diversity of - biomarksbiomarks.eu/sites/default/files/wiebe-light.pdf · 2014-10-29 · diversity...
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Diversity of Leptocylindrus and Tenuicylindrus in BioMarKs samples
Deepak Nanjappa (PhD student, Now PhD. SZN) Wiebe HCF Kooistra (SZN) Adriana Zingone (SZN) Stephane Audic (SBR) The whole BioMarKs team BioMarKs London Meeting
Questions: can nuclear SSU rDNA V4-sequences, obtained from BioMarKs samples, be used to
• Detect species, and
• Circumscribe the biodiversity
in the planktonic diatom genera Leptocylindrus and Tenuicylindrus?
Project BioMarKs Diversity of Leptocylindrus and Tenuicylindrus in BioMarKs samples
What are Leptocylindrus and Tenuicylindrus? • Radial centric diatom genera
• Simple morphology
Project BioMarKs Diversity of Leptocylindrus and Tenuicylindrus in BioMarKs samples
Leptocylindrus danicus
Tenuicylindrus belgicus
Why explore biodiversity of Leptocylindrus and Tenuicylindrus
• Generally at the base of the nuclear SSU rDNA tree, as
sister lineage to all the remainder of the diatoms,
• Apparently species-poor “remnant”-lineage, but species diversity is poorly known,
• Many ancestral morphological features,
• The only diatom lineage that forms resting spores immediately following sexual reproduction,
• Ecologically important and widespread.
Project BioMarKs Diversity of Leptocylindrus and Tenuicylindrus in BioMarKs samples
The nuclear SSU rDNA tree
Project BioMarKs Diversity of Leptocylindrus and Tenuicylindrus in BioMarKs samples
Achlya apiculataPhytophthora undulata
Developayella elegansPirsonia punctigeraHyphochytrium catenoides
Tetrasporopsis fuscescensGiraudyopsis stellifera
Antarctosaccion applanatumPleurochloridella botrydiopsis
Heterococcus pleurococcoidesBotrydiopsis constricta
Botrydiopsis alpinaMischococcus sphaerocephalus
Phaeothamnion confervicolaStichogloea doederleiniiChoristocarpus tenellusDiscosporangium mesarthrocarpum
Spatoglossum pacificumDictyota dichotoma
Bodanella lauterborniSporochnus comosusSaccharina japonica
Polypodochrysis teissieriPinguiochrysis pyriformis
Glossomastix chrysoplastaFibrocapsa japonica CCMP1661
Chattonella antiquaHaramonas viridis
Rhizochromulina marinaHelicopedinella tricostata
Pteridomonas danicaDictyocha fibula
Dictyocha speculumFlorenciella parvula
Verrucophora farcimenSarcinochrysis marina
Aureococcus anophagefferensPelagomonas calceolata
Pseudotetraedriella kamillaeVischeria helveticaEustigmatos magna
Synchroma grandeChrysochaete britannica
Chromophyton rosanoffiiChrysocapsa paludosa
Chrysonebula flavaHibberdia magna
Ochromonas danicaOchromonas tuberculataChrysolepidomonas dendrolepidota
Chromulina chionophilaChrysosphaera parvulaPhaeoplaca thallosaMallomonas adamasTessellaria volvocina CCMP1781
Mallomonas annulataSynura curtispina CCMP847
marine eukaryote NW414.14Uncultured marine diatom DH22_2A36
freshwater eukaryote LG22-09Uncultured stramenopile Bothnian Bay
Uncultured eukaryote DSGM-35Uncultured stramenopile 1-E8
Uncultured eukaryote A10Uncultured stramenopile 3b-A8
Marine eukaryote BLACKSEA_cl_48Bolidomonas mediterranea MINB11E5
Bolidomonas pacifica OLI31SE3Bolidomonas pacifica OLI51103Bolidomonas pacifica var. eleuthera OLI46SE
Triparma sp. TOY-0807Uncultured Bolidomonas clone MALINA_St320
Tenuicylindrus belgicus DN97S3Leptocylindrus hargravesii CCMP1856
Leptocylindrus danicusLeptocylindrus aporus p372
Leptocylindrus minimus BB951Leptocylindrus convexus B768
Hyalodiscus sp.Hyalodiscus sp. CCMP1679
Paralia sulcataStephanopyxis nipponicaStephanopyxis palmeriana CCMP814
Melosira sp.Melosira cf. octogona CCMP483
Melosira varians X85402Melosira varians CHMP7Melosira varians AJ243065Melosira varians MV13
Aulacoseira crenulataFDCCL639Aulacoseira distansAulacoseira baicalensis
Aulacoseira islandicaCHMP1Aulacoseira subarcticaWLB3
Aulacoseira ambiguaAulacoseira alpigenaCCAP1002/6Proboscia indica CCMP1896
Proboscia alata KTActinoptychus sinensis p394
Actinocyclus actinochilusCCMP107Aulacodiscus sp.
Stellarima microtrias CCMP1471Stellarima microtrias AnM0003
Coscinodiscus radiatusCoscinodiscus granii
Coscinodiscus sp. GGM-2004Rhizosolenia pungens
Rhizosolenia similoidesRhizosolenia setigera
Guinardia flaccida p788Guinardia delicatula p146
Rhizosolenia shubsoleiRhizosolenia fallaxCorethron pennatumCorethron hystrix
Corethron pennatum L29-A3bigCorethron pennatum L29-B2 pollutionCorethron inerme p534Corethron inerme L8-A1Corethron inerme L5-B5Corethron inerme L7-C4Corethron inerme L6-B5Corethron inerme L8-A4
Lampriscus kittoniiToxarium undulatumToxarium hennedyanum ECT3648
Ardissonea baculusClimacosphenia moniligera s0253
Calyptrella robustaBiddulphiopsis titiana
Trigonium formosumTerpsinoe musica
Hydrosera sp.Hemiaulus sinensis CCH-2
Acanthoceras sp. 6vii09-1AUrosolenia eriensis Y98-8Urosolenia eriensis Y98-8
Guinardia sp uncultured ECS_Sp10Eucampia antarctica
Eucampia zoodiacus CCMP386Chaetoceros muelleriiChaetoceros muellerii2
Chaetoceros didymusPhaeoceros sp.
Chaetoceros peruvianusChaetoceros rostratus
Chaetoceros calcitrans f. pumilusCCMP1315Chaetoceros radicans CCMP197
Chaetoceros socialisChaetoceros sp2
Chaetoceros neogracile ArM0004Chaetoceros sp.
Chaetoceros muelleriCCMP1316Chaetoceros gracilisChaetoceros debilis Ch4Chaetoceros curvisetis ch5
Brockmanniella brockmannii CCMP151Papiliocellulus elegans
Minutocellus polymorphus CCMP497Minutocellus sp. RCC967
Leyanella arenariaExtubocellulus cribriger
Cymatosira belgicaOdontella sinensis
Pleurosira laevisOdontella aurita CCAP1054.1
Lithodesmium undulatumDitylum brightwelliiHelicotheca tamesisBellerochea malleus
Porosira pseudodenticulataLauderia annulata
P. pseudodenticulataThalassiosira gessneri ANO2-08
Thalassiosira weissflogii CCMP1010Thalassiosira pseudonana CCMP1057Detonula confervacea
Cyclotella menegheniana WC03-01Cyclotella striata CCMP1586Skeletonema menzelii B82Skeletonema grevilleii CCMP1685Skeletonema dohrnii B104
Skeletonema costatum B211Skeletonema grethae CCMP780Skeletonema pseudocostatumSkeletonema tropicum CCMP2070
Thalassiosira oceanica CCMP1001Minidiscus trioculatus CCMP495Discostella pseudostelligera ROR01-1Cyclostephanos bodanica J98-1
Stephanodiscus hantzschii WTC21Stephanodiscus niagarae OKA-AThalassiosira anguste_lineata MHta1Planktoniella sol p354Detonula pumila NB48Thalassiosira tumida CCMP1469Shionodiscus ritscheri LC01-12Bacterosira bathyomphala NB04-B6
Thalassiosira eccentricaThalassiosira rotula
Attheya cf. septentrionalis strain CCMP2084Attheya longicornis CCMP214
Podocystis spathulata ECT3733Delphineis sp. CCMP1095
Rhaphoneis cf. belgicaRhaphoneis amphiceros
Asteroplanus karyanaAsterionellopsis glacialisAsterionellopsis glacialis WK20
Plagiogramma atomus s0330Neofragilaria nicobarica s0371
Talaroneis posidoniae WK59AB430598.1.1862_U Dimeregramma;Dimeregramma_minor_var._nanumPsammoneis senegalensis s0387Psammoneis japonica s0305Psammoneis pseudojaponica s0354
Araphid pennate sp1.Araphid pennate sp2.
Synedropsis cf.recta CCMP1620Fragilaria barbararum ROSD99Grammonema striatula CCAP1029.18Grammonema islandicaGrammonema striatula CCMP1094
Synedropsis minuscula CCMP845Synedropsis hyperborea CCMP1423Synedropsis fragilaroides CCMP844Grammonema sp.
Hyalosynedra cf.laevigata WK52Catacombas gaillonii WK37
Tabularia fasciculata L778Tabularia tabulata CCMP846Synedra toxoneides WK57
Thalassionema bacillare WK54Thalassionema nitzschioidesThalassionema frauenfeldii WK3Thalassionema frauenfeldii WK5
Synedra marineSynedra sp. p517
Ulnaria ulna WK36Synedra acus At182gel3Synedra angustissima D-16
Fragilaria vaucheriae L1496Fragilaria crotonensisFragilaria rumpens At114Fragilaria capucina D113Fragilariforma virescens V18Fragilaria bidens At170.4
Fragilaria delicatissima At135.13Hyalosira sp. WK50Hyalosira sp. CCMP469Hyalosira delicatula WK51
Grammatophora oceanicaGrammatophora oceanica WK72Grammatophora gibberula WK48Grammatophora marina WK43Grammatophora oceanica CCMP410
Rhabdonema arcuatumLicmophora reichardtii WK63
Licmophora grandis WK62DN82S1 CCMP470 pennate deepak
Synedra berolinensis L1722Staurosira construens
Staurosirella pinnata D91Staurosira elliptica L948
Nanofrustulum shiloi CCMP1306Pseudostaurosira zeilleri v.elliptica D48
Pseudostaurosiropsis sp. I-01Diatoma hyemalis
Asterionella formosaTabellaria flocculosa L1839Diatoma vulgare L1535Diatoma tenue L1251Diatoma tenue v.elongatum L1346Staurosira subsalinaCyclophora tenuisLicmophora juergensii P367
Licmophora flucticulataLicmophora abbreviata
Licmophora gracilisConvoluta endosymbiont
Licmophora communisLicmophora flabellata WK58Licmophora flabellata WK47Licmophora flabellata
Pseudostriatella pacifica s0384Striatella unipunctata
Protoraphis atlantica WK42Pseudohimantidium pacificum
Amphora laevis v.laevissimaGU385679.1.1033_U Bacillariophyta_XXX_sp.
Eunotia monodon v.asiaticaEunotia pectinalisEunotia formica v.sumatrana
Eunophora oberonicaActinella pulchella
Actinella parvaUndatella sp.
AY180015.1.1715_U Bacillariophyta_XXX_sp.Gyrosigma limosum C092Pleurosigma planktonicum C090
Pleurosigma intermedium C089Pleurosigma sp.
Haslea nipkowii C083Haslea crucigera C082Haslea ostrearia C079Haslea pseudostrearia C081
GU385626.1.1698_U Bacillariophyta_XXX_sp.GU385631.1.1691_U Bacillariophyta_XXX_sp.
GU385685.1.1700_U Bacillariophyta_XXX_sp.GU385690.1.1340_U Bacillariophyta_XXX_sp.
GU385609.1.1711_U Bacillariophyta_XXX_sp.GU385639.1.1696_U Bacillariophyta_XXX_sp.
GU385674.1.1694_U Bacillariophyta_XXX_sp.Navicula lanceolata C086
Navicula arenaria v.rostellataNavicula perminuta
Navicula gregaria AT-117Gel05Navicula sclesviscensis C085
Navicula cryptocephala v.venetaPseudogomphonema sp.
Navicula phyllepta HPSeminavis robusta
Navicula salinarumHQ912404.1.1728_U Hantzschia;Hantzschia_amphioxys
JN033248.1.1192_U Cymbella;Cymbella_cistuliformisPseudo-nitzschia pungens
Pseudo-nitzschia multiseriesGU823436.1.1207_U Bacillariophyta_XXX_sp.
Fragilariopsis cylindricus CCMP1102Fragilariopsis sublinearis
JF791075.1.1777_U Nitzschia;Nitzschia_apiculataAchnanthes bongrainii
Achnanthidium cf. longipesAchnanthes brevipes CCMP100Psammodictyon pandurifore UTCC162
Bacillaria paxilliferTryblionella constrictaNitzschia cf. frustulum
HM799827.1.1722_U Bacillariophyta_XXX_sp.Durinskia baltica endosymbiont
Nitzschia thermalis HPKryptoperidinium foliaceum endosymbiont
FN598376.1.1059_U Bacillariophyta_XXX_sp.Cylindrotheca fusiformis CCMP339
EU333092.1.800_U Bacillariophyta_XXX_sp.Cylindrotheca closterium
Cylindrotheca closterium2FN690543.1.842_U Bacillariophyta_XXX_sp.
HQ912597.1.1724_U Diploneis;Diploneis_subovalisGU823641.1.1061_U Bacillariophyta_XXX_sp.
GU822956.1.1203_U Bacillariophyta_XXX_sp.GU385662.1.1670_U Bacillariophyta_XXX_sp.
HQ680564.1.876_U Cymbella;Cymbella_pelagicaHQ680563.1.876_U Cymbella;Cymbella_parva
HQ912644.1.1591_U Climaconeis;Climaconeis_riddleaeLyrella atlantica
HQ912413.1.1746_U Campylodiscus;Campylodiscus_sp.HQ912609.1.1719_U Scoliopleura;Scoliopleura_peisonis
HQ912591.1.1742_U Neidium;Neidium_bisulcatumHQ912583.1.1742_U Neidium;Neidium_affine
HQ912582.1.1733_U Neidium;Neidium_productumAM501955.1.1727_U Neidium;Neidium_affine
Anomoeoneis sphaerophora L1222Encyonema angulatum L1313
Gomphonema parvulumGomphonema gracile
AM502016.1.1724_U Cymbella;Cymbella_asperaHQ655885.1.1724_U Reimeria;Reimeria_sinuataHQ655880.1.1724_U Cymbella;Cymbella_navicula
Phaeodactylum tricornutumCocconeis placentula AT-212Gel11
HQ912626.1.1750_U Lemnicola;Lemnicola_hungaricaPauliella taeniata CCMP1115
Dickieia ulvaceaPlanothidium lanceolatum L1249
Diadesmis gallicaDQ310321.1.1054_U Bacillariophyta_XXX_sp.
HQ912637.1.1736_U Berkeleya;Berkeleya_rutilansDQ310229.1.1384_U Bacillariophyta_XXX_sp.DQ310238.1.1381_U Bacillariophyta_XXX_sp.Biremis ambigua
Brachysira cf. styriacaParlibellus cf.plicatus
Navicula pelliculosa CCMP543Eolimna subminuscula
Pinnularia microstauron AT-69.06Pinnularia acrosphaeria AT-161.03
Caloneis amphisbaena AT-177.07Pinnunavis cf.elegans
Fallacia forcipataHQ912605.1.1745_U Fallacia;Fallacia_pygmaea
Amphora coffeaeformis CCAP1008/1Eolimna minimaSellaphora auldreekie DUN1
Sellaphora pupula GER1Amphora lybica AT-117.10Amphora cf. proteus p386Amphora.montana
Amphora cf. capitellata p363Amphora cf. coffeaeformis CCAP1008/1
HQ912415.1.1727_U Surirella;Surirella_splendidaStenopterobia cf.curvataHQ912416.1.1725_U Stenopterobia;Stenopterobia_curvula
HQ912414.1.1752_U Surirella;Surirella_sp.Amphiproa paludosa CCMP125
Amphiprora alata CCAP1003/3Entomoneis cf. alata CCMP1522
Surirella sp.HQ912658.1.1741_U Surirella;Surirella_minuta
Surirella angusta SANG1Surirella brebissoni SBRE3
0.01 substitutions/site
NJ
Heterotrophic Stramenopiles
“other” autotrophic Stramenopiles
Bolidomonads and Parmales
Other Radial Centrics
Bi- and Multipolar Centrics
Pennates
T. belgicus L. hargravesii
L. danicus L. aporus
L. minimus L. convexus
Project BioMarKs Diversity of Leptocylindrus and Tenuicylindrus in BioMarKs samples
2µm
Triangular “wings”
Annulus
Subcentral pore
Striae
Valve Valve face
Valve mantle
Chain of cells
Butterfly shaped cingular band
Leptocylindrus
Project BioMarKs Diversity of Leptocylindrus and Tenuicylindrus in BioMarKs samples
Tenuicylindrus
Scale Bar: A–B 5µm C–J 1µm
Project BioMarKs Diversity of Leptocylindrus and Tenuicylindrus in BioMarKs samples
Basilicostephanus
Harwood and Gersonde, 1990 Gersonde and Harwood, 1990
Gladiopsis
Kerkis
Rhynchopyxis
Project BioMarKs Diversity of Leptocylindrus and Tenuicylindrus in BioMarKs samples
Scale bars 1µm
Leptocylindrus danicus
Leptocylindrus danicus, L. hargravesii and L minimus form resting spores immediately upon sexual reproduction.
Project BioMarKs Diversity of Leptocylindrus and Tenuicylindrus in BioMarKs samples
L. aporus L. danicus L. dan
L. convexus
L.h L.h
Diversity and abundance of Leptocylindrus spp. in the plankton of the Gulf of Naples
Project BioMarKs Diversity of Leptocylindrus and Tenuicylindrus in BioMarKs samples
Diversity and abundance of Tenuicylindrus belgicus in the plankton of the Gulf of Naples
Can we use BioMarKs V4 sequence data gathered all across European coasts to assess the biodiversity of these genera?
Two questions: • Can we find the same species in other places?
• Can we find yet undiscovered diversity within the two genera?
Project BioMarKs Diversity of Leptocylindrus and Tenuicylindrus in BioMarKs samples
Can we use BioMarKs V4 sequence data gathered all across European coasts to assess the biodiversity of these genera?
Two questions: • Can we find the same species in other places?
– Yes! Based on highly similar or identical V4-sequences
Project BioMarKs Diversity of Leptocylindrus and Tenuicylindrus in BioMarKs samples
Project BioMarKs Diversity of Leptocylindrus and Tenuicylindrus in BioMarKs samples
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Naples 2009 Naples 2010 Oslo 2009 Oslo 2010 Barcelona 2010 Gijon 2010
Leptocylindrus minimus Leptocylindrus danicus Tenuicylindrus belgicus Leptocylindrus convexus Leptocylindrus aporus
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Naples 2009 Naples 2010 Oslo 2009 Oslo 2010 Barcelona 2010 Gijon 2010
Leptocylindrus minimus Leptocylindrus danicus Tenuicylindrus belgicus Leptocylindrus convexus Leptocylindrus aporus
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Naples 2009 Naples 2010 Oslo 2009 Oslo 2010 Barcelona 2010 Gijon 2010
Leptocylindrus minimus Leptocylindrus danicus Tenuicylindrus belgicus Leptocylindrus convexus Leptocylindrus aporus1
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Naples 2009 Naples 2010 Oslo 2009 Oslo 2010 Barcelona 2010 Gijon 2010
Leptocylindrus minimus Leptocylindrus danicus Tenuicylindrus belgicus Leptocylindrus convexus Leptocylindrus aporus
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Naples 2009 Naples 2010 Oslo 2009 Oslo 2010 Barcelona 2010 Gijon 2010
Leptocylindrus minimus Leptocylindrus danicus Tenuicylindrus belgicus Leptocylindrus convexus Leptocylindrus aporus
Leptocylindrus dancius/hargravesii
Leptocylindrus minimus
Tenuicylindrus belgicus
Leptocylindrus convexus
Leptocylindrus aporus
Can we use BioMarKs V4 sequence data gathered all across European coasts to assess the biodiversity of these genera?
Two questions: • Can we find the same species in other places?
• Can we find yet undiscovered diversity within the two genera? – Yeah, but with some trouble!
Project BioMarKs Diversity of Leptocylindrus and Tenuicylindrus in BioMarKs samples
The V4-tree
Pirsonia punctigeraHyphochytrium catenoidesDevelopayella elegans
Achlya apiculataPhytophthora undulata
Fibrocapsa japonica CCMP1661Chattonella antiqua
Haramonas viridisPolypodochrysis teissieri
Pinguiochrysis pyriformisGlossomastix chrysoplasta
Dictyocha fibulaSynchroma grandeSarcinochrysis marina
Aureococcus anophagefferensPelagomonas calceolata
Florenciella parvulaVerrucophora farcimen
Dictyocha speculumRhizochromulina marina
Helicopedinella tricostataPteridomonas danica
Pseudotetraedriella kamillaeVischeria helveticaEustigmatos magna
Ochromonas danicaChrysolepidomonas dendrolepidota
Chromulina chionophilaChrysosphaera parvula
Mallomonas adamasOchromonas tuberculata
Phaeoplaca thallosaTessellaria volvocina CCMP1781
Mallomonas annulataSynura curtispina CCMP847
Tetrasporopsis fuscescensGiraudyopsis stelliferaAntarctosaccion applanatum
Pleurochloridella botrydiopsisStichogloea doederleinii
Botrydiopsis alpinaBotrydiopsis constricta
Heterococcus pleurococcoidesMischococcus sphaerocephalus
Phaeothamnion confervicolaChoristocarpus tenellus
Discosporangium mesarthrocarpumDictyota dichotomaBodanella lauterborni
Spatoglossum pacificumSporochnus comosusSaccharina japonica
Bolidomonas mediterranea MINB11E5marine eukaryote NW414.14
Uncultured marine diatom DH22_2A36Uncultured eukaryote DSGM-35
Uncultured stramenopile 1-E8Uncultured eukaryote A10
Uncultured stramenopile 3b-A8freshwater eukaryote LG22-09
Uncultured stramenopile Bothnian BayMarine eukaryote BLACKSEA_cl_48
Bolidomonas pacifica OLI31SE3Bolidomonas pacifica OLI51103
Bolidomonas pacifica var. eleuthera OLI46SETriparma sp. TOY-0807Uncultured Bolidomonas clone MALINA_St320
Tenuicylindrus belgicus DN97S3Porosira pseudodenticulata
Lauderia annulataLeptocylindrus hargravesii CCMP1856Leptocylindrus danicus
Leptocylindrus aporus p372Leptocylindrus minimus BB951
Leptocylindrus convexus B768Aulacodiscus sp.
Actinoptychus sinensis p394Actinocyclus actinochilusCCMP107Chrysochaete britannica
Chromophyton rosanoffiiChrysocapsa paludosa
Chrysonebula flavaHibberdia magna
HQ680564.1.876_U Cymbella;Cymbella_pelagicaHQ680563.1.876_U Cymbella;Cymbella_parva
Melosira sp.Melosira cf. octogona CCMP483
Melosira varians X85402Melosira varians CHMP7Melosira varians AJ243065Melosira varians MV13
Terpsinoe musicaHyalodiscus sp. CCMP1679
Hyalodiscus sp.Rhizosolenia pungensRhizosolenia similoides
Corethron pennatumCorethron hystrix
Corethron pennatum L29-A3bigCorethron pennatum L29-B2 pollutionCorethron inerme p534Corethron inerme L5-B5
Corethron inerme L8-A4Corethron inerme L8-A1Corethron inerme L7-C4Corethron inerme L6-B5
Lampriscus kittoniiToxarium undulatum
Toxarium hennedyanum ECT3648Ardissonea baculusClimacosphenia moniligera s0253
Hydrosera sp.Acanthoceras sp. 6vii09-1AUrosolenia eriensis Y98-8
Urosolenia eriensis Y98-8Hemiaulus sinensis CCH-2
Guinardia sp uncultured ECS_Sp10Lithodesmium undulatum
Bellerochea malleusHelicotheca tamesisDitylum brightwellii
Chaetoceros didymusPhaeoceros sp.Chaetoceros peruvianus
Chaetoceros rostratusChaetoceros muellerii
Chaetoceros muellerii2Chaetoceros calcitrans f. pumilusCCMP1315
Chaetoceros radicans CCMP197Chaetoceros socialis
Chaetoceros sp2Chaetoceros neogracile ArM0004Chaetoceros sp.
Chaetoceros muelleriCCMP1316Chaetoceros gracilisChaetoceros debilis Ch4Chaetoceros curvisetis ch5
Attheya cf. septentrionalis strain CCMP2084Attheya longicornis CCMP214Paralia sulcata
Stephanopyxis nipponicaStephanopyxis palmeriana CCMP814
Thalassiosira gessneri ANO2-08Thalassiosira weissflogii CCMP1010
Thalassiosira pseudonana CCMP1057Detonula confervaceaCyclotella menegheniana WC03-01Cyclotella striata CCMP1586Skeletonema menzelii B82
Skeletonema grethae CCMP780Skeletonema grevilleii CCMP1685Skeletonema costatum B211Skeletonema dohrnii B104
Skeletonema pseudocostatumSkeletonema tropicum CCMP2070
Discostella pseudostelligera ROR01-1Cyclostephanos bodanica J98-1
Stephanodiscus hantzschii WTC21Stephanodiscus niagarae OKA-ABacterosira bathyomphala NB04-B6
Thalassiosira eccentricaThalassiosira rotulaThalassiosira anguste_lineata MHta1Detonula pumila NB48
Planktoniella sol p354Thalassiosira tumida CCMP1469
Minidiscus trioculatus CCMP495Thalassiosira oceanica CCMP1001Shionodiscus ritscheri LC01-12
Stellarima microtrias CCMP1471Stellarima microtrias AnM0003Trigonium formosum
Eucampia antarcticaEucampia zoodiacus CCMP386
Calyptrella robustaCoscinodiscus radiatus
Coscinodiscus graniiCoscinodiscus sp. GGM-2004
Proboscia indica CCMP1896Proboscia alata KT
Rhizosolenia setigeraGuinardia flaccida p788
Pleurosira laevisOdontella aurita CCAP1054.1
Guinardia delicatula p146Rhizosolenia shubsolei
Rhizosolenia fallaxBiddulphiopsis titiana
Odontella sinensisP. pseudodenticulata
Brockmanniella brockmannii CCMP151Leyanella arenaria
Extubocellulus cribrigerCymatosira belgica
Papiliocellulus elegansMinutocellus polymorphus CCMP497Minutocellus sp. RCC967
GU385679.1.1033_U Bacillariophyta_XXX_sp.Aulacoseira crenulataFDCCL639
Aulacoseira distansAulacoseira ambiguaAulacoseira subarcticaWLB3Aulacoseira alpigenaCCAP1002/6Aulacoseira baicalensisAulacoseira islandicaCHMP1
Podocystis spathulata ECT3733HQ912644.1.1591_U Climaconeis;Climaconeis_riddleaeGomphonema parvulum
Gomphonema gracileAnomoeoneis sphaerophora L1222
Pinnularia acrosphaeria AT-161.03HQ912413.1.1746_U Campylodiscus;Campylodiscus_sp.
Lyrella atlanticaAM502016.1.1724_U Cymbella;Cymbella_asperaHQ655885.1.1724_U Reimeria;Reimeria_sinuataHQ655880.1.1724_U Cymbella;Cymbella_navicula
HQ912609.1.1719_U Scoliopleura;Scoliopleura_peisonisHQ912591.1.1742_U Neidium;Neidium_bisulcatumHQ912583.1.1742_U Neidium;Neidium_affine
HQ912582.1.1733_U Neidium;Neidium_productumAM501955.1.1727_U Neidium;Neidium_affine
Undatella sp.Haslea nipkowii C083
Haslea crucigera C082Haslea ostrearia C079Haslea pseudostrearia C081
AY180015.1.1715_U Bacillariophyta_XXX_sp.Gyrosigma limosum C092
Pleurosigma planktonicum C090Pleurosigma intermedium C089Pleurosigma sp.
Araphid pennate sp1.Pseudogomphonema sp.
Navicula phyllepta HPNavicula gregaria AT-117Gel05Navicula sclesviscensis C085
GU385674.1.1694_U Bacillariophyta_XXX_sp.Navicula lanceolata C086Navicula arenaria v.rostellataNavicula perminuta
Navicula cryptocephala v.venetaSeminavis robustaNavicula salinarum
GU823641.1.1061_U Bacillariophyta_XXX_sp.Araphid pennate sp2.
HQ912637.1.1736_U Berkeleya;Berkeleya_rutilansDQ310238.1.1381_U Bacillariophyta_XXX_sp.
DQ310229.1.1384_U Bacillariophyta_XXX_sp.GU385690.1.1340_U Bacillariophyta_XXX_sp.
Cocconeis placentula AT-212Gel11Encyonema angulatum L1313
GU385626.1.1698_U Bacillariophyta_XXX_sp.Planothidium lanceolatum L1249
HQ912626.1.1750_U Lemnicola;Lemnicola_hungaricaPauliella taeniata CCMP1115Dickieia ulvacea
HQ912597.1.1724_U Diploneis;Diploneis_subovalisDiadesmis gallica
Amphora coffeaeformis CCAP1008/1Eolimna minimaSellaphora auldreekie DUN1
Sellaphora pupula GER1Fallacia forcipata
HQ912605.1.1745_U Fallacia;Fallacia_pygmaeaBiremis ambigua
Pinnularia microstauron AT-69.06Caloneis amphisbaena AT-177.07Pinnunavis cf.elegans
Amphora cf. coffeaeformis CCAP1008/1Amphora.montana
Amphora cf. capitellata p363Amphora lybica AT-117.10
Amphora cf. proteus p386Parlibellus cf.plicatus
Navicula pelliculosa CCMP543Eolimna subminuscula
HQ912415.1.1727_U Surirella;Surirella_splendidaStenopterobia cf.curvata
HQ912416.1.1725_U Stenopterobia;Stenopterobia_curvulaHQ912414.1.1752_U Surirella;Surirella_sp.
Amphiproa paludosa CCMP125Amphiprora alata CCAP1003/3
Entomoneis cf. alata CCMP1522Surirella sp.
HQ912658.1.1741_U Surirella;Surirella_minutaSurirella angusta SANG1Surirella brebissoni SBRE3
HQ912404.1.1728_U Hantzschia;Hantzschia_amphioxysJN033248.1.1192_U Cymbella;Cymbella_cistuliformis
GU823436.1.1207_U Bacillariophyta_XXX_sp.Fragilariopsis cylindricus CCMP1102Fragilariopsis sublinearisPseudo-nitzschia pungens
Pseudo-nitzschia multiseriesFN598376.1.1059_U Bacillariophyta_XXX_sp.GU385662.1.1670_U Bacillariophyta_XXX_sp.
GU385685.1.1700_U Bacillariophyta_XXX_sp.Eunotia monodon v.asiaticaEunotia pectinalis
Eunotia formica v.sumatranaEunophora oberonica
Actinella pulchellaActinella parvaCylindrotheca closteriumCylindrotheca closterium2
Cylindrotheca fusiformis CCMP339FN690543.1.842_U Bacillariophyta_XXX_sp.
EU333092.1.800_U Bacillariophyta_XXX_sp.Amphora laevis v.laevissimaAchnanthes bongrainii
Psammodictyon pandurifore UTCC162GU385639.1.1696_U Bacillariophyta_XXX_sp.
Bacillaria paxilliferAchnanthidium cf. longipes
Achnanthes brevipes CCMP100Tryblionella constricta
Kryptoperidinium foliaceum endosymbiontNitzschia cf. frustulumDurinskia baltica endosymbiontGU385631.1.1691_U Bacillariophyta_XXX_sp.
GU385609.1.1711_U Bacillariophyta_XXX_sp.Nitzschia thermalis HP
GU822956.1.1203_U Bacillariophyta_XXX_sp.HM799827.1.1722_U Bacillariophyta_XXX_sp.
Phaeodactylum tricornutumDQ310321.1.1054_U Bacillariophyta_XXX_sp.
JF791075.1.1777_U Nitzschia;Nitzschia_apiculataHyalosira sp. WK50Hyalosira sp. CCMP469Hyalosira delicatula WK51
Brachysira cf. styriacaProtoraphis atlantica WK42
Pseudohimantidium pacificumSynedra marine
Synedra sp. p517Ulnaria ulna WK36Synedra acus At182gel3Synedra angustissima D-16
Thalassionema nitzschioidesThalassionema bacillare WK54Thalassionema frauenfeldii WK3
Thalassionema frauenfeldii WK5Hyalosynedra cf.laevigata WK52
Synedra toxoneides WK57Catacombas gaillonii WK37
Tabularia fasciculata L778Tabularia tabulata CCMP846
Fragilaria vaucheriae L1496Fragilaria crotonensisFragilaria rumpens At114Fragilaria capucina D113
Fragilariforma virescens V18Fragilaria bidens At170.4Fragilaria delicatissima At135.13
Pseudostriatella pacifica s0384Striatella unipunctata
Licmophora flucticulataLicmophora abbreviata
Licmophora gracilisLicmophora grandis WK62Convoluta endosymbiontLicmophora communis
Rhabdonema arcuatumGrammonema striatula CCAP1029.18
Grammonema islandicaSynedropsis cf.recta CCMP1620Grammonema striatula CCMP1094Fragilaria barbararum ROSD99Synedropsis minuscula CCMP845Synedropsis hyperborea CCMP1423Synedropsis fragilaroides CCMP844Grammonema sp.
Grammatophora oceanicaGrammatophora oceanica CCMP410Grammatophora oceanica WK72Grammatophora marina WK43Grammatophora gibberula WK48
Cyclophora tenuisLicmophora reichardtii WK63
DN82S1 CCMP470 pennate deepakLicmophora juergensii P367Licmophora flabellataLicmophora flabellata WK47Licmophora flabellata WK58
Asteroplanus karyanaAsterionellopsis glacialisAsterionellopsis glacialis WK20
Delphineis sp. CCMP1095Rhaphoneis cf. belgica
Rhaphoneis amphicerosStaurosirella pinnata D91
Staurosira construensSynedra berolinensis L1722Staurosira elliptica L948Nanofrustulum shiloi CCMP1306
Pseudostaurosira zeilleri v.elliptica D48Pseudostaurosiropsis sp. I-01
AB430598.1.1862_U Dimeregramma;Dimeregramma_minor_var._nanumTalaroneis posidoniae WK59
Plagiogramma atomus s0330Neofragilaria nicobarica s0371Psammoneis japonica s0305Psammoneis pseudojaponica s0354Psammoneis senegalensis s0387
Asterionella formosaDiatoma hyemalis
Diatoma vulgare L1535Tabellaria flocculosa L1839Staurosira subsalinaDiatoma tenue L1251Diatoma tenue v.elongatum L1346
0.01 substitutions/site
NJ
L. hargravesii L. danicus
L. aporus L. minimus
L. convexus
Project BioMarKs Diversity of Leptocylindrus and Tenuicylindrus in BioMarKs samples
Heterotrophic Stramenopiles
“most” autotrophic Stramenopiles
Bolidomonads and Parmales
Other Radial Centrics and a few Stramenopiles
Bi-Multipolar, and a few radial Centrics
Pennates
Porosira & Lauderia T. belgicus
10% dissimilarity
Ldan_382b87d93ce960f24faeee43512a1758_1 Lharg dcefcbb168cf68d9d61b94967207ed0b_1 Lharg 81e5d1a40e3b1f2e6e76ff87a5fad302_2 Ldan_49de97debf5a4bdf34a02e745eea4331_1 Lharg_2807c3131b3356d4d03c8a015bf1e64b_1 Lharg 5d681a0f9e2f894241f70363d81e50b1_1
Lharg 847f436624de6a861121d8fdcbcbd035_1 Lharg_294c718549e6d44740195c3020cbbdad_2 Ldan_403d4e8bef56dc568bb0dd04619a6709_2
Lharg 48a4f876508ee8b7e69fe60741c33e6b_1 Ldan_fb349487abc6abf2bfe46e42c95db8ca_1
Lharg_2a68188090520caa8d81c45a0a443188_4 GU824887
Lharg_ff503d2534c999af974bf427c46df44e_8 Leptocylindrus hargravesii Lharg Leptocylindrus hargravesii_635 GU825201
GU824332 Lharg_76607d003c01e727ee17b4630cbf842a_1
GU825183 Leptocylindrus danicus Ldan Leptocylindrus danicus_68
GU824143 Leptocylindrus convexus_17 Leptocylindrus convexus GU823919
GU825668 Leptocylindrus minimus NCBI
Lmini_22170c8c5b9a7a1daf53f4d7066987f0_185 Lcon_355d80c0a1f0b317384480a48cae4cb0_42
Lmini 42da7abc101a949c189137e89ca987ab_16 Lmini 64e6ca789c83f3a1258cc18cd8defb78_3
Lapo_e9dede35a7076885146edd819db18e25_1 Naples2009 Lapo_a88ade6550b62188339dbf44ab3e56c3_24
Lapo_12388426d16bccda4ab619cd18fe6792_1 Lapo_Leptocylindrus aporus_48820
Lapo_e93487fb115d07d82421cc106f2a9283_1 Lapo_442fa2332449e0593cd2d47b6f3ed7f6_5
Lapo_8b3d183186a056081e73ef80cf8c53fe_3 Lapo_bab055b2f4b42689a9490af9e726fed3_20
Lapo_987d26ffc34dbf1088def1f5e9cc6190_1 Lapo_c90e2b3ba682d9e90c980689ae962986_1
Tbel d879885476c520772996a2a230446060_1 Tbel 6755f0b4633e96ebb8ea275997fd3935_1
Tbel 0b38196304d4cf65273e7800b2b1494a_1 Tbel c20e86a160753ce12126e809c385537a_1
Tbel 37ae65a8517aa36e460bf8812a378802_1 Tbel c9b3c284663d66bd56fed02b5cec97dc_1
Tbel bc86d8d21a404eabe412968d81b0ab85_1 Tenuicylindrus belgicus Tbel_0577c4678c8eec0a9ad9693527326f03_165
Other Centrics
L. hargravesii and L. danicus
L. aporus
L. minimus
L. convexus
T. belgicus
L. something new?
Bolidomonas
10% dissimilarity