chapter 3: fossils and the diversity of lifefaculty.chas.uni.edu/~groves/ehch03lecture.pdf · earth...
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Earth History, Ch. 3 1
Chapter 3:
Fossils and the diversity of life
• Art objects
• Incredible variety
– From “nanno” scale (0.001 mm) to gigantic
– From bacteria to T. rex and everything in between
• Objects of scientific study
– Age dating
– Environmental indicators
– Objective record of evolution
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Earth History, Ch. 3 2
Fossils and the diversity of life
• Our knowledge of past life is derived
mostly from fossils
• Fossil = the preserved remains or traces of a
once living organism
– Usually thousands or millions of years old (if it
still smells, it ain’t a fossil)
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Earth History, Ch. 3 3
Fossil preservation
• “Hard parts” are easily preserved
– Bones, teeth, shells or other skeletal material
• Soft tissue is rarely preserved
– This requires a burial environment in which there is no
oxygen for bacteria
• Permineralization is the filling in of void spaces
with minerals
– Spaces within plant cells can be filled with chert to
form petrified wood
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Earth History, Ch. 3 4
Fossil preservation (continued)
• A mold is a three-dimensional negative imprint of an organic structure (e.g., press your fist into clay)
• A cast is a filled-in mold—it’s an exact replica of the original structure, but without the original composition
• An impression is a two-dimensional imprint, like a very shallow mold (e.g., imprint of a leaf in mud)
• A trace fossil is the preserved evidence of animal activity (e.g., tracks, trails, burrows)
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Earth History, Ch. 3 5
Completeness of the fossil record
Death
Scaveng-
ingDecay
of soft
tissues
Decreasin
g p
robab
ility
100%
?%
Continuous removal of “potential” fossils
The probability of preservation depends on the anatomy and habitat of the
organism—marine organisms with hard parts are most likely to be preserved
Abrasion
And
transport
BurialFinal
Preser-
vation
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Earth History, Ch. 3 6
Taxonomy = the science of classification
and evolutionary relationships among groups
• Groups of organisms are called taxa
(singular, taxon)
• “Natural clusters” of taxa are called clades
– A clade is a group of taxa that share a single
common ancestor
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Earth History, Ch. 3 7
• The taxonomic hierarchy has seven levels
Kingdom (e.g., Animalia)
Phylum (e.g., Chordata)
Class (e.g., Mammalia)
Order (e.g., Primates)
Family (e.g., Hominidae)
Genus (e.g., Homo)
Species (e.g., sapiens)
Taxonomic hierarchy
Incre
asi
ng i
nclu
sive
ness
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Earth History, Ch. 3 8
Clades
Morphology continuum Morphology continuum
tim
e
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Earth History, Ch. 3 9
How are evolutionary
relationships known?
• “Cladistics” is the science of inferring
genealogical relationships among groups of
organisms
• “Cladogram” is a diagram that depicts
inferred genealogical relationships
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Earth History, Ch. 3 10
Cladistics
• All living things are
related (like cousins)
• Any two taxa share a
common ancestor
ASSUMPTIONS
memy
cousins
grandpagrandma
mysister
myparents
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Earth History, Ch. 3 11
Cladistics
Branch tips are taxabeing investigated
Node representscommon ancestor
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Earth History, Ch. 3 12
Cladistics
When more than two taxa are being considered,there are multiple possible branching arrangements
Two taxa are considered to be closely related if they share derived traits that are not present in the third taxon
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Earth History, Ch. 3 13
Cladogram of vertebrates
inve
rtebr
ates
Jawle
ss fi
sh
Shark
s, ra
ys
amphi
bian
s
Bony
fish
mam
mal
s
Turtle
s, c
rocs
birds
origin of vertebral columnVertebrata
origin of jaws with teeth; paired appendagesGnathostomata
origin of bony skeleton
Osteichthyes
Tetrapodaorigin of limbs
Amniota
Reptilia
origin of amniotic egg
origin of skull fenestrae
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Earth History, Ch. 3 14
The Six Kingdoms
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Earth History, Ch. 3 15
Prokaryotes vs. Eukaryotes
• Prokaryotes lack a well defined cell
nucleus and intracellular organelles
• Eukaryotes possess cell nucleus and
specialized organelles
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Earth History, Ch. 3 16
Prokaryotes
• Archaeobacteria are the most primitive life
forms, probably ancestral to all other life
forms—some can tolerate extremely high
temperatures and hostile chemical
environments; others can live in the absence
of oxygen
• Eubacteria are slightly more advanced—
some are capable of photosynthesis
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Earth History, Ch. 3 17
Stromatolites(built by Eubacteria)
Bacterial filaments
Modern stromatolites
Precambrian stromatolites
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Earth History, Ch. 3 18
Protists
• Single-celled or simple
multicellular organisms,
including:
– Algae
– Dinoflagellates
– Diatoms
– Calcareous nannoplankton
– Radiolarians
– Foraminifera
The preserved microfossilsof protists are extremelyuseful for dating sedimentaryrocks.
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Earth History, Ch. 3 19
Protists
dinoflagellate
Calcareous
nannoplanktondiatoms
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Earth History, Ch. 3 20
Protists
Planktonic foram
Radiolarians
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Earth History, Ch. 3 21
Fungi
• Fungi are primitive organisms that typically absord their food from dead plants and animals
– Mushrooms
– Yeasts
• Fungal spores are readily preserved—when they occur in great concentrations they may signal a catastrophic event
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Earth History, Ch. 3 22
Plants
• Non-vascular plants lack conductive tissue and reproduce by means of spores
– Mosses
• Vascular plants possess conductive tissue and reproduce by means of spores or seeds
– Ferns
– Gymnosperms (e.g., conifers)
– Angiosperms (flowering plants)
• Because their tissues are relatively tough, vascular plants have a superb fossil record!!
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Earth History, Ch. 3 23
Plants
Psilotum, the
simplest living
vascular plant
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Earth History, Ch. 3 24
Animals
• Sponges are primitive animals whose cells
are not organized into tissues
• Cnidarians have simple tissues—an inner
and outer body layer
– Jellyfish
– corals
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Earth History, Ch. 3 25
Sponges
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Earth History, Ch. 3 26
Cnidarians
(corals, jellyfish)
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Earth History, Ch. 3 27
Coelomates are more advanced
animals that possess a body cavity
• Segmented worms
• Arthropods (jointed appendages)
– Trilobites (extinct)
– Crustaceans (crabs, shrimp, lobsters)
– Insects (you know ‘em)
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Earth History, Ch. 3 28
Segmented worms; Arthopods
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Earth History, Ch. 3 29
Coelomates (continued)
• Mollusks
– Cephalopods (octopus, squid, Nautilus)
– Gastropods (snails)
– Bivalves (clams, oysters, mussels)
• Brachiopods
– Lampshells
• Bryozoans
– “Moss animals”
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Earth History, Ch. 3 30
Mollusks
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Earth History, Ch. 3 31
Bryozoa
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Earth History, Ch. 3 32
Brachiopods
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Earth History, Ch. 3 33
Coelomates (continued)
• Echinoderms
– Starfish
– Sea urchins
– Crinoids (“sea lillies”)
• Chordates
– Fish
– Amphibians
– Reptiles
– Birds
– Mammals
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Earth History, Ch. 3 34
Echinoderms
Sea urchin
Crinoid
(“sea lily”)
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Earth History, Ch. 3 35
Chordates
Amphioxus
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Earth History, Ch. 3 36
Chordates