bio f4_root transport 1011

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  • 7/28/2019 Bio f4_Root Transport 1011

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    Review: Root Anatomy

    Three regions

    Meristematic

    Elongation

    Maturation

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    Root hairs and

    branching greatly

    increase the

    surface through

    which water and

    dissolved minerals

    can be absorbed.

    http://botit.botany.wisc.edu/images/130/Root/Grass_Seedling_root/Root_hair.php
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    Functions of Roots

    Absorption

    Anchorage

    Storage

    Conduction of water Hormone synthesis

    TODAY

    TOMORROW

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    Uptake ofIons by the Roots WHAT?

    Root hairs bring

    nitrates,

    phosphates,sulphates, and a

    variety of other

    essential inorganicnutrients into the

    plant

    Why are these

    nutrientsneeded?

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    Uptake of Ions by the Roots HOW? Root interception

    Root grows and intercepts ions

    Simple diffusion Ions move down their concentration gradient

    No energy expense by plant

    Example: K+

    Mass flow Bulk flow of water into the root carries ions to root

    Delivers N, Ca, Mg, S

    Active transport Ions move against their concentration gradient

    Requires a specific protein pump in the cell membrane

    Energy expense by plant

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    Water Transport into the Roots

    Water flows from

    SOIL

    EpidermisRoot CortexEndodermisXylem

    via osmosis

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    Water Transport into the Roots

    Cells are usuallyinterconnected via

    plasmodesmata.

    Their plasma forms a

    continuum (a

    symplast).

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    Water Transport into the Roots

    The transport of substances from cell to cell via

    plasmodesmata is called symplastic

    transport.

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    Water Transport into the Roots

    Apoplastic transportis when water is transported between

    cells through the extra-cellular space.

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    Water Transport into the Roots

    The endodermis

    functions as a last

    checkpoint forthe passage of

    dissolved minerals

    into the vasculartissue

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    Water Transport into the Roots

    Minerals already in the symplast when they reach the

    endodermis continue through the endodermis and

    pass into the xylem

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    Water Transport into the Roots

    Minerals that reach the

    endodermis via the

    apoplast encounter a

    waxy layer on the

    endodermis that blocks

    their entry into the

    xylem called theCasparian strip.

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    Water Transport into the Roots

    The only way to get through the endodermis is

    through the symplastic route.

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    Water Transport into the Roots

    Water and minerals

    following the

    apoplastic route

    must first cross theplasma membrane

    of the endodermis

    cells and enter thesymplast system. This ensures that no minerals

    can enter the vascular systemwith out first being checked by

    the plasma membrane

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    Water Transport into the Roots

    Once through

    the endodermis

    layer, the water

    and minerals are

    now free to

    enter the xylem

    tracheids and

    vessels.

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    Uptake of water through

    root epidermis by osmosis

    Water that crosses the

    root hair plasma membrane

    enters the symplast system

    Water that does not cross the

    root hair plasma membrane

    enters the apoplast system

    The Casparian Strip blocks

    passage of water unless it

    is moving via the symplast

    Water moves into the

    xylem for transportup the plant