.5 - 5 mm

CONOESUCI DAE: CONUXIA

gunni (McFARLANE, 1956) (f)

.. ". "......... -~

m:5mm

. ' . .... ~ ", ...........

CONOESUCI DAE: PERIWINKLA

NZ

childi McFARLANE, 1973 (f) m: 5.5 - 6 mm; f: 7.6 mm NZs

NZs

I I I ! ........ }

o

2:2:4-

2:2:4-

2:2:4-

194

CONOESUCIDAE: OLINGA 2:2:4-

fereday; (McLACHLAN,1868)(!) m:9-10.5mm;f:10-11 mm NZ

... , . , ,

v \.: ""- -~'

jeanae McFARLANE, 1966 m: 10.5; f: 11.5 mm NZs

fumosa WISE, 1958 m:8mm NZ (s7) Syn. (?) of feredavi (McLachlan)

195

ANTIPODOECIIDAE

This is a monotypic famiLy, known from one smaLL,

dark coLoured species with wing span of 8 to 10 mm from

eastern AustraLia.

OceLLi absent. MaxiLLary paLpi 3-segmented in maLe,

curved and heLd upright in front of the face, apicaL

segment short; in femaLe the maxiLLary paLpi 5-segmen

ted. Antennae about as Long as the forewing, basaL

segment sLightLy shorter than the head, posterior se

taL warts Large, rounded, eLongate.

Mesoscutum setaL warts smaLL, ovaL, often indis-

tinct, or sometimes entireLy absent; scuteLLum with

one median, dome shaped wart. Wings sLender, poin

ted apicaLLy; discoidaL and median ceLLs absent; Large

hyaLine area in the centre of the wing at anastomosis;

basaL section of costaL margin of hindwing with sparse

Long bri stLes.

TibiaL spurs 2:2:4, the midLeg with a row of short,

dark spines on femur, tibia and tarsus.

Synopsis of genera


Antipodoecia MoseLy, 1934 (1)

196

ANTIPODOECIIDAE: ANTIPODOECIA 2:2:4-

turneri MOSELY, 1934 (j) m: 3.5 - 4 mm; f: 4 - 4.5 mm AUse

197

CALOCIDAE

Most of the species in this famiLy are medium size

with wing span of 12 to 25 mm, but a few are moderate

Ly Large, reaching wing span of approximateLy 30 mm;

forewings bLackish, usuaLLy mottLed with whitish irre

guLar spots. The species occur aLong smaLL cooL-water

streams.

OceLLi absent. MaxiLLary paLpi 3 to 5-segmented in

maLes, frequentLy modified; 5-segmented in femaLes.

Antennae stout, about as Long or sLightLy shorter than

forewing; basaL segment Long, in maLes often with ex

panded Lobes or other modifications. In some genera

head with expandabLe membranous fiLaments dorsomesaLLy

and Long posterior setaL warts; in femaLes posterior

setaL warts shorter.

Pronotum usuaLLy with two pairs of setaL warts, the

mesaL pair smaLL, rounded, the LateraL ones eLongate,

(except in maLes of Caenota, which have onLy one pair

of eLongate warts, simiLar to those in Conoesucidae).

Mesoscutum without setaL warts; scuteLLum with a

pair of eLongate warts. Forewings eLongate, ovaL, dis

coidaL ceLL present, cLosed, narrow hyaLine area above

thyridiaL ceLL and surrounding cross-vein r-m; hindwing

vein R1 usuaLLy joins Sc for a short distance before

ending separateLy at wing margin.

TibiaL spurs 2:2:4.

The famiLy is known onLy from the AustraLian region

and incLudes the New ZeaLand famiLy PycnocentreLLidae

[77]; at present a totaL of 7 genera with 20 species

are known.

Synopsis of genera

Alloecentrella Wise, 1958 (1)

Caenota MoseLy, 1953 (5)

Caloca MoseLy, 1953 (6)

Tismana MoseLy, 1953

Calocoides Neboiss, 1984

Pliocaloca Neboiss, 1984

Pycnocentrella MoseLy, 1953

Tamasia MoseLy, 1936 (3)

(1)

(3)

(1 )

198

CALOCI DAE: CALOCA 2:2:4-

straminea MOSELY, 1953 (f) m( 6.5 mm; f: 6.5 -7 mm AU

ascita NEBOISS, 1977 m:6.5mm TA

tertia MOSELY, 1953 m:8mm TA

fallia MOSELY, 1953 m:6.Smm AUse

199

CALOCI DAE: CALOCA 2:2:4-

saneva (MOSELY, 1953) m: 7" 7.5 mm; f: 7.5 mm TA

eba MOSELY, 1953 m:5mm AUse

CALOCI DAE: PYCNOCENTRELLA 2:2:4-

eruensis MOSELY, 1953 @ m:8.5-10mm;f:9-11 mm NZ

200

CALOCI DAE: TAMASIA 2:2:4-

variegata MOSELY, 1936 (i) m: 6.5 - 8 mm; f: 7.5 - 9 mm AU se, TA

acuta NEBOISS, 1984 m: 6.5 - 8 mm; f: 7.5 - 8.5 mm AUse

furcilla NEBOISS, 1984 m: 7 - 7.5 mm; f: 8 mm AUse

201

CALOCI DAE: CALOCOIDES 2:2:4-

aquilonia NEBOISS, 1984 (j) m: 5.5 - 6 mm; f: 6 - 7 mm AU ne

CALOCI DAE: ALLOECENTRELLA 2:2:4-

magnicornis WISE, 1958 (j) m: 5 - 6 mm; f: 6.5 mm NZ

., . ' .. -.. .- .................. /

CALOCI DAE: PLIOCALOCA

mucronata NEBOISS, 1984 CD m: 6.5 -7 mm

dasodes NEBOISS, 1984 m: 6 - 6.5 mm; f: 7.5 mm

fastigiata NEBOISS, 1984 m: 6 mm; f: 6.5 mm

mucronata NEBOISS, 1984 6.5 - 7 mm

202

AU ne

AU ne

I

~ '~ "

: :

~~ ~~

dasodes NEBOISS, 1984 6 - 6.5 mm

2:2:4-

203

CALOCIDAE: CAENOTA 2:2:4-

plicata MOSELY, 1953(i) m:11-13mm;f:12-14mm AUse, TA

simu/ans MOSELY, 1953 m: 14 mm AU ne

nemorosa NEBOISS, 1984 m: 7 - 8.5 mm; f: 8 - 8.5 mm AUne

204

CALOCIDAE: CAENOTA 2:2:4-

monteithi NEBOISS, 1984 m: 8.5 - 11 mm; f: 9 - 11 mm AU ne

v galeata NEBOISS, 1984 m: 7 - 7.5 mm; f: 7 - 8 mm AU ne

.... ----~ :..~ -~--J --,,-\-. -.. ---' I~ \: \ ~'

I~ .' I, \\. __ J

205

HELICOPHIDAE

The aduLts of this famiLy are smaLL to medium size,

inconspicuous, dark greyish species with wing span of

8 to 15 mm; known to inhabit cLear, swift fLowing

streams in forested areas.


sexes. Antennae not exceeding the Length of forewing,

basaL segment about as Long, or sLightLy shorter than

the head. Large posteroLateraL setaL warts on the head.

Pronotum with one pair of eLongate setaL warts;

mesoscutum without setaL warts; scuteLLum with one

or a pair of smaLL setaL warts. Forewing eLongate ovaL,

with discoidaL ceLL either cLosed or open,hyaLine area

at vein junction just beLow discoidaL ceLL, jugaL Lobe

reduced, venation often differing in sexes; hindwing

with a row of hamuLi aLong the basaL section of costaL

margin, Large, vein-free area discaLLy.

TibiaL spurs 2:2:4.

The distribution of the famiLy is Limited to Aus

traLia and New ZeaLand, where a totaL of 3 genera and

8 species are known.

Synopsis of genera

Alloecella Banks, 1939 (3)

Helicopha MoseLy, 1953 (3)

Zelolessica McFarLane, 1956 (2)

~Li,"Ph'

ALLoeceLLa

206

HELICOPHIDAE: ALLOECELLA 2:2:4-

grisea BANKS, 1939(i) m: 5.5 - 6 mm; f: 6 - 6.5 mm AU se, TA

v

/ongispina JACQUEMART, 1965 m: 6 - 7 mm; f: 6.5 - 7.5 mm TA

pi/osa NEBOISS, 1977 m: 5 - 6 mm; f: 5.5 - 6.5 mm TA

207

HELICOPHIDAE:. HELICOPHA 2:2:4-

astia MOSELY, 1953 (i) m: 6 - 7 mm; f: 6 - 7.5 mm TA

~m ~ .. vag

v

\

delamarei JACQUEMART, 1965 m: 4.5 - 6.5 mm; f: 6 - 7 mm TA

v

hortena MOSELY, 1953 m:5.5 mm AU

208

HE LlCOPHI DAE: ZELOLESSICA 2:2:4-

cheira McFARLANE, 1956(i) m: 3 - 4.5 mm; f: 4 - 4.7 mm NZ

meizon McFARLANE,1981 m: 5 - 5.5 mm; f: 5 - 6 mm NZ S, ST

209

ODONTOCERIDAE

The famiLy in this region is represented by medium

size caddis-fLies with wing span of 14 to 25 mm; some

robust, bLackish species with stout antennae, others

are sLender, brownish species with Long, fiLiform an

tennae and distinctLy enLarged eyes in the maLes.


sexes, segments Long, cyLindricaL. Antennae are as Long

as or considerabLy Longer than forewing, basaL segment

Large. In maLes of some species the eyes are much en

Larged and aLmost meeting mesaLly.

Mesoscutum either with a pair of smaLL, ovaL scutaL

warts or with two indistinct LongitudinaL bands or

groups of setiferous punctures; scuteLLum dome-shaped,

rounded anteriorLy, a singLe setaL wart mesaLLy, some

times indistinctLy outLined. Forewings moderateLy broad

to sLender; discoidaL ceLL cLosed, median ceLL absent;

veins R1 and R2 frequentLy joined or connected by

cross-vein near wing margin; venation sometimes irre

guLar in maLes, differing from females.

TibiaL spurs 2:4:4 (AustraLian species).

The famiLy has a scattered distribution throughout

aLL faunaL regions except the Ethiopian; there are 10

genera with about 70 species recorded so far; of those

2 genera and 4 species are recorded from AustraLia,but

there are no records from other SW Pacific LocaLities.

Barynema

Marilia

Synopsis of

Banks, 1939

MuLLer, 1878

genera

(2)

(2)

"" - -- -

210

ODONTOCERI DAE: BARYNEMA 2:4:4-

australicum MOSELY,1953 m: 7.5 mm AUse

costa tum BANKS, 1939G) m: 9.5 -10 mm; f: 9.5 mm AUse

v

ODONTOCERIDAE: MARiLlA 2:4:4-

bola MOSELY, 1953 m: 8.5 mm; f: 7.5 - 8.5 mm AUse

v

fusca KIMMINS, 1953 m: 7 mm; f: 6 -7 mm AUse

211

ATRIPLECTIDIDAE

The aduLts of this famiLy are medium size,with wing

span of 20 to 28 mm, sLender buiLd, brown with subdued

mottLed appearance and Long, sLender antennae.

OceLLi absent. MaxiLLary paLpi stout, 5-segmented

and simiLar in sexes. Antennae Longer than forewing,

basaL segment short, buLbous.

ProsternaL setaL warts transverseLy eLongate; meso

scutum with two LongitudinaL bands of setiferous punc

tures; scuteLLum with a singLe mesaL wart. Forewings

narrow, discoidaL ceLL present, wing coupLing with ha

muLi aLong the anterior margin of hindwing.

TibiaL spurs 2:4:4.

The distribution of this famiLy is restricted to

AustraLia and the SeycheLLes IsLands, comprising 2 ge

nera with a singLe species each. In AustraLia the spe

cies is wideLy distributed in the SE coastaL regions

and in Tasmania, but distribution in Western AustraLia

is restricted to a smaLL coastaL area south-west of the

State.

Synopsis of genera


Atriplectides MoseLy, 1936 (1)

212

ATRIPLECTIDIDAE: ATRIPLECTIDES 2:4:4-

dubius MOSELY, 1936 ([) m: 10 -12.5 mm; f: 12 - 13 mm AU se,sw, TA

o~=======~

D

213

PHiLORHEITHRIDAE

The famiLy contains moderateLy Large caddis-fLies

with wing span of 16 to 35 mm, and irreguLarLy varie

gated wings.

OceLLi absent. MaxiLLary paLpi 3 to 5 segmented in

maLes; 5-segmented in femaLes; basaL segment short with

apicomesaL expansion bearing a tuft of setae. In most

genera maLes with a pair of piLifers arising above the

base of maxiLLary paLpi and heLd upright in front of

the face. Antennae stout, about as Long as forewing;

10 or 15 basaL segments often excavated LateraLLy, bea

ring setae or pectinate.

Mesonotum usuaLLy with setaL warts, or if there are

no warts, then one or severaL setate punctures are

present in their pLace; scuteLLum usuaLLy with a pair

of rounded setaL warts. Wings eLongate, rounded or ob

LiqueLy truncate apicaLLy, discoidaL ceLL cLosed in

both wing pairs; in forewing R1 joins R2 near the api

caL margin.

TibiaL spurs 2:4:4.

The famiLy is restricted to the AustraLian and Neo

tropicaL regions. A totaL of about 9 genera with 20

species are known, and of those 6 genera with 15 spe

cies occur in AustraLia and New ZeaLand; the famiLy is

not recorded from other parts of the SW Pacific region.

Synopsis of genera

Aphilorhei thrus MoseLy, 1936 (4)

Austrheithrus MoseLy, 1953 (3)

Kosrheithrus MoseLy, 1953 (3)

Philorheithrus Hare, 1910 (2)

Ramiheithrus Neboiss, 1974 (2)

Tasmanthrus MoseLy, 1936 (1)

Tasmanthrus Austrheithrus

~I~t

Kosrheithrus AphiLorheithrus

214

PHI LORHEITHRIDAE: PHILORHEITHRUS 2:4:4-

agilis (HUDSON,1904)(i) m:11-13mm;f:14-15mm NZ

o

lacustris TILLYARD,1924 m:12-14mm;f:14-15.5mm NZs

PHI LORHEITHRI DAE: TASMA NTHR US 2:4:4-

angustipennis MOSELY,1936(i) m:9-12mm;f:11-14mm TA

(t.·o •• ' • ·0 ......---------~ .•... • ..... .

.y-0 .... /[ /~ . : 0.· •. .0.

'. . . . . .. 0

215

PHI LORHEITHRIDAE: A USTRHEI THRUS 2:4:4-

dubitans MOSELY, 1953 (i) m: 11 mm AUse

g/ymma NEBOISS, 1977 m: 10 -12 mm; f: 11 -13 mm TA

ronewa MOSELY, 1953 m: 10 -12 mm; f: 12 -14 mm AU se, TA

216

PHI LORHEITHRIDAE: RAMIHEITHRUS 2:4:4-

virgatus NEBOISS, 1974 (j) m:10-11mm AUse

kacinus NEBOISS, 1974 m:9mm TA

. . J ..... : .. (\()\"/;i> \ ,1 .•

Il ',. . : . .

.. rd v

PHI LORHEITHRIDAE: APHILORHEITHRUS 2:4:4-

lutealus NEBOISS, 1977 m: 13 -13.5 mm; f: 15.5 mm TA

D

217

PHI LORHEITH R I DAE: APHILORHEITHRUS 2:4:4-

stepheni MOSELY, 1936(i) m: 12-14mm;f: 13-17mm AUse, TA

~ ......... J cf~ ••.

\~,-----

v

decoratus NEBOISS, 1977 m: 10 -12 mm; f: 13 -15 mm TA

pauxillus NEBOISS, 1977 m: 7.5-8 mm TA

218

PHI LORHEITHR I DAE: KOSRHEITHRUS 2:4:4-

til/yardi MOSELY,1953(i) m:15mm;f:19-21 mm AUse

remu/us NEBOISS, 1977 m: 10.5 -12.5 mm TA

boorarus NEBOISS, 1982 m: 14 mm AUsw

, t~ , ,---_ ..

219

HELICOPSYCHIDAE

The members of this family mostly are small, grey

ish-brown to black, with rather slender wings and wing

span of 10 to 16 mm. The integument of at least fi rst few

abdominal sternites has reticulate pattern.

Ocelli absent. Maxillary palpi 2 or 3-segmented in

males; 5-segmented in females. Head with large, ovoid,

posterior setaL warts.

Mesoscutum and scutellum each with a pair of small,

rounded setal warts. Wings narrow, wing coupling by

row of hamuli; discoidal cell closed in forewing, open

in hindwing.

Tibial spurs 2:2:4 (species from Australian regi

on).

The larvae construct helical, snail shell like tube

cases of sandgrains.

The fami ly is known from all faunal regions with 6

genera and approximately 100 species; 2 genera with 11

species are recorded from the Australian region, how

ever, more species are known to exist in the area.

Synopsis of genera

Helicopsyche Siebold, 1856 (10)

Saetotricha Brauer, 1865

Rakiura McFarlane, 1973 (1)

220

HELICOPSYCHIDAE: HELICOPSYCHE 2:2:4-

zealandica HUDSON,1904 m: 4.5 - 5.5 mm; f: 5 - 6.5 mm NZ

albescens TILLYARD,1924 m: 4 - 4.5 mm; f: 4.5 - 5.5 mm NZ

howesi TILLYARD,1924 m:5.5mm NZs

poutini McFARLANE,1964 m: 4.5 mm; f: 6 mm NZ

221


murrumba MOSELY, 1953 m: 5 - 6 mm; f: 6 - 7 mm AU se, TA

cochleaetesta KORBOOT, 1964 m: 4.5 - 5 mm; f: 4.5 - 6 mm AU ne

bartona MOSELY, 1953 m: 5-6mm TA

tillyardi MOSELY, 1953 AUse

1//

222


heacota MOSELY, 1953 m:5mm AUse

ptychopteryx (BRAUER, 1865) m: 4.5 - 5 mm; f: 5.5 - 6 mm AUse

I

v

HELICOPSYCHIDAE: RAKIURA 2:2:4-

vernale McFARLANE, 1973 G) m: 5 - 5.5 mm; f: 7.3 mm

223

CALAMOCERATIDAE

The adults of this family are medium sized insects

with wing span of 15 to 28 mm, forewings rather broad,

somewhat triangular, densely covered with short, some

times distinctly coloured pubescence.

Ocelli absent. Maxillary palpi 6-segmented in both

sexes (genus Anisocentropus), long and densely covered

with long erect hairs, apical segment simple. Antennae

filiform, longer than forewing, basal segment shorter

than the head, subsequent segments slender.

Mesoscutum without setal warts, but with two, more

or less distinct irregular bands of setiferous punc-

tures, mesal separation of which is often indistinct;

scutellum small,without setal warts. Forewings broadly

triangular, discoidal and median cells both present;

venation usually similar in sexes; there is an elon

gate, hyaline line between discoidal and median cells.

Tibial spurs 2:4:3 (genus Anisocentropus).

The family, although represented in all faunal re

gions, is more common in the subtropical zone; it is

known by 6 genera and about 100 species, of which only

one genus with 23 species is recorded fron the Austra

lian region.

Synopsis of genera

(Only one genus in the region)

Anisocentropus McLachlan, 1863 (23)

224

CALAMOCERATIDAE: ANISOCENTROPUS 2:4:3~

dilucidus McLACHLAN, 1863 m: 7 - 9 mm; f: 8 -10 mm NG

iIIustris McLACHLAN,1863(j) m:11-13mm NG

---=~-.:' ...

fulgidus NAVAS, 1933 m: 13.3 mm NG

Syn. (?) of iIIustris McLachlan

maclachlani ULMER,1929 m: 9.5 mm; f: 9 - 10 mm NG

D

225

CALAMOCERATIDAE: ANISOCENTROPUS 2:4:3-

triangulatus ULMER, 1907 m: 9-10 mm NG (Fergusson Is.)

banghaasi ULMER, 1909 m: 8.5 -10 mm; f: 9 -10.5 mm NG, AU n,ne

tristis ULMER, 1929 m:8mm NG (Ambon Is.)

D

cretosus McLACHLAN, 1875 m: 7.5 - 9 mm; f: 8 - 9.5 mm SL, NG

226

CALAMOCERATI DAE: AN/SOCENTROPUS 2:4:3-

corvin us NEBOISS, 1980 m: 7.5 - 9.5 mm; f: 8 -10.5 mm AUnw

/atifascia (WALKER,1852) m: 8.5 -10.5 mm; f: 9 -12 mm AU ne,se, TA

muricatus NEBOISS, 1980 m: 7 - 7.5 mm; f: 7.5 - 8 mm AU n

vag.

ph

va/gus NEBOISS, 1980 m: 9 - 10.5 mm; f: 9.5 - 11.5 mm AUse

1/

ph

227

CALAMOCERATIDAE: A NISOCEN TROPUS

bicoloratus (MARTYNOV, 1914) m: 7.5 -10.5 mm; f: 7.5 -10.5 mm

• ____ --=-'-::"~~;1 ph

v

eungel/us NEBOISS, 1980 m:9mm AU ne

torulus NEBOISS, 1980 m: 7.5 - 8.5 mm; f: 7.5 - 8.5 mm AU ne

AU ne,se

" ,. r-/\"-)

..-'---.:

2:4:3-

228

CALAMOCERATIDAE: A NISOCENTROPUS

kirramus NEBOISS, 1980 m: 9 -10.5 mm; f: 9 -10.5 mm

pictilis

... -.,": . .~-

NEBOISS, 1986

io KIMMINS, 1962

semiflavus BANKS, 1939

m: 10.5 mm NG

m: 10 mm NG

m: 7.5 - 9 mm; f: 7.5 - 9 mm

2:4:3-

AUne

vag.

AU ne

229

CALAMOCERATIDAE: A NISOCEN TR OP US

hyboma NEBOISS, 1986

fijianus BANKS, 1936

s%monis BANKS, 1939

immunis McLACHLAN, 1863

yellow

antenna of immunis type

m: 8 - 8.5 mm; f: 8.7 mm NG

m:7mm;f:7mm FI

m: 8 mm Solomon Is.

(sex unknown; abdomen lost) 9.5 mm NG

Figure of LikeLy _immunis male specimen

These figures published by Ulmer 1907 [107] as immunis (from Japan) represent another species

2:4:3-

230

LEPTOCERIDAE

Members of this family are easily recognized by

their long, slender wings and long filiform antennae.

The size of species varies from moderately small to

rather large with wing span ranging from 10 to 40 mm.

A number of species are active during daytime and can

be seen flying along waters edge among vegetation.

Ocelli absent. Maxillary palpi long, 5-segmented;

the apical segment incompletely sclerotized, flexible

but not annulate; segment 2 the longest. Antennae usu

ally two or three times longer than the forewing, in

males longer than in females; basal segment bulbous,

rarely slender and longer than the head.

Mesoscutum elongate, without setal warts, but in

stead, with two longitudinal bands of setiferous punc

tures; scutellum small with few setiferous punctu

res. Forewings are long and narrow, discoidal cell

present, closed; median cell absent; hindwings are

either narrow and acute apically, or considerably ex

panded, sometimes with long fringe.

Tibial spurs 0-2:2:2-4.

The family is very common and known from all faunal

regions; there are about SO genera with more than 800

species which are placed in two subfamilies [SO]: Tri-

plectidinae Ulmer and Leptocerinae Leach. At present

18 genera and 135 species are recorded from the SW Pa

cific - Australian region.

Synopsis of genera

TRIPLECTIDINAE

Condocerus Neboi ss, 1977 (2)

Hudsonema Mosely, 1936 (2)

Lectrides Mosely, 1953 (2)

Notalina Mosely, 1936 (11 )

Notoperata Neboiss, 1977 (5)

Triplexina Jacquemart, 1965

Symphitoneuria Ulmer, 1906 (5)

Loticana Mosely, 1936

Symphitoneurina Schmid, 1950 (1)

Triplectides Kolenati, 1859 (33)

Notanatolica McLachlan, 1866

Triplectidina Mosely, 1936 (2)

Triplexa Mosely, 1953 (2)

Gracilipsodes Sykora, 1967

Westriplectes Neboiss, 1977 (3)

TRIPLECTIDINAE: hidwing discoidal cell closed

LEPTOCERINAE: hindwing discoidal cell open

LEPTOCERINAE

Adicella McLachlan, 1877 (1)

Leptocerus Leach, 1815 (5)

Leptorussa Mosely, 1953 (1)

Oecetinella Ulmer, 1907 (2)

Oecetis McLachlan, 1877 (33)

Oecetina Banks, 1899

Setodes Rambur, 1842 (4)

Triaenodes McLachlan J 1365 (21 )

231

LEPTOCERIDAE: WESTRIPLECTES 2:2:4-

pedderensis NEBOISS, 1977 (i) m: 14 -15 mm; f: 10 -12.5 mm TA

albanus (MOSELY, 1953) m: 10.5 mm AUsw

angelae NEBOISS, 1978 m: 10.5 mm AU ne

232

LEPTOCERIDAE: TRIPLECTIDES 2:2:2-

australis NAVAS,1934 m:11-14mm;f:12-14mm AU

ph

helvolus MORSE & NEBOISS, 1982 m:11-13mm;f:12-14.5mm AU nW,ne

e~ ph

pal/idus (BANKS, 1936) m: 13 mm FI

233


magnus (WALKER, 1852) m: 15 -18 mm; f: 17 -18 mm AU se, TA

~---------'.' ...•..... , ,~/ ~---'--'- . . . .:""'''--,''

ph

valda MOSELY, 1953 m: 10 - 12 mm; f: 10 - 12 mm AU ne,se

~---.---- •• =-.". ':,,,,., ....... / ..

~ ph

'c\/;~ , \

~, .' ///':-". j ) ·····'''9··· " !

. ,

parvus (BANKS, 1939) m: 10 -13 mm; f: 10 -13 mm AU n, ne

~--"" ........ ;;,',

ph

234


similis MOSELY,1953 m: 14 -18 mm; f: 14 -18 mm AU ne,se, TA

'-

ciuskus ciuskus MOSELY, 1953 m: 12 -14 mm; f: 12.5 -15 mm AU,TA

ciuskus seductus MORSE & NEBOISS, 1982 m: 10- 12 mm; f: 11 - 12.5 mm AUnw

australicus BANKS, 1939 m: 11.5 - 13 mm; f: 12.5 - 16 mm AU nW,ne,se 2:2:2/2:2:4-

ina V

/;-~ f,,/ ph

235

LEPTOCERI DAE: TRIPLECTIDES 2:2:4-

bilobus NEBOISS,1977 m: 15 -18 mm; f: 15 -18 mm TA

proximus NEBOISS, 1977 m: 13 -16 mm; f: 14 -16 mm AU se, TA

236

LEPTOCE R I DAE: TRIPLECTI DES 2:2:4-

niveipennis MOSELY, 1953 m: 12-14mm

enthesis NEBOISS, 1982 m: 13.5 mm; f: 13 -14 mm AUsw

~ ph

dolabratus MORSE & NEBOISS, 1982 m:11-13mm AU ne

237


prolatus MORSE & NEBOISS, 1982 m: 10 -11 mm AUne

liratus MORSE & NEBOISS, 1982 m: 9 -10 mm; f: 9 -10 mm

c· , ~{??!> ph

liratellus MORSE & NEBOISS, 1982 m: 9 - 9.5 mm AUne

elongatus BANKS, 1939 m: 12 -15 mm; f: 14 -16 mm AU se, TA

c------<~ ph

238

LEPTOCER IDAE: TRIPLECTIDES

varius KIMMINS,1953 m: 15 -16 mm

ph

truncatus NEBOISS, 1977 m: 10- 12 mm; f: 11 - 13 mm

tambina MOSELY, 1953 m: 11 - 12 mm; f: 12 - 17 mm

, I

I I

I , I

,/

AU se, TA

,

AU ne,se

, , , ,

2:2:4-

LEPTOCERIDAE: TRIPLECTIDES

insperatus MORSE & NEBOISS, 1982

-:~ .• C- ........... . •...........

ph

gonetalus MORSE & NEBOISS, 1982

pft' "':~ ~--__ ~I ph

,. , . : ~~~;''N

," .)"

altenogus MORSE & NEBOISS, 1982

239

2:2:4-

m: 10 mm AUne

m: 16 -18 mm; f: 17 -18 mm AUne

m: 15 -16 mm; f: 15.5 -16 mm AU ne,se

240


hamatus MORSE & NEBOISS, 1982 m: 12 mm AUse

ross; MORSE & NEBOISS, 1982 m: 10 -12 mm; f: 12 - 13.5 mm

~, ..........•

. , ...... -..•... ...:

ph o

241

LEPTOCE R IDAE: TRIPLECTI DES 2:2:2-

gilolensis (McLACHLAN, 1866) m:8-10mm JA, GI, NG

v

cepahlotes (WALKER, 1852) m: 14 -16 mm; f: 13.5 -15 mm NZ, ST,NO

v

cognatus (McLACHLAN,1862) m:14-15mm NZ

242


obsoleta McLACHLAN,1862 m: 12.5 -14.5 mm; f: 15 -17.5 mm NZ

dolichos McFARLANE, 1981 m: 13 - 17 mm; f: 17 - 19 mm NZ, ST

latipennis MOSELY, 1936 m: 11.5 mm; f: 10 mm NH 2:2:3-

243

LEPTOCERIDAE: TRIPLECTIDINA 2:2:4-

oreolimnetes (TILLYARD, 1924) (!) m: 9.5 -12 mm; f: 10 -12.5 mm AU S9, TA

nigricornis MOSELY, 1936 m: 9 - 10 mm; f: 9.5 - 11 mm NZs

244

LEPTOCERIDAE: LECTRIDES 2:2:4-

varians MOSELY,1953(!) m: 9 -12.5 mm; f: 8 -12 mm AUse, TA

!i¥#% .. -::.~

parilis NEBOISS, 1982 m: 10 - 11 mm; f: 10 - 11 .5 mm AUsw

245

LEPTOCE R I DAE: SYMPHITONEURIA 2:2:2-

opposita (WALKER, 1852) (j') m:11-12mm;f:10.5-11mm AUse,TA

exigua (McLACHLAN, 1862) m:8-10mm;f:8-10mm AUne

amp/a KORBOOT, 1964 m:15mm;f:15-16mm NG

246

LEPTOCERIDAE: SYMPHITONEURIA 2:2:2-

wheeleri BANKS, 1939 m: 10 - 11 mm; f: 11 - 11.5 mm AUsw

licmetica NEBOISS, 1986 m:11 mm;f:10mm NC

LEPTOCE RI DAE: SYMPHITONEURINA 2:2:2-

fulva (NAVAS, 1932)(j) m: 8-9.5 mm; NG

247

LEPTOCE R I DAE: NO TOPERA TA 2:2:4-

maculata (MOSELY, 1953) m:12-15mm;f:11-14mm AU ne,se, TA

sparsa (KIMMINS, 1953)(!) m:8-10mm;f:8-9mm TA

syncope NEBOISS,1982 m: 8 - 9 mm; f: 8 - 8.5 mm AUsw

tenax NEBOISS,1982 m: 9.5 - 11 mm AUsw

248

LEPTOCERIDAE: NOTOPERATA 2:2:4-

diversa NEBOISS, 1982 m: 9 - 9.5 mm; f: 8.5 - 9.5 mm AUsw

LEPTOCERIDAE: HUDSONEMA 2:2:4-

amabilis (McLACHLAN, 1868) (!) m:11-12mm;f:11-12.Smm NZ,ST

aliena (McLACHLAN, 1868) m:11-14mm;f:10.Smm NZ, ST

LEPTOCERIDAE: NOTALINA

mose/yi KIMMINS, 1953 m: 9 mm; f: 6 - 7 mm

parkeri MOSELY, 1936 CD

nigra (MOSELY, 1953)

.........

flava (ULMER, 1908)

m:11-13mm;f:9-11 mm

m: 7.5 - 9 mm; f: 7 - 8 mm

v

m: 8 - 9 mm; f: 7.5 - 8..5 mm

249

2:2:4-

AUse

TA

:: .. . : ..... :i·-·:· : .... ."'

TA

~ ~

r! ," ..

250

LEPTOCERIDAE: NOTALINA 2:2:4-

fulva KIMMINS,1953 m:9-15mm;f:7-11 mm AU sW,se, TA

~ if ..

)

til/yardi KIMMINS, 1953 m:12-13mm;f:10-11 mm TA

bifaria NEBOISS, 1977 m:12-13mm TA

251


delicatu/a (ULMER. 1908) m: 7 - 7.5 mm; f: 5 - 6 mm AUsw

~~«~ .. S?2<~~i&~wi~·_ if .

v

pseudodelicata NEBOISS. 1982 m: 7.5 - 8 mm; f: 5.5 - 6.5 mm AUsw

dwe/linga NEBOISS. 1982

•....• m9.9.5mm;fB ••. 5mm ~ :~

if

252


gungarra NEBOISS, 1984 m: 11.5 - 12 mm AUse

253

LEPTOCERIDAE: TRIPLEXA

villa MOSELY, 1953 (j) m: 9.5 mm; f: 8 mm AUne 2:2:2-

psocopterus (SYKORA, 1967) m: 5.5 - 6.5 mm NC 0:1:1-

254

LEPTOCERIDAE: CONDOCERUS 2:2:4-

paludosus NEBOISS, 1977 (j) m:10-11 mm;f:9.5-11 mm AU se, TA

,~£5kfP 9

D

aptus NEBOISS, 1982 m: 10.5 -11 mm; f: 10.5 -12 mm AUsw

LEPTOCERIDAE: LEPTORUSSA 2:2:4-

darling toni (BANKS, 1939) (j) m: 8 - 9 mm; f: 8.5 mm

255

LEPTOCERIDAE: LEPTOCERUS 0:2:2-

souta MOSELY, 1953 m: 8 - 9 mm; f: 8.5 mm AU se

ph

atsou SCHMID,1986 m: 4 - 5 mm; f: 4 - 4.5 mm AU n,ne

ousta SCHMID,1986 m:4mm AU ne

256

LEPTOCERIDAE: LEPTOCERUS 0:2:2-

cheesmanae KIMMINS, 1962 m:5.5mm NG

assimu/ans (ULMER, 1916) m:5mm AU

LEPTOCERIDAE: AD/CELLA 1:2:2-

pu/cherrima ULMER, 1906 m: 6.5 " 7 mm; f: 6.5 mm JA, SM, NG

257

LEPTOCERIDAE: TRIAENODES 1:2:2-

costalis KIMMINS, 1962 m:8mm

nigrolineata KIMMINS, 1962 m: 6 mm NG

excisa KIMMINS, 1957 m: 4.5 mm GU

picea KIMMINS. 1957 m: 5 mm GU

/anceo/ata KIMMINS, 1957 m:8.2mm GU

258


ustulata KIMMINS, 1962 m: 6 mm; f: 5 - 6 mm NG

tafana KIMMINS, 1962 m:6.5mm NG

trifida KIMMINS, 1957 m:3.2mm GU

longispina KIMMINS. 1962 m:8mm;f:7mm NG

259


aurea KIMMINS, 1962 m:8.5mm

telefominicus KUMANSKI, 1979 m:7.5mm NG

coral/ina KIMMINS, 1962 m:8.5mm NG

mondoana KIMMINS, 1962 m:5.5mm NG

260

LEPTOCE R I DAE: TRIAENODES 1 :2:2-

jubatus NEBOISS, 1982 m: 6 -7 mm; f: 6 -7 mm AU sw

~ ~~

intricata NEBOISS, 1977 m: 7 - 8 mm; f: 7.5 - 8 mm TA

bernaysae KORBOOT, 1964 m:6.5mm AU ne

261

LEPTOCERIDAE: TRIAENODES 1 :2:2-

volda MOSELY, 1953 m: 7 - 7.5 mm; f: 7 mm AU ne,se

fijianus MOSELY, 1941 m:5mm FI

dubia MOSELY, 1934 m:9mm FI

manni BANKS, 1936 m:6.5mm FI

insulana ULMER, 1951 m:6mm NG (Kei Is.)

262

LEPTOCERIDAE: OECETINELLA

hemerobioides (McLACHLAN, 1866) m: 8 mm;f: 6-7 mm SL, NG 1:2:2-

pulchel/a BANKS, 1936 m:5mm FI 0:2:2-

263

LEPTOCERIDAE: OECETIS 0:2:2-

mambia KIMMINS, 1962 m: 9 mm; f: 8 mm NG

ornata KIMMINS, 1962 m:6mm NG

buitenzorgensis ULMER, 1951 m: 8 mm; f: 7.5 mm JA, NG

kimminsi KUMANSKI, 1979 m: 11 mm; f: 11.4 mm NG

264


acuminata KIMMINS, 1962 m: 7.5 mm; f: 7 mm NG

/ongiterga KIMMINS, 1962 m:8.5mm

marginata KIMMINS, 1962 NG

L'l , , ,

," .... j.... I

0",: , ' ! i. " I, " "'- -~

asymmetrica KIMMINS, 1962 m: 8 mm; f: 7.5 mm NG

265

LEPTOCERIDAE: OECETIS

iti McFARLANE, 1964 m: 6.2 mm; f: 7 mm NZ g 1 :2:2-

arctipennis KIMMINS, 1962 m: 8 mm NG

reticulata KIMMINS, 1957

~'~~ ~iLJj; ~

~

m:4.5mm GU

0:2:2-

266

LEPTOCERIDAE: OECETIS 1 :2:2-

burtoni NEBOISS, 1978 m: 5 - 5.5 mm; f: 5.5 - 6 mm AU ne

australis (BANKS, 1920) m: 6 - 8 mm; f: 7 - 8.5 mm AU c,ne,se, TA

squamosa KIMMINS, 1962 m:6mm

267

LEPTOCERIDAE: OECETfS 1 :2:2-

pechana MOSELY, 1953 m: 8.5 - 10,5 mm; f: 7.5 - 11 mm AU sW,c,ne,se, TA

waJpolica NEBOISS, 1982 m: 6.5 - 8.5 mm; f: 6 - 7.5 mm AUsw

unicoJor (McLACHLAN 1868) m: 8 - 10 mm; f: 8 - 11 mm NZ, ST

268


chathamensis TILL YARD, 1925 m: 9-9.5 mm CH

umbra NEBOISS, 1977 m: 12 mm; f: 10.5 mm TA

gi/va NEBOISS, 1977 m: 10 -10.5 mm; f: 9.5 mm AU ne,se, TA

lurida KIMMINS, 1953 f: 11 mm AUse

269

LEPTOCERIDAE: OECETIS

aeoloptera KIMMINS, 1953 m: 5.5 - 6.5 mm; f: 5.5 - 6 mm AUne 1:2:2-

scirpicula NEBOISS, 1977 m:9mm;f:9mm TA 2:2:2-

laustra MOSELY, 1953 m: 7 - 9 mm; f: 8 - 8.5 mm AU sW,ne,se, TA 1:2:2-

270

LEPTOCERIDAE: DECETIS

arcada MOSELY, 1953

parka MOSELY, 1953

multipunctata ULMER, 1916

complexa KIMMINS, 1953

.

m: 6,5 - 8,5 mm; f: 7 - 8 mm TA

~ ~

&\ -------D

m:7mm AUse

m:5.5-6mm AUne

--------

m:7mm AUse

1:2:2-

271


asmanista MOSELY, 1953 m: 7.5 - 9 mm; f: 8 - 8.5 mm TA

minasata MOSELY, 1953 m: 6.5 - 7.5 mm; f: 7 mm TA

cymu/a NEBOISS, 1982 m: 6.5 - 7.5 mm; f: 6 -7 mm AUsw 1 :2:2-

LEPTOCER IDAE: OECETIS

atarpa MOSELY, 1953 m: 7 mm; f: 7 mm AUse

inscripta KIMMINS, 1953 m: 8 - 11 mm; f: 9 - 10 mm

272

v

AU sW,se, TA

I J I J J I I

1 :2:2-

273

LEPTOCERIDAE: SETODES 0:2:2-

bracteatus NEBOISS, 1982 m: 4.5 - 5.5 mm; f: 4.5 - 5.5 mm AU ne

~ ~";,,~

niveolineata KIMMINS, 1962 m: 5 - 6.2 mm; f: 4.7 - 5.4 mm NG

niveogra mmicus SCHMID,1986 m:7mm NG

papuana KIMMINS, 1962 m: 5 - 6 mm; f: 5 - 6 mm NG

2

3

4

5

6

7

8

LITERATURE CITED

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BANKS, N. 1924 Descriptions of new Neuropteroid insects. Bull. Mus. Compo Zool. 65: 421-455, 4 pLs

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9 COWLEY, D.R. 1976 Additions and amendments to the New ZeaLand Trichoptera. New Zealand J. Zool. 3: 21-26

10 HAGEN, H.A. 1859

11

12

13

Synopsis der Neuroptera CeyLons II. Verh.Zoo1.Bot. Ges. Wien 9: 199-212

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274

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21 KIMMINS, D.E. 1958

22

Miss L.E.Cheesman's expedition to New Hebrides, 1955. Orders Odonata, Neuroptera and Trichoptera. Bull. Br. Mus. Nat. Hist. (Entomol.) 6: 237-250

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24 KOLENATI, F.A. 1859

25

Genera et species Trichopterorum. Pars aLtera, AequipaLpidae. Nouv. Mem. Soc. (imp.) Nat. Mosc. 1'1: 143-296, 5 pL

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26 KORBOOT, K. 1964

27

Eight new species of caddis-fLies (Trichoptera) from the AustraLian region. Pap. Dept. Ent. Univ. Qd 2(2): 47-56

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28 KUMANSKI, K. 1979 Trichoptera (Insecta) from New Guinea. Aquatic Insects 1(4): 193-219

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33 McFARLANE, A.G. 1951 A note on the genus Neurochorema Till. and the additionof a species thereto. Rec. Cant. Mus.5: 253-254

34 McFARLANE, A.G. 1951

35

36

37

Additions to the N.Z. Rhyacophilidae. Part 2 Rec. Cant. Mus. 5: 255-265

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38 McFARLANE, A.G. 1966 New Zealand Trichoptera. Part 6. Rec. Cant.Mus. 8: 137-161

39 McFARLANE, A.G. 1973 Five new species of Trichoptera from New Zealand. J. Roy. Soc. NZ. 3: 23-34

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49 MARTYNOV, A.V. 1914

50

51

52

Notice sur quelques formes nouvelles de Trichopteres, provenant de differentes localites.Ezheg. Zool. Muz. (Ann. Mus. Zool. Acad. Sci. St.Petersburg) 19: 125-133

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55 MOSELY, M.E. 1933 The genus Smicridea McLachlan (Trichoptera) in Tasmania. Ann. Mag. Nat. Rist. (10)12: 216-221

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58 MOSELY, M.E. 1934 New Fijian and African Trichoptera. Eos 9: 17-27

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276

62 MOSELY, M.E. 1951 Descriptions of new Sarawak Trichoptera. Ann. Mag. Nat. Hist. (12)4: 480-490

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65 NAVAS, L. 1924 Neue Trichopteren. Konowia 3: 204-209

66 NAVAS, L. 1930

67

68

Insectos deL Museo de Paris. Broteria Serie Zool. 26: 19-23

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74 NEBOISS, A. 1974 Additions to the famiLy Kokiriidae (Trichoptera) Vict. Nat. 91: 175-179

75 NEBOISS, A. 1974

76

A new caddis-fLy genus from Victoria and Tasmania (PhiLorheithrinae: Trichoptera). Vict. Nat. 91: 322-325

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77 NEBOISS, A. 1977

78

A taxonomic and zoogeographic study of Tasmanian caddis-fLies (Insecta: Trichoptera). Mem. Natl. Mus. Vict. 38: 1-208

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79 NEBOISS, A. 1980 AustraLian species of the genus Anisocentropus McLachLan (CaLamoceratidae: Trichoptera). Aust. J. Mar. Freshwat. Res. 31: 193-213

80 NEBOISS, A. 1980 First record of the subfamiLy HyaLopsychinae from AustraLia (Trichoptera: PoLycentropodidae). Arch. Hydrobiol. 90: 357-361

81 NEBOISS, A. 1981 Tasmanian Caddis-fLies (Fauna of Tasmania Handbook No.4) Hobart: University of Tasmania 180pp

82 NEBOISS, A. 1981

83

Distribution of Trichoptera famiLies in AustraLia, with comments of fauna in the south-west. Proc. 3rd Int. Symp. on Trichoptera. Perugia 1980. (Series Entomologica 20: 265-272) The Hague: W. Junk

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89 NEBOISS, A. 1984

90

91

Review of taxonomic position of AustraLian and New Guinean species previousLy ascribed to Macronema (Trichoptera: Hydropsychidae). Proc.Roy. Soc. Vict. 96: 127-139

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93

94

95

96

97

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98 SCHMID, F. 1969

99

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109 ULMER, G. 1908

110

111

112

113

Trichopteridae und Ephemeridae. Fauna SudwestAust. 2(3): 25-46

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E. Jacobson auf Java geNotes Leyden Mus. 32:

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114 ULMER, G. 1916

115

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124 WELLS, A. 1981 The genera Oxyethira Eaton, Gnathotrichia Ulmer and Stenoxyethira Kimmins (Trichoptera: Hydroptilidae) in Australia. Aust. J. Zool. 29: 103-118

125 WELLS, A. 1982

278

Tricho1eiochiton Kloet and Hincks and new genera in the Australian Hydroptilidae (Trichoptera) Aust. J. Zool. 30: 251-270

126 WELLS, A. 1983 New species in the Australian Hydroptilidae (Trichoptera), with observations on relationships and distributions. Aust. J. Zool. 31: 629-649

127 WELLS, A. 1984 Hydropti1a Dalman and Orthotrichia Eaton (Trichoptera: Hydroptilidae) from the islands of New Guinea and New Britain, with observations on relationships. Aust. J. Zool. 32: 261-282

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129 WISE, K.A.J. 1958 Trichoptera of New Zealand 1. A catalogue of

Works in preparation

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136 WELLS, A. 1985 Four new species of Hydroptilidae (Trichoptera) from the Alligator Rivers region, Northern Territory. Trans. Roy. Soc. S. Aust. 109:

137 SCHMID, F. 1986 Considerations diverses sur quelques genres leptocerins. Mem. Entomo1. Soc. Can.

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130 WISE, K.A.J. 1962 A new genus and three new species of Trichoptera. Rec. Auck. Inst. Mus. 5: 247-250

131 WISE, K.A.J. 1965 An annotated list of the aquatic and semi-aquatic insects of New Zealand. Pacific Insects 7: 191-216

132 WISE, K.A.J. 1970 Trichoptera of New Zealand 2. The present status of R.J.Tillyard's species of New Zealand Trichoptera, with notes on the type specimens. Rec. Auck. Inst. Mus. 7: 201-215

133 WISE, K.A.J. 1972 Trichoptera of the Auckland islands. Rec. Auck. Inst. Mus. 9: 253-267

134 WISE, K.A.J. 1977 A synonymic checklist of the Hexapoda of the New Zealand Sub-region. The small orders. Bull. Auck. Inst. Mus. 11: 1-176

135 WISE, K.A.J. 1982 Two new species of Trichoptera from the Murchison Mountains, South Island, New Zealand. Rec. Auck. Inst. Mus. 19: 149-151

138 NEBOISS, A. 1986 Taxonomic changes and descriptions of new caddis-fly species from SW Pacific - Australian region (Insecta: Trichoptera). Mem. Mus. Vict.

139 NEBOISS, A. 1986 Identity of Trichoptera species described by Korboot 1964-65. (Insecta: Trichoptera). Mem. Mus. Vict.

Abacaria Acanthotrichia Achorema see Apsilochorema Acritoptila Adectophylax Adicella Aethaloptera Agapetus Allobiosis Allochorema Alloecella Alloecentrella Anachorema see Ulmerochorema

120 73

69 138 256 136

55 53 14

206 201

Anisocentropus 224 Anomalostoma see Philanisus Antipodoecia 196 Aoteapsyche 123 Aphilorheithrus 216 Apsilochorema 11 Archaeophylax 165 Asmicridea 127 Atrachorema 31 Atriplectides 212 Austratrichia 68 Austrecnomina see Austrheithrus Austrochorema Austropsyche

Plectrocnemia 215

15 132

Bachorema see Apsilochorema Baliomorpha 133 Barynema 210 Beraeoptera 191

Caenota Caledonotrichia Caledopsyche Caloca Calocoides Chathamia Cheumatopsyche Chimarra Chimarrha see Chimarra Coenoria Condocerus Confluens Conia see Conuxia

203 70

121 198 201 160 117 104

190 254 191

Conoesucus 186 Conuxia 193 Costachorema 29 Costora 184 Cryptobiosella 98

Dampfi trichia (subgenus) 82 Dinarthropsis 169 Diplectrona 128 Dolophilus see Hydrobiosella

Ecnomina 152 Ecnomus 150 Edpercivalia 39 Ethochorema 26

Ganonema see Anisocentropus Gnathotrichia see Oxyethira Goera Goerodes Gracilipsodes see Triplexa

Hampa

167 169

188

279

INDEX OF GENERIC NAMES Synonyms are in italics

Helicopha Helicopsyche Hellyethira Herbertorossia Hudsonema Hyalopsyche Hyalopsychella Hydrobiosella Hydrobiosis Hydrochorema Hydropsyche Hydropsychodes Hydroptila

207 220

76 118 248 148 148

99 32 41

115 122

60

Ipsebiosis 18

Koetonga 18 Kokiria 177 Kosrheithrus 218

Lectrides 244 Leptocerus 255 Leptopsyche 135 Leptorussa 254 Liapota 182 Lingora 189 Loticana see Symphitoneuria

Macronema (-um) see Macrostemum Marilia Matasia Maydenoptila Mecynostomella Megogata

Baliomorpha 135 210 188

Molanniella see Tasimia Monopseudopsis cf. Macronemum

zebra tum Hagen (N.America)

71 177

22

Moruya 51 Mulgravia 68

Nanoplectrus 182 Neobiosella 103 Neolepidostoma 169 Neureclipsis 141 Neurochorema 42 Notalina 249 Notanatolica see Triplectides Notidobia cf. Anisocentropus

latifascia (Walker) Notiobiosis see Taschorema Notoperata 247 Nyctiophylax 138

Oecetina see Oecetis Oecetinella 262 Oecetis 263 Oeconesus 171 Oestropsyche 136 Olinga 194 Ornatus see Tanorus Orphninotrichia 65 Orthopsyche 125 Orthotrichia 85 Oxyethira 82

Paroxyethira 80 Percivalia see Edpercivalia Periwinkla 193 Philanisus 160

Philorheithrus Plectrocnemia Plectrotarsus Pliocaloca Polycentropus Polyplectropus Pseudoeconesus Psilochorema Psyllobetina Ptychobiosis Pycnocentria Pycnocentrella Pycnocentrodes

214 139 181 202 142 145 172

43 52 28

192 199 190

Rakiura 222 Ramiheithrus 216

Saetotricha see Helicopsyche Sciops see Diplectrona Setodes 273 Smicridea see Smicrophylax Smicrophylax 126 Sortosa see Hydrobiosella Stenopsychodes 111 Stenoxyethira see Oxyethira Symphitoneuria 245 Symphitoneurina 246 Synagapetus see Agapetus Synchorema 50

Tamasia 200 Tanjilana 22 Tanjistomella 179 Tanorus 38 Tarapsyche 174 Targatrichia see Orthotrichia Taschorema 23 Tascuna 174 Tasiagma 163 Tasimia 162 Taskiria 178 Taskiropsyche 179 Tasmania see Liapota Tasmanoplegas 141 Tasmanthrus 214 Tinodes 158 Tiphobiosis 47 Tismana see Caloca Traillochorema 31 Triaenodes 257 Trichoglene (subgenus) 83 Tricholeiochiton 74 Triplectides 232 Triplectidina 243 Triplexa 253 Triplexina see Notoperata

Ulmerochorema 19 Westriplectes 231

Xanthochorema 14 Xenobiosella 103 Xuthotrichia 73

Zelandopsyche Zelandoptila

175 158

Zelobiosella see Hydrobiosella Zelo1essica 208 Zelomyia see Zelandoptila Zepsyche 175

280

INDEX OF SPECIES NAMES

Showing source of illustrations; synonyms are in italics

aberrans Kinunins (Tinodes) [22] f.34 158 aberrans Wells (Orthotrichia) [122] f. 76, 77 95 ablusus Neboiss Orig./p-ty 58 acheir McFarlane [41] f.24-27 43 acinaces McFarlane (Zepsyche) [36] f.7-10 175 acinacis Wells (Hydroptila) [120] f.30-34 63 acta Neboiss [123] f.9-11 66 aculeata Wells [122] f.38-40 87 acuminata Kinunins [22] f.53 264 acuta Neboiss [86] f.1-4 200 adornata Wells [122] f.1-9 85 advena Wells [127] f.35,36 89 aeoloptera Kinunins [63] f .201 269 aeris Wise [9] f.4 190 agilis (Hudson) [38] f.36-43 214 aiyura Korboot Orig./p-ty 105 akroreia Neboiss [72] f. 78-82 22 albanus (Mosely) [63] f.150,151 231 albescens Tillyard [63] f.47 220 a I b ice p s (McLachlan) [63] or f. 346; [29] ~ f. 5 84 albodecorata Jacquemart - see arcada Mosely aliena (McLachlan) [63] 6' f.165;0 wings Orig. 248 alloneura Brauer - see plebeius W~lker allynensis Wells [121] f.14,15 78 alpinum Neboiss [72] f.22-26 15 altenogus Morse & Neboiss [51] f.81,99 239 altera McFarlane (Polyplectropus) [41] f.1-4 146 altera Neboiss (Plectrocnemia) [77] f.279-284 140 alveata Wells [122] f.52,53 91 ama b il i s (McLachlan) [63] f .167; 0 wings Orig. 248 amblyopia Neboiss [83] f.12-15 + 99 amiena Sykora [100] f.6 117 ampla Korboot [14] f.9,10 245 anasina Neboiss [77] 210,211 102 anei tyuma Neboiss Orig./h-ty d'; p-ty ~ 167 angelae Neboiss [78] f .23-25 231 angusta Banks Orig. 129 angustipennisMosely [77] f.673,678-681 214 annulata Wells [127] f.29,30 90 anoac1ana (Malicky) [30] f.4f-j 104 antennifera Wells [126] f .3,4 71 aorere Henderson [12] f.8 100 apalapsili (Malicky) [30] f.4a-e 56 apobamum Neboiss [77] f .104-108 23 aptus Neboiss [83] f.111-113 254 aquilonia Neboiss [87] f.1-6 201 arcada Mosely [77] f.796-800 270 arctipennis Kinunins [22] f .50 265 arc ua t a Kinunins [63] f. 271 103 armata Jacquemart(Hydrobiosella)[77] f.218,219 101 armata Wells (Orthotrichia) [122] f.17,18 86 armstrongi McFarlane [33] f .36,37 42 artuvillosus (Wells) [124] f.29-34 82 asci ta Neboiss [77] f.463-467 198 asmanista Mosely [77] f. 792-795 271 asmanum Mosely [77] f.99-102 23 assimulans (Ulmer) Orig. by Schmid 256 astia Mosely [63] f.97-99 207 asymmetrica Kinunins [22] f.57 264 atarpa Mosely [63] f.202,203 272 atra (Banks) [63] a' f.84; Orig./h-ty ~ 162 atraseta Wells [122] f.14-16 85 atratus Mosely [54] f.15,16 151 at sou Schmid Orig. by Schmid 255 attenuata Wells [126] f.33-35 93 attunga Neboiss [72] f.170-174 53 aurata Mosely [77] f.570-575 189 aurea Kinunins [22] f.66 259 a ur eo f u sca Kinunins [20] f. 6 106 aureola (McLachlan) [63] f.51,52 190

aureonigra Schmid [98] p1.2 f.1,2 112 auricollis Kinunins [22] f.30 143 auricoma Hare [40] species not recognizable au rife r urn Neboiss [89] f. 32-34 135 a ur i fu sc a McFarlane [35] f.14-17 146 austera Neboiss [77] f.394-404 178 australica Banks(Plectrocnemia)[77] f.287,288 140 australica (Ulmer) (Chimarra) [63] f.274 105 australicum Mosely (Barynema) [63] f .109 210 australicus Banks (Triplectides)[51] f.61,90 234 australiensis Wells - see incana Ulmer australis (Banks) (Oecetis) [77] f. 777-783 266 australis (-e) McLachlan (Macronema) cf.Macro-

sternum zebratum Hagen of N.America australis (Navas) (Chimarra) [63] species not

recognizable, no figures available australis Navas (Triplectides) [51] f.54,83 232 au s t r a lis Ulmer (Polycentropus)[ 113] f. 9,10 143 australis (Ulmer) (Smicrophy1ax) Orig. 126

banghaasi Ulmer [79] f.1,10-15 225 banksi (Mosely) (Apsilochorema) [61] ~ f.29,30

[93] or f.3 13 [89] f.9,10,35-38 133 banksi (Mosely) (Baliomorpha)

bank s i en sis (McFarlane) [63] barretti Korboot [26] f.7-9; bartona Mosely [63] f.43 basilobata Wells [121] f.1l-13

f.305 39 ~ Orig./p-ty 120

22l

batyle Neboiss [77] f.272-276 baynesi Wells [126] f.1,2 bellicosa Wells [122] f.72,73 benambrica Wells [126] f.38,39,41 bernaysae Korboot Orig.d" beroni Kumanski [28] f.24-28 b i color a t u s (Martynov) [79] f. 2,50-55 bid ens McFarlane [34] f. 29,30 bifaria Neboiss [77] f. 741-743 bifurca Wells [125] f.17-21 bifurcata (Kinunins) Orig. 6' bilamina Wells [125] f.51-57 bilobus Neboiss [51] f.59,88 b i ramo sa Kinunins [20] f. 5 bishopi Wells [122] f.19-22 b i spina Kinunins (Hydrobiosella) [63] f. 270 bispina Kinunins (Hydroptila) [22] f.6,7 bispinosa (Jacquemart) [77] f .363 bistirpis Wise [129] f.4 boera Mosely [63] f .14 bola Mosely [63] f.llO,l11 boorarus Neboiss [83] f. 71-73 borealis (McFarlane) [63] f.363 bourina Mosely - see spinata Banks brac hy p ter a McFarlane [41] f. 40-42,44 bracteatus Neboiss [84] f.1-9 bradleyi Kinunins [20] f. 7 breve (Mosely) (Ulmerochorema) [77] f.65,66 brevipennis Tillyard [92] f.16-21 brevis Wells (Oxyethira) [124] f.17,18 b ron tens i s Neboiss [77] f. 605, 608, 609 brunneum (Mosely)[63] or f.295;[72] ~ f.107-109 bryanti Banks [118] f.318-320 budgei McFarlane [36] f.14-18 bugata Wells [127] £'15,16 bu i t enzor gens i s Ulmer [1l8] f. 708-712 bulla Wise Orig. bullata Wells [1221 f.36,37 burgersi Ulmer (Apsilochorema) Orig./h-ty 0' burgersi Ulmer (Hydropsyche) [1l3] f.16-18 burtoni Neboiss [78] f.37-40

76 153

72 94 65

260 117 227

46 250

75 132

73 235 108

86 102

61 132 172 190 210 218 40

30 273 147

21 160 84

187 27

116 33 62

263 129 88 12

115 266

ca1cara Wells [120] f.23-29 63 ca1edon Kimmins [18] f.6-10 14 callistum McFarlane [41] f.37-39,43 29 canarus Neboiss [71] f.15-18 165 caenescens McLachlan - see magnus (Walker) caparti Jacquemart (Lingora) - see aurata Mosely caparti Jacquemart (Tasmania) - see 1avara Neboiss caperata Wells [127] f.3,4 64 capillata Wells [122] f.74,75 94 carteri Wells [126] f.44-46 68 caudata Neboiss [77] f.307-310 139 caudicea Neboiss [87] f.12-14 134 cassico1a (McFarlane) [63] £.302 41 castanea Kimmins Orig./h-ty 130 ca ther inae (McFar1ane)[ 36 ]f.11-13; [40 ]f.12, 20 124 cepha10tes (Walker) [63] f.137 241 ceru1a Neboiss [77] f.208,209 102 cha1codes McFarlane [41] f. 7-11 33 chapmani Kumanski [28] f.48,50 145 charadra Neboiss [77] f.134-136, 138, 139 51 charadraea McFar1ane[34]d' f.19-21; [38]~ f.60,61 36 cha thamens i s Tillyard [104] f. 5,6 268 cheesmanae Kimmins (Apsilochorema) [21] f.6 13 cheesmanae Kimmins (Ca1edopsyche)[18] f.13-16 121 chee smanae Kimmins (Chimarra) [22] f .18 107 cheesmanae Kimmins (Hydropsychodes) [22] f.41 122 cheesmanae Kimmins (Leptocerus) [22] £.48 256 cheira McFarlane [35] f.9,11-13 208 cheirodes McFarlane [41] £,18-22 44 chi1di McFarlane (Periwink1a) [39] f.3-5 193 chi1di McFarlane (Tiphobiosis) [41] f.56-58 49 chi10ma Neboiss [89] f.23,24 134 chiltoni Ti11yard - see evecta McLachlan (part)

and aureola (McLachlan) (part) chorisis Neboiss [78] f.4-6 154 ciliata Neboiss [77] f.444-450 163 cinctigera Wells [127] £.39,40 90 ci uskus c i u skus Mosely [51] f. 62,91 234 ciuskus seductus Morse & Neboiss [51] £.63 234 c1a v igera McFarlane [34 Jar £'16,17; [38]~ f. 56,57 36 clivico1a Neboiss [72] f.144-151 18 coch1eaetesta Korboot Orig. 221 cognata Kimmins (Dip1ectrona) Orig./p-ty 130 cognata Kimmins (Hydrobiose11a) [77] f.220-224 101 cognatus McLachlan (Trip1ectides) Orig.by Morse 241 cohi b i li s Neboiss [83] f. 26-28,38 154 co10nica (McLachlan) [63] f.223; [40] f.ll,19 123 columba (Neboiss) [124] f.l-l0 83 commi s ta Wells [123] £. 33,34 72 comp1exa Jacquemart (Austrochorema) [77] f .49-51 17 comp1exa Kimmins (Oecetis) [63] f.200 270 concubium Neboiss [72] f.29,30 16 conferta Wells [126] f.12-14 95 confluens Ulmer - see hemerobioides McLachlan confusum (McLachlan) [63] f.316 42 continentalis Ulmer [83] f.16,17 150 coomata Mosely [63] f.61 189 copis McFarlane [36] f.20-22 37 corallina Kimmins [22] f.65 259 corinna Neboiss [77] f.204,205 102 cornuta Wells [121] f.37-40 78 coronata Wells [125] f.35-38 68 corvinus Neboiss [79] f.29-34 226 costalis Kimmins [22] f.60 257 costatum Banks Orig. 210 cra1us (Mosely) [77] f.154-159 57 crassicaudatum Tillyard [63] f .318 41 creektona Neboiss Orig. 126 c re to sus McLachlan [118] f. 513-516 225 crinitum Neboiss [77] f.36-38,40-42 17 cubitans Wells [121] f.16-18 76

281

cucullata Wells [127] f.21,22 c umbe r 1 and i ca Neboiss [72] f.164-168 cuneo1a Neboiss [123] f.23-30 c yc lop i ca Kimmins (Chimarra)[ 22] f. 9 cyc10picus Kimmins (Ecnomus)[22] f.35 cygnitus Neboiss [83] f.18,19 cymu1a Neboiss [83] f .132-136

89 53 71

107 151 150 271

daabanum Ulmer [118] f.802,803 169 dar1ingtoni (Banks) [63]d' f.192; [77]~ f. 758 254 dasodes Neboiss [87] f.10,18-20 202 dayi Ross [95] f.350 56 decoratus Neboiss [77] f.662-665 217 decussatum Ti11yard - see confusum McLachlan de1amarei Jacquemart [77] f .497-504 207 de 1i ca t u 1a (Ulmer) [83] f. 93-96 251 de 1 or a Mosely [S3] f, 26; [77]~ f. 543 185 densus (Korboot) Orig./p-ty 38 dentata Wells [121] f.24-26 78 denticu1ata Jacquemart [77] f.433-438 162 diacanthus Neboiss Orig./p-ty 56 digitata Wells [127] f.49,50 91 digitiferus Jacquemart [77] f.594,596-599 187 di1gri Wells [126] f.24-26 86 dilucidus McLachlan [115] f.ll 224 dis j unc ta Neboiss [80] f.1-8 148 dispara1is Wells [127] f.31,32 90 divaricata Wells [126] f.22,23 95 diversa Neboiss [83] f.91,92 248 do1abratus Morse & Neboiss [51] f.33,72 236 do1ichos rkFar1ane [41] f.81-D6 242 do na1 d son i McFarlane [36] f. 27-33 46 drepana Neboiss [77] f.439-443 162 drummondi Illies [14] f.2,3 144 dubitans Mosely [63] f.128 215 dubia Mosely (Triaenodes) [58] f.2-4 261 dubia (U1mer)(Ba1iomorpha) [89] f.7,8,46,47 133 dubius Mosely (Atrip1ectides) [63] f.113,114 212 dubius Mosely (Trip1ectides) - see e10ngatus Bks dulce (McLachlan) type lost in mail 1984 dwe1linga Neboiss [83] f.104-106 251

eatoni Mosely [63]ar f.348; [29]0 f.8 80 eba Mosely [63] f.105 + 199 ebenina Neboiss [77] f.546-550 185 echinata Neboiss [89] f.15-17 134 edmondsi Wells [125] f.22-25 74 ed war d si (lkLach1an) [63] f. 230 127 e1egans Kumanski (Po1ycentropus) [28] f.42-44 144 elegans (Walker) - see 1atifascia (Walker) e10ngatus Banks [51] f.73,95 237 el'lbersoni Wise [135] f.1-3 45 empheres Neboiss [88] f.10-13 38 ensiformis Wells [127] f.43-45 94 enthesis Neboiss [51] f.64,92 236 eremica Neboiss Orig./h-ty 163 erratica Mosely c! Orig.; [63]~ f.327 53 eruensis Mosely [63] f.95;[38] f.25-27 199 eskensis (Mosely) [121] f.27-30 79 eungellus Neboiss [79] f.56-59 227 eurhinata Wells [136] f.11,12 88 evansi (Mosely) (Austrochorena) [77] f.32-34,46 17 evansi Mosely (Taschorema)[63]~ f.297;[72]~f.126 25 evansi Ross (Agapetus) [95] f.349 58 evecta McLachlan [63] f.16 192 exca va ta Kimmins [22] f. 37 119 excisa Kimmins (Stenoxyethira) - see incana(U1mer) excisa Kimmins (Triaenodes) [20] f.14 257 exigua (McLach1an)(Symphitoneuria) [63] f.181 245

exigua Wells (Orthotrichia) [122] f.67-69 92 eXlmla Neboiss [83] f.61,62 139 expe d i ti on i s Ulmer [28] f. 29-32 122 explicata Wells (Hydroptila) [127] f.1,2 61 explicata Wells (Maydenoptila) [123] f.31,32 72 exserta Wells [121] f.19-23 77

falcata Kimmins [22] f.20 105 falcis Wise [129] f.13 37 fal1ia Mosely [63] f.106 198 fasciatus Riek [92] f.38-43 160 fastigiata Neboiss [87] f.12-14 202 feredayi (McLachlan) [63] f.85,86 194 ferulum Neboiss [77] f.111-114 23 fidelis Wells [125] f.1-10 74 fijiana Banks (Goera) Orig./d' h-ty;~ p-ty 167 fijiana (Mosely) (Abacaria) [58] f. 6-9 120 fijianus Banks (Anisocentropus) Orig./h-ty 229 fi j ian u s Banks (Polyplectropus) Orig. /h-ty 145 fijianus Mosely (Triaenodes) [61] f.6-8 261 fimbriata (McLachlan) (Orthopsyche)[63] f .222 125 fimbriata (Mosely) (Hellyethira) [63] f.352 78 flabel1a Wells [126] f.18,19 96 flava (Ulmer) (Notalina) [83] f.100,101 249 flavus Ulmer (Nyctiophylax) [113] f.11,12 138 flavicaput McLachlan - see latifascia (Walker) flynni Korboot Orig./p-ty 115 folioharpax McFarlane [35] f.33-35 45 forcipata Mosely [63] f.17 192 forsteri McFarlane [37] f.47-51 42 fragilis Wells [127] f.25,26 89 frater McLachlan [63] f.282,283 32 fromus Mosely [77] f.584-588 186 fulgidus Navas Orig./h-ty 224 fulva Kimmins (Notalina) [77] f.737-740 250 fulva (Navas) (Symphitoneurina) [97] f.111-114 246 fulva Tillyard (Tiphobiosis)[41] not recognised 49 fumosa Wise [129] f.6 194 funerea McLachlan [63] f.18 192 furcata Henderson [12] f.3 98 furcilla Neboiss [86] f.8,9 200 furcula Wells [127] f.l7,18 61 fusca Kimmins (Marilia) [63] f.112 210 fusca Kimmins (Mecynostomella) [18] f.17-24 177 fusca (McFarlane) (Edpercivalia) [63] f.304 39

ga.lbinomaculatus Jacquemart - see angustipennis Mos. galeata Neboiss [87] f.21-24,31,32 204 geevestonia Neboiss - see asmanista Mosely gilolensis (McLachlan) [63] f.139 241 giluweana Neboiss [88] f.9 38 gilva Neboiss [77] f.774-776 268 gisbum (Mosely) [77] f.1O-16 11 glebula Wells [127] f.33,34 90 globosa Wells [125] f.39-42 69 glymma Neboiss [77] f.633-640 215 gol1anis Mosely [63) f.276 34 gonetalus Horse & Neboiss [51] f.78,97 239 goroca Sykora [100] f.4 106 g r a c iIi s McLachlan (Leptopsyche) [107] f. 25 l35 gracilis Wells (Orthotrichia) [122] f.54-57 92 grandis Kimmins [22) f.26,27 142 gravenhorsti Ko1enati [77] f.365,366 181 greeClwoodi Hosely [61] f .14-20 145 gressitti Sykora [100] f.5 109 grisea Banks (A1loecel1a) [77) f.507,509-511 206 grisea Mosely (Asmicridea) [55] f.1-3 127 gungarra Neboiss [86] f .12-15 252 gunni (McFarlane) [35] f .5-8 193

282

hamatus Morse & Neboiss (Triplectides)[51]f .80 240 hamatus Iolells (Acritoptila) [125) f.44-46 69 hamil toni (Tillyard) [38] f.3-5 191 har p i d i 0 sa McFarlane [34]d" f. 7,8; [38]~ f. 48,49 35 harrisoni Wise [l35] f.4-7 39 hastata Henderson [12] f.2 98 hawdonia McFarlane [35] f.2-4 193 heacota Mosely [63] f.42 222 hebdomon McFarlane [41] f.33-36 30 hecton McFarlane [41] f.30-32 30 helvolus Morse & Neboiss [51] f.55,84 232 hemerobioides (McLachlan) [118] f.713-718 262 hendersoni Mosely [63]d' f.347; [29]0 f.6 80 hesperium (Neboiss) [72] f.121-125 + 28 hi emali sTillyardOrig.111 hienghene Malicky [31] f.1 109 hi n ton i Leader [29] f. 1 , 2 80 hobbyi Mosely [62] f.9-12 116 hortena Mosely [63] f.100 207 howesi Tillyard [132] f.2-4 220 hudsoni Mosely [63] f.74 172 hyboma Neboiss Orig./h-ty 229 hystricosa Neboiss [78] f .14-19 131

iena ~1osely [63] f.25 ignota Neboiss [77] f.421,423-425;0 Orig. il1iesi Sykora [100] f.1A,B +

184 174

ill u s t r i s McLachlan [79] f. 4-7; forewing Orig. immunis McLachlan antenna Orig./h-ty Barnard impluvii Wise [130] f.4

70 224 229 147

iltona Mosely - see zealandica 1udson incana Ulmer [124] f.23-28 incertula Mosely [63] f.343 incisus Mosely [63] f.68 indigota Mosely [61] f.22-26 inermis (Banks) Orig. ingens Tillyard [63] f. 78-80 in g e n u a Hare [63] type destroyed in ornata Wells [122] f.41,43,44

82 61

171 104 131 175

inscripta Kimmins (Oecetis)[83] f.127, 128 inscriptus Walker (Monopseudopsis) cf.australis

88 272

McLachlan (i·lacronema) insperatus Morse & Neboiss [51] f.79 insulana Ulmer [118] f.693-695 integripennis Riek [92] f.22-27 in terme d ia Mosely [63] f. 332 intricata Neboiss Orig. io Kimmins [22] f.44,45 i r ro rata Kimmins (Ecnomina)[ 63] f. 264; [77]~ irrorata Wise (Neobiose11a) [129] L12 isabellina Ulmer - see incana Ulmer

239 261 160

47 260 228

258 155 103

it i McFarlane redrawn from original 265

jabirel1a Wells [136] f.4,6,7 74 jafiwi Ross Orig. 55 japenensis Kimmins [22] f. 72-74 169 jeanae McFarlane [38] L17-22 194 johnsi McFarlane [41] L13-17 33 jubatus Neboiss [83] f.115-120 260 justini Wells [126] f.36,37,40 67

karoriensis Mosely [63] f.77 173 kelion Neboiss [77] f.96,98 27 kenampi (Korboot) Orig./p-ty 144 kholoensis Wells [122) f.61,62 92 !<iddi HcFarlane [34] L9-11 32 kimminsi Kumanski (Herbertorossia)[28] f.l3-15 118 k i mm ins i Kumanski (Oecetis) [28) L 54-58 263

kimminsi Leader (Paroxyethira) [29] f .3,4 81 kimminsi Neboiss (Taschorema) [72] L110-114 24 kimminsi Ross (Agapetus) [95] L351 55 kirramus Neboiss [79] L16-21 228 kocinus Neboiss [77] L650-652 216 kokodana Kimmins (Chimarra) [22] L24 109 kokodana Kimmins (Orthotrichia) [22] f. 5 89 koropa Wells [127] L10,11 64 krene Neboiss [77] L554,555 184 krokale Neboiss [78] L 7-9 154 kurandica Wells [123] L35,36 71 kuscheli Wise [133] L17-23 48

lacuna Neboiss [77] L291,292 140 lac us t r i s Neboiss (Taskiropsyche)[ 77]L 412-420 179 lacustris Tillyard (Philorheithrus)[63] L121 214 lac usal b inae Kimmins [63] L 249 147 lanceolata Kimmins [20] L16 257 laparus Neboiss [77] L161,165 57 latifascia (Walker) [79] L23-28 226 la t i penn i s Mosely [60] L15-22 242 latosus Ross Orig./p-ty 56 laustra Mosely [83] f.129-131;[77]~ f.790,791 269 lavara Neboiss [77] L374-378; [70] ~ L9,1O 182 legula Neboiss [77] f.261-268 152 lentigo Wells [127] L27,28 90 lentum Neboiss [72] f.35-39 20 lepnevae Jacquemart [81] L33,34 17 leptoharpax McFarlane [63] L360 44 1 e t t i Korboot Orig. /h-ty 101 licmetica Neboiss Orig./p-ty 246 lindsayi Tillyard [104] L1 34 lineata Neboiss [77] L234-242 113 liratellus Morse & Neboiss [51] L70 237 liratus Morse & Neboiss [51] L69,94 237 litotes Wells [127] L23,24 90 litua Wells [121] L51,52 77 lobata Jacquemart (Triplexina) - see maculata(Mos.) lobatus Wise (Oeconesus)[129] L3a-c 171 locula Neboiss [72] f.176-180 52 1 ongi s p i na Jacquemart(Alloecella)[ 77]L 512-522 206 longispina Kimmins (Triaenodes) [22] L59 258 longiterga Kimmins [22] L54 264 loriai (Nav's) (Baliomorpha)[89] f.11,48 134 loriai (Navas) (Macrostemum) [89] f.29-31 135 loriana (Navas) Orig./h-ty 108 loripes Wells [121] L33,34 76 losida Mosely [120] f.35-38 60 lurida Kimmins [63] L210 268 1 u teo 1 u s Neboiss [77] L 670-672 216 luxata Neboiss [77[ L561-563 184 luxaturum Neboiss [72] L66-69 20 lyella Neboiss [77] L356-358 130

maclachlani Mosely (Plectrocnemia)[63] L245 139 maclachlani Ulmer (Anisocentropus)[115]L14-16 224 maclellani McFarlane [41] L71-74 175 mac r oha r pax McFarlane [63] L 290 46 maculata (Mosely) (Notoperata) [77] L717-720 247 maculata Mosely (Orphninotrichia) [123] Ll-8 65 rna fu 1 ua Kimmins [22] L 43 128 magnicornis Wise [129] L7,8;[38] ~ L30-32 201 magnus (Walker) [51] L5,56,85 233 mall eo forma Wells [121] f. 41-46 77 mambia Kimmins [22] f.52 263 mangu McFarlane [37] L34-40 31 manicata Neboiss [77] f .298-301 139 manni (Banks) (Chimarra) Orig./h-ty 104 manni Banks (Polyplectropus) Orig./h-ty 145 manni Banks (Triaenodes) Orig./h-ty 261

283

maori McLachlan [63] f.63-65 171 margaretae Wells [125] L47-50 69 marginata Kimmins [22] f.55,56 264 marlieri Jacquemart - see astia Mosely mataura McFarlane [35] L28-30 45 maxima (McFarlane) [63] f .303 39 mccubbini Neboiss [77] L405-407 178 media Wells [123] L15,16 66 meizon McFarlane [41] f.75,77-79 208 melanochrysa Tillyard d" Orig.; [98]~ p1.2.L9 112 membrum Neboiss [72] L55-60 19 merga Neboiss [83] L57-59 155 mesembria Neboiss [83] L29-31;40,41 156 michaelseni (Ulmer) [83] f .8-11 99 mienica Wells [124] L14-16 83 miharo McFarlane [37] f.59-67 177 mimicum McLachlan [63] L287,28G 43 mimus McLachlan [63] L75,76 173 minasata Mosely [77] L 784-787 271 minor Mosely (Plectrotarsus) [63] f.10 181 minor Sykora (Caledonotrichia) [100] f.2a-c 70 mixta (Cowley) Orig. 100 mjobergi Ulmer [63] L254 111 modesta Cowley [9] f.3 190 modica (McLachlan) [63] L225; ~ Orig. 117 mondoana Kimmins [22] L64 259 montana Tillyard (Stenopsychodes)[98] f.5,6 113 montana Tillyard (Tiphobiosis)[41] f.52-54 47 monteithi Neboiss [37] L28-30,36,37 204 monticola Kimmins (Chimarra) [63] f .275 105 monticolus Banks (Agapetus) Orig. 56 morula Wells [122] L48-51 91 moselyella Kimmins [19] f.2-4 12 moselyi Jacquemart(Conoesucus) - see fromus Mosely mo s ely i Kimmins (Hydropsyche) [22] L 36 116 moselyi Kimmins (Notalina) [63] L174,175 249 moselyi Kimmins (Polycentropus) [22] L29 142 mo s ely i Ross (Apsilochorema) [93] L 4 13 moselyi Tillyard (Zelandoptila) [35] L21-25 158 motueka Henderson [12] L 7 103 mounthageni Kumanski [28] L38,40 142 muc rona ta Neboiss [87] L 8,15-17 202 mul tilobata Wells [121]L47-50 77 multipunctata Ulmer [63] L206 270 muricatus Neboiss [79] L37-42 226 murrumba Mosely [63] L45; [77] ~ L458,459 221 muscari Wells [126] L20,21 95

nama Neboiss [72] L31-34 16 napaea Neboiss Orig./p-ty 141 natasia Mosely [63] L83 163 neboissi Wells [125] L26-34 68 necopina Neboiss [72] L 70-76 22 nemorale HcFarlane [63] L289 45 nemorosa Neboiss [87] L25-27,33,34 203 nepotulus Neboiss [77] L600,603 187 nervosa (Brauer) no figs. sp.not recognisable nesydrion (Neboiss) [77] f .86-95 26 niger Korboot - see similis Kimmins nigra (Mosely) (Notalina) [77] f.744-746 249 nigra Mosely (Taschorema) - see nigrita (Banks) nigricornis Hosely [63] L162,163 243 n i gr ita (Banks) [63] L 294; [72] ~ L138, 139 28 nigrolineata Kimmins [22] f.61 257 nivea Ulmer see edwardsi (McLachlan) niveipennis Mosely [51] L23,67 236 niveogrammicus Schmid Orig. by Schmid 273 ni veolineata Kimmins [22] L69 273 nokowoula Neboiss Orig./h-ty 58 norelus Mosely [63] L57; [77] ~ L591 186 notoptera Wise [133] L3-1O 29

obliquum (Mosely) [77] L8,9 11 obscura Kimmins (Orthotrichia) [22] L4 89 obscura Wells (Hydroptila) [120] L39,42-46 62 obscure11a (Banks) Orig./h-ty 104 obsoleta McLachlan [63] L144 242 occ u I ta Hare [40] type destroyed, not recognisable ochracea Jacquemart (Oecetis)-see asmanista Mosely ochracea Mosely (Xuthotrichia)[63]L350 73 ochraceum (Mosely) (Ethochorema)[63] L298,299 26 ochreus Mosely [77] L386-388;[63] ~ L12 165 olingoides (Ti11yard) [38] L 7-11 191 olis McFarlane [36] Ll-4 174 onychion Neboiss [77] f. 73-78 21 opora Neboiss [77] f.140-142,145 51 opposita (Walker) [63] f.184,185 245 opsia Neboiss [73] Ll-4 113 orakaivai Kimmins [22] L40 119 orba Neboiss [77] L206,207 102 orbostensis Wells [122] L23,24 87 oreolimnetes (Ti11yard) Orig. (McFarlane) 143 orkeni Illies [14] L5,6 121 ornata Kimmins [22] LSI 263 o twa yens i s Neboiss [86] L 10, 11 178 ousta Schmid OL"ig. by Schmid 255 oxypages Neboiss-[88] f.1-3 12

pa11escens (Banks) [72] L92-99 pallidus (Banks) Orig./h-ty (Morse) palpata Mosely [77] L426-432 paludosus Neboiss [77] L747-756 pansus Neboiss [83] L21,23 papi11ata Wells [123] L17-20 papuana Kimmins (Chimarra) [22] f.ll papuana Kimmins (Setodes) [22] f. 70, 71 papuana (Kimmins) (Tanorus) [22] L2

24 232 162 254 150

67 107 273

38 129 papuana Kumanski (Diplectrona) [28] L 7 ,8

papuana Kumanski (Hydropsyche) - see flynni papuanus Ulmer (Ecnomus) Orig./h-ty

Korboot 151

paranga Wells [122] L63,64 parilis Neboiss [83] L109, 110, 114 parka Mosely [63] L205 parkeri Mosely [77] L 732-735 parumbripennis McFarlane [63] L279,280 parva Banks (Smicridea) see australis Ulmer parvula (Mosely) (Smicrophylax) Orig. parvus Banks (Triplectides) [51] f .57 ,86 parvus Mosely (Nyctiophylax) [63] L246 patona Mosely [63] L22; [77]~ L529-533 patulum Neboiss [72] L20, 21 pauxi11us Neboiss [77] L658-661 pechana Mosely [83] L121,122,126 pectine11a Wells [126] L15-17 pedderensis Neboiss [77] L683-689 pe d unc ul a t urn Jacquemart [77] L118-120 pegidion Neboiss [72] L27,28;[77] ~ L28,29 penicillus Wise - see puerilis (McLacnlan) perkinsi Neboiss [72] f .188-192 pezidion Neboiss [89] L18-22 philpotti (Tillyard) [63] L226;[40] L13,21 picea (Brauer) (Abacaria) no figs available picea Kimmins (Triaenodes) [20] f.13 picea Ulmer (Dinarthropsis) [118] L 780, 781 piceus Kimmins (Polycentropus) [22] f.31 pictilis Neboiss Orig./h-ty pilosa Neboiss (Alloecella) [77] f.524-528 pilosum McFarlane (Neurochorema) [37] L42-44 plebeius Walker [63] f.87 ,88; [92] ~ f.28,30 plicata Banks (Lingora) Orig./h-ty p li cat a Mosely (Caenota) [77] L 468-479 plicosta McFarlane [36] f .44-47

92 244 270 249

34

126 233 138 188

16 217 267

96 231

25 15

52 134 123

257 169 142 228 206

42 160 189 203

49

284

plumosa (Wells) [124] L35-37 82 plutonis Banks [72] L181-186 52 pontona (Mosely) [63] L339 57 poutini McFarlane [37] L3-7 220 productus Kimmins [22] L3 55 prolatus Morse & Neboiss [51] L26,68 237 proximus Neboiss [51] L60,89 235 psaroptera McFarlane [63] rJ' f.324;[41]0 f.45 29 pseudodelicata Neboiss [83] L97-99 + 251 pseudorupina Wells [123] f.43-45 73 psocopterus (Sykora) [100] f.8 253 ptychopteryx (Brauer) [90] Orig.6"; a h-ty 222 puerilis (McLachlan [63] L248 + 146 puhia McFarlane [35] f.18-20 147 pulchella Tillyard (Pycnocentrodes) - see

aureola (McLachlan) p u lche 11 a Banks (Oecetinella) Orig. /h-ty 262 pu I cher rima Ulmer [118] L 678-682 256 pulchripenne (Tillyard) [89] Ll-3,43,44 133 pulvina Wells [121] f.35,36 76

quadrifida Wells [127] f.5,6 61

rakiura McFarlane [41] f.47-51 ramosa Jacquemart (Costora) [77] f.553 ramosa Wells (Hellyethira) [126] L9-11 rapsoni Korboot Orig./p-ty 11', 2 r aru ra ru (McFarlane) [39] L 25, 2li; [40] L14, 22 reclivatum Neboiss [72] f.13-19 regia Wells [123] L12-14 remulus Neboiss [77] f.644-646 repandus Neboiss [77] f.317 ret ic ula ta Kimmins [20] f.ll retracta Wells [124] f.19-21 rieki Neboiss [72] f.140-143 ri valis (Betten) [6] f.5 robusta Wells [120] L1-11 r onewa Mosely [77] f. 627-630 roria Mosely [63] 0' f.31;[38] ~ L13-16 rosselinus Navas [22] f.32 rossi Kimmins (Apsilochorema) [20] f.2 rossi Kimmins (Diplectrona) [63] f.233 rossi Morse & Neboiss (Triplectides)[51] f.82 rostrata Wells [122] f.25-28 rotosca Mosely [63] L28;[77] ~ f.569 rubiconum Neboiss [72] L46 - 51 ruficeps (Brauer) [61] f.11-13 rugu1um Neboiss [72] f.117-120 rupina Neboiss [123] f.37-42 russata Mosely - see dar1ingtoni (Banks) russellius Neboiss [77] f.250-252

sa b rona Kimmins [22] L14 sabronensis Kimmins [22] f.38 sagitta Neboiss [77] f.225,226 salmoni McFarlane [41] L60-64 salomonis (Kimmins) [20] f.3 saneva (Mosely) [81] f.368,369

31 184

78 119 123

14 65

218 138 265 84 28

148 60

215 191 143

11 131 240 86

185 20

120 24 72

150

107 119 101

48 58

199 sarothropus Brauer - see obsoleta (McLachlan) satana Mosely (Diplectrona) [63] f.236 131 satana Mosely (Matasia) [63] f.20;[77] ~ f.537 188 saundersi (McLachlan) [89] f.25-28,42 135 scamandra Neboiss [120] f.16-22 63 schmi d i Kimmins [22] f.13 107 scindens Neboiss [83] f.44,45,50-53 155 scirpicula Neboiss [77] f.804-807 269 scutata Wells [122] L29-32 87 secutum Neboiss [77] L83-85 26

sed 1 ac ek i Sykora [100] L 3 seductus Morse & Neboiss - see ciuskus semiauratus Mosely [63] L58 s emif la vu s Banks [79] L 66-71 sentisa Wells [121] L31,32 sen to sa Neboiss [83] L 24,25,35-37

106 seductus

186 228

79

seona (Mosely) [77] f.67-69;[72] 0 f.64,65 seposi ta Neboiss [77] L557-559 +

153 21

184 serrata Kimmins - see atra (Banks) setigera Wells [127] L19,20 62 sexpunctata (Kolenati) [5] L41-45 136 shandi (McFarlane) [63]6' L362;[41] ~ L28,29 40 signata (Banks) Orig./h-ty 108 silicis Wells [123] L21,22 66 s il vic 0 la McFarlane [63]6' L 281; [38] ~ L 50, 51 32 simi 1 i s Kimmins (Polycentropus) [22]L 25, 33 143 similis Mosely (Oeconesus) [63] L67 171 similis Mosely (Triplectides) [51] L65,93 234 simplex (Jacquemart) (Smicrophylax) Orig. 126 simplex' (Mosely) (Hellyethira)[121] L3-10 76 simulans Mosely [63] L39 203 sinuosa Kimmins (Chimarra) [22] f.22 105 sinuosa Wells (Hydroptila) [120] L49-51 62 sinuosus Kimmins (Polycentropus) [22] a" L28

[28] ~ f.37 144 situlus Korboot - see australis (Banks)(Oecetis) soena Kimmins - see seona Mosely solomonis Banks Orig./h-ty ~ 229 soror Mosely [63]6' L285;[38] ~ L54,55 36 souta Mosely [63] L196 255 spaini McFarlane [39] f.27-29 40 sparsa (Kimmins) [77] f.712-715 247 spa t ula ta McFarlane [34]6' L14, 15; [38]oL 52,53 35 spicula Neboiss [77] L 54-60 + 18 spilota Neboiss [77] L310-314 141 spinata (Banks) Orig./h-ty 130 spinosa Henderson (Cryptobiosis) [12] f.6 08 spinosa Jacquemart (Hydrobiosella) - see

cognata Kimmins spinosa Kimmins (Ecnomina) [63] f .262 152 spinosum Neboiss (Austrochorema) [72] f .193 16 squamosa Kimmins (Oecetis) [22] L49 266 squamosus Mosely (Pseudoeconesus) [63] f.73 172 stenocerca Tillyard Orig. 100 stepheni Mosely [77] L653-657 217 stigma (Ulmer) [72] L40-45 19 stipa Wells [122[ L65,66 92 straminea Mosely (Caloca) [63] L102,103 198 stramineus McLachlan (Pseudoeconesus)[63] f.71 172 striata Kimmins [22] L39 118 styliferus Jacquemart - see cralus (Mosely) styracine McFarlane [36] L24-26 37 styx McFarlane [34] L22-24 35 subalbidus Kimmins - see elongatus Banks subfusca Kimmins [22] L42a-c 120 subtriangulata Kumanski [28] L9,10 128 subulata Wells [126] L42,43 66 suteri Wells [122] L45-47 88 sylvestris McFarlane [39] L 7-9 192 syncope Neboiss [83] L88,89 247 syrdena Neboiss [73] f .6-9 112

tafana Kimmins [22] L62 258 talea Wells [127] L41,42 90 tambina Mosely [51] L40,41,76,98 238 tapena Kimmins [20] f.9 115 tasmanica Jacquemart (Diplectrona)[77] L364 129 tasmanica (Jacquemart) (Moruya) [81] L93 51 tasmanica Mosely (Hydrobiosella)[77] L216,217 101

285

tasmanica Mosely (Hydroptila) [120] L12-15 60 tasmanicum Mosely (Allochorema) [77] L19-21 14 tasmanicum (Mosely) (Ulmerochorema) [77] f .80;

[72] 0 f.52,53 19 tasmanft:us (Mosely) (Agapetus) [77] L147-151 55 tasmanicus Mosely (Plectrotarsus)[77]L367-373 181 tautoru McFarlane [37] L25-32 43 te 1 e f omini c us Kumanski [28] L 61,63,64 259 tenax Neboiss [83] L85,86 247 tenuicaudatum Tillyard [103] L9a,IOb 41 tepoka (Mosely) [63] L224;[40] L15,17,23 123 tertia Mosely [81] f.363,364 198 testacea (Navas) Orig./h-ty 116 thinotes Neboiss (78] Ll-3 154 thomasi (Wise) [40] L4,5,9,26 125 thomasoni (McFarlane) [36] L35-38 40 ti11yardi Banks (Stenopsychodes) [98] p1.2,

f.3,4,7,8 III t i 11 Y a r d i Kimmins (Notalina) [63] L 179 250 ti11yardi McFarlane (Synchorema) [37] L55-58 50 ti11yardi Mosely (Ecnomus) [63] L179 151 ti11yardi Mosely (Helicopsyche) [63] L44 221 ti11yardi Mosely (Kosrheithrus) [63] L132,133 218 ti11yardi Mosely (Paroxyethira) [63] if L349;

[29] 0 L 7 81 tipua +(McFarlane) [37] f.17-19;[40]f.16,18,24 124 tonela (Mosely) [63] L272;[12] 0 L9 100 torrenticola Korboot - see banksi+(Mosely) tortuosa Wells [122] L58-60 94 torulus Neboiss (79] L60-65 227 triangulata Sykora (Diplectrona)[100] f. 7 l28 triangulata Wells (Oxyethira) [124] Ll1-13 83 triangulatus Ulmer (Anisocentropus)[108] L31 225 tridens Henderson (Cryptobiosis) [12] f.1 98 tridens Wells (Tricholeiochiton) [125] Ll1-16 75 trifida Kimmins [20] LIS 258 trifurca McFarlane [41] f.65-69 47 trifurcata Kimmins [63] L263 155 trilobata Ki'1lIllins [20] LlO 62 tristirpis Wise [129] L5 172 tristis Banks (Macronema)- see banksi Mosely tristis Ulmer (Anisocentropus) [115] f.17,18 225 truchanasi Neboiss [77] f .379-382 182 trulla McFarlane (Zelomyia)-see mose1yi Tillyard trulla Neboiss (Ecnomina) [83] f.54-56 156 truncatus Neboiss [51] L74,96 238 tuarua McFarlane [38] f.68-70 31 tumoris Wells [127] L37,38 89 turbidum (Neboiss) [72] LI00-106 27 turgidus Neboiss [83] L20 151 turneri Mosely [63] f .32; Orig.o 196 turrita Wells [122] L33-35 + 87 tyleri Wells [122] L 70, 71 94

ulmeri (Banks) (Smicrophylax) Orig. u1meri Kimmins (Chimarra) [22] f .16 ulmeri Ross (Agapetus) Orig.

126 106

57 268

34 100 118

[63] f.211,212 267

umbra Neboiss [77] L 770-773 umbripennis McLachlan [63] f.277,278 unc i na t a Kimmins [18] L12 ung ul a ta (Ulmer) [106] L 81-83 unicolor (McLachlan) (Oecetis) unicolor Wise (Pycnocentrodes) -uranka Mosely [63] L273

see aureola (McL) 106

urbana Neboiss [89] f.4-6,39-41 urdalum Neboiss [83] a' L3; Orig.~ urimica Wells [127] L46-48 ustulata Kimmins [22] f .67

133 11 93

258

valgus Neboiss [79] f.43-48 226 vallecula Neboiss - see simplex (Mosely) variegata Mosely [86] L5,6;[77] L489,490 200 varians Mosely [63] L190; [77] L 728-730 244 varius Kimmins [51} L31,32, 71 238 vega Neboiss [77] L269-271 152 velata Wells [126] f.27-29 88 veniflex McFarlane [36] L40-42 48 venusta Kimmins - see montana Tillyard (Steno-

psychodes) verna Neboiss (Tanjistomella) [74] f.1-7;Orig.~ 179 vernale McFarlane (Rakiura) [39] f.12-16 222 vernalis Neboiss (Archaeophylax) [77]L390-392 165 vernoni Wells [126] f.5-8 79 veruta Wells [136] Ll-3 79 vesca Neboiss [77] f.580-583 189 viatica Neboiss [83] L32-34,42,43 153 victoriana Banks Orig./h-ty 132 vi 11 a Mosely [63] L155, 156 253 virgatus Neboiss [75] f .6, 7 216 vi ri dar i um Neboiss [72] L130-135 25 vitrina (Hagen) [5] L8-18 136 vittata Wells [127] L12-14 64 volda Mosely (Triaenodes) Orig. 261 volda Mosely (Triplectides) [51] f.58,87 233 volutus Neboiss [83] L63-65 138 vomerharpax McFarlane [37] L21-23 44 vunida Mosely - see fijiana Banks (Goera)

286

waddama Mosely [77] L230-233 99 waitakerensis Wise [130] L3 146 wallacei (McLachlan) type lost in mail 1984 VI walpolica Neboiss [83] L123-125 267 warisa Wells [127] L 7-9 64 warneria Mosely - see grisea Banks (Alloecella) warramunga Wells [136] L5,8,9 82 wentum Mosely [63] f.325,326a 15 wheeleri Banks [63] L186-188 246 wi nkle ri Ulmer [U8] LU9-121 148

xanthoptera McFarlane [63] L322;[41] ~ L46 30 xeni11ion Neboiss Orig./h-ty 108

yabbaca Wells [126] L30-32 93

zealandensis Mosely (Diplectrona) Orig. 128 zealandica Hudson (Helir:opsyche) [63] f .46 220 zealandica Wise (Ecnomina) [129] L1l 156 zelandensis Mosely (Oeconesus) - see maori McL. zelandica Mosely (Synchorema) - see zygoneu~a

Tillyard zimmermani Ross [93] L3 12 zonata (Neboiss) [122] LlO-13 85 zothecula Neboiss [72] f .84-90 22 zygoneura Tillyard [37] L52-54 50

atlas of trichoptera of the sw pacific — australian region ||

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