ecosystem ecology. i. introduction - ecosystem: an assemblage of organisms, together with their...

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Ecosystem Ecology

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Page 1: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

Page 2: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

- Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Page 3: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

Page 4: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A. Productivity

1. Gross Primary Productivity

Total photosynthetic productivity:

CO2 + H20 -----> Glucose + O2

Page 5: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A. Productivity

1. Gross Primary Productivity

Total photosynthetic productivity:

CO2 + H20 -----> Glucose + O2

Metabolism

Growth

Reproduction

Page 6: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

1. Gross Primary Productivity

Total photosynthetic productivity:

CO2 + H20 -----> Glucose + O2

Ecosystem Ecology

I. Introduction

II. Energy Flow

A. Productivity

Page 7: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

2. Net Primary Productivity:

- energy stored in biomass

Ecosystem Ecology

I. Introduction

II. Energy Flow

A. Productivity

Page 8: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A. Productivity

2. Net Primary Productivity:

- energy stored in biomass

- measurements

Page 9: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A. Productivity

2. Net Primary Productivity:

- energy stored in biomass

- measurements

- factors affecting NPP

Page 10: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A. Productivity

2. Net Primary Productivity:

- energy stored in biomass

- measurements

- factors affecting NPP

- light

Page 11: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A. Productivity

2. Net Primary Productivity:

- energy stored in biomass

- measurements

- factors affecting NPP

- light

- water

Page 12: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A. Productivity

2. Net Primary Productivity:

- energy stored in biomass

- measurements

- factors affecting NPP

- light

- water

- temp

Page 13: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A. Productivity

2. Net Primary Productivity:

- energy stored in biomass

- measurements

- factors affecting NPP

- light

- water

- temp

- nutrients

Page 14: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

2. Net Primary Productivity:

- factors affecting NPP

- Nutrients: Nutrient Use Efficiency =

grams of dry mass produced/gram of nutrient absorbed

Lower NUE for a nutrient means it is more limiting (need more to produce the same biomass).

Page 15: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

2. Net Primary Productivity:

- energy stored in biomass

- measurements

- factors affecting NPP

- light

- water

- temp

- nutrients

- Global Patterns

Page 16: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments
Page 17: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

2. Net Primary Productivity:

- energy stored in biomass

- measurements

- factors affecting NPP

- light

- water

- temp

- nutrients

- Global Patterns

Page 18: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments
Page 20: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A. Productivity

3. Net Secondary Productivity

- assimilations efficiencies – A/I

seed eaters: 60-80%

browsers: 30-40%

detritivores: 15%

Page 21: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A. Productivity

3. Net Secondary Productivity

- assimilations efficiencies – A/I

seed eaters: 60-80%

browsers: 30-40%

detritivores: 15%

herbivores: 60-70%

carnivores: 80-90%

Page 22: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Low AE? Must eat more to get energy needed.

Horse – ‘hindgut ruminant’ – less efficient, high throughput

Cattle – ‘foregut ruminant’ – more efficient, can eat less.

Page 23: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A. Productivity

3. Net Secondary Productivity

- affected by nutrient ratios, growth rates, and most limiting variable. May need to eat a lot to get enough of the limiting variable.

N:P :: 50:1

N:P :: 15:1

Fast growing; need higher ratio of Phosphorus for DNA synthesis.

Page 24: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A. Productivity

3. Net Secondary Productivity

- Net Production Efficiency = P/A

Page 25: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

What might this depend on???

NPP

NSP

Page 26: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A. Productivity

3. Net Secondary Productivity

- net production efficiency = P/A

0.7%Shrews

0.5%Birds

6-10%MostMammals

Up to 75% for sedentary poikilotherms

Page 27: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A.Productivity

B.Trophic Pyramids

Page 28: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A.Productivity

B.Trophic Pyramids

- ecological efficiency: NSP/NPP (5-20%)

NPP of Producers (PLANTS)

NPP of HERBIVORES

Loss due to 2nd Law

NPP of Primary Carnivores

NPP of Secondary Carnivores

Page 29: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

a. trophic "pyramids"

NPP of Producers (PLANTS)

NPP of HERBIVORES

Loss due to 2nd Law

NPP of Primary Carnivores

NPP of Secondary Carnivores

This is why large carnivores are RARE, and why they have large RANGES

Page 30: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A.Productivity

B.Trophic Pyramids

C.Detrital Foodchains

Herbivores

Predators

Page 31: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A.Productivity

B.Trophic Pyramids

C.Detrital Foodchains

HerbivoresNPPDetritivores

Temperate forest: 1.5% - 2.5%Old-field Habitat: 12%Plankton: 60-99%

Page 32: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A.Productivity

B.Trophic Pyramids

C.Detrital Foodchains

D.‘Biomass Accumulation Ratios’

If we know the mean ‘standing crop’ of biomass from year to year, and we know the net productivity, we can calculate how long, on average the biomass persists:

BAR (per year) = (biomass/m2) / (np of biomass / m2 / yr)

Page 33: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A.Productivity

B.Trophic Pyramids

C.Detrital Foodchains

D.‘Biomass Accumulation Ratios’

If we know the mean ‘standing crop’ of biomass from year to year, and we know the net productivity, we can calculate how long, on average the biomass persists:

BAR (per year) = (biomass/m2) / (np of biomass / m2 / yr)

Forests: ~ 20 years Tropical leaf litter: 3 monthsPhytoplantkon: ~20 days Temperate leaf litter: 2-20 years

Page 34: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

Ecosystem Ecology

I. Introduction

II. Energy Flow

A.Productivity

B.Trophic Pyramids

C.Detrital Foodchains

D.BAR

E.Human Concerns

Page 35: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

E. Human Concerns

Page 36: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

E. Human Concerns

Page 37: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

E. Human Concerns

Page 38: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

E. Human Concerns

500% increase in 50 years, with population increase of 250%

Page 39: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

E. Human Concerns

A doubling of meat production per capita

Page 40: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

E. Human Concerns

25% of catch by weight discarded

Page 41: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

E. Human Concerns

Page 42: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

E. Human Concerns

6-10 lbs of feed for 1 lb increase in cattle weight2-5 lbs of fish meal for 1 lb increase in farmed fish weight

Page 43: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

E. Human Concerns Edible kilocalories produced from kilocalories of energy required for cultivation are:

18.1% for chicken,

6.7% for grass-fed beef,

5.7% for farmed salmon

0.9% for shrimp.

123% for potatoes

250% for corn

415% for soy

input calories converted to calories able to be utilized by humans

So, for every 100 calories of energy we put in to raise chickens, we get 18 calories of energy produced in chicken meat. 100 cal into soy, 415 calories out.

Page 44: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

E. Human Concerns

Food production, per capita(400 kg per year is healthy minimum)

Page 45: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

SO HOW DID WE DO IT?

Page 46: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

E. Human Concerns

EXTENSIFICATION – MORE AREA

Page 47: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

E. Human Concerns

EXTENSIFICATION – MORE AREA

Page 48: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

E. Human Concerns

The best land has already been used; further expansion in marginal areas is costly and requires more supplementation

Page 49: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

E. Human Concerns

47% of historical forested land has been cut

Page 50: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

E. Human Concerns

INTENSIFICATION

Page 51: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

E. Human Concerns

Page 52: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

E. Human Concerns

The best land has already been used; further expansion in marginal areas is costly and requires more supplementation

Page 53: Ecosystem Ecology. I. Introduction - Ecosystem: an assemblage of organisms, together with their chemical and physical environments

E. Human Concerns

Global NPP (dry mass) = 224 billion tons. 59% is terrestrial, and of this, humans 35-40% is controlled by humans, either eaten directly or fed to animals we will consume