c harles d arwin : t heory of e volution as a m echanistic p rocess jrmg bio1 2008
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JRMGBio1 2008
DARWIN’S IDEA OF COMMON DESCENT
Descent with modification A.K.A
EVOLUTION
common ancestor/prototype
Accumulated diverse modifications or adaptations
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DARWIN’S IDEA OF COMMON DESCENT
Life history is like a tree Common trunk: multiple
branching and re-branching
Common ancestor in each fork of branching
lineage of common descent
Extinct species
TAXONOMY: TREE OF LIFE
Carolus Linnaeus Species are fixed Ordered the great diversity of organisms into “groups
subordinate to groups” Kingdom
Phylum Class Order
Family Genus
Species
Reflected the genealogy of the tree of life
DARWIN’S IDEA OF MULTIPLICATION OF SPECIES species either split
into or bud off other species
geographical isolation of a founder species.
Founder effect. The frequency of the a allele is low in the initial population, but a small subset, in which one individual is Aa, is removed from the large population and founds a new population. The frequency of a is markedly higher in this new population, due to its relatively high frequency in the founders
DARWIN’S IDEA OF MULTIPLICATION OF SPECIES different ecological niches provide different ways
of living different plants and animals come to fill different niches with
different shapes and behaviors.
DARWIN’S IDEA OF GRADUALISM
changes through the gradual change of population rather than the sudden production of new individuals
species arise: Through gradual accumulation of adaptations to a
different environment
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Rapid evolution may arise: Isolation of small population Migration of small group in a new environment Through mass extinction Because of geological barriers
DARWIN’S IDEA OF NATURAL SELECTION Compared processes in nature with artificial selection
Developed a scientific hypothesis to explain how evolution occurs
Struggle for existence (Malthus) Members of each species compete regularly to obtain
food and living space and other necessities in life Central to his Theory of Evolution
Survival of the Fittest
key factor in the struggle for existence
Fitnessability to survive and reproduceresult of adaptationcentral to the process of evolution by natural selection
LOW FITNESS Either DIE or LEAVE FEW OFFSPRINGS
HIGH FITNESS LEVEL many OFFSPRINGS
referred to as: NATURAL SELECTIONaccumulation of changes that differentiate groups from one another, such that a new species may arise
DARWIN’S MISSING INGREDIENT
Darwin did not understand the genetic basis for variation
variations mutations genetic recombination
mutation as a raw material for evolution
OTHER THINGS TO BE CONSIDERED
Natural vs. Artificial selection
Importance of population in evolution smallest unit that can evolve Natural selection acts on individuals but INDIVIDUALS
DO NOT EVOLVE
DARWINIAN VIEW OF LIFE
Diverse forms have arisenDESCENT WITH MODIFICATION FROM
ANCESTRAL SPECIES Biological diversity
Mechanism of modification NATURAL SELECTION
PATTERNS OF EVOLUTION
Mass extinction 99% of the species ever lived are now extinct wiped out whole ecological systems* Cretaceous extinction
Large asteroid struck the earth Can be caused by eruptions of many large volcanoes (Permian
and Cretaceous) , changing of positions of continents and changing of sea levels
PATTERNS OF EVOLUTION
Adaptive Radiation single species or small group of species has evolved
into several different forms that live in different ways Ex. Darwin’s Finches
Dinosaurs Mammals
PATTERNS OF EVOLUTION Convergent Evolution
unrelated organisms come to resemble one another
Natural selection may mold different body structures structures tend to function the same way and look
similar
Analogous structures same look and function Different embryonic origin
PATTERNS OF EVOLUTION Coevolution
two species evolve in response to the changes in each other over time
Ex. Fig trees and wasps
PATTERNS OF EVOLUTION
Developmental Genes and Body Plans
Hox genes
PROCESS OF SPECIATION
Species group of organisms that can interbreed and produce a
fertile offspring Share a common gene pool
As new species evolve populations become reproductively isolated from each other
Reproductive isolation Can be: Behavioral, Geographical and Temporal
PROCESS OF SPECIATION Behavioral isolation
Capable of interbreeeding but has different courtship rituals or behavior
Ex. Eastern and western meadowlark
Geographical isolation Two populations are separated by geographic barriers Acted by natural selection Ex. Abert and Kaibab Squirrels
PROCESS OF SPECIATIONBehavioral Geographical
Temporal isolation Two or more species reproduce at the different times
PROCESS OF SPECIATION
DIANE DODD’S EXPERIMENT