BRAIN RESEARCH INTERNATIONAL MULTIDISCIPLINARY JOURNALDEVOTED TO FUNDAMENTAL RESEARCH IN THE BRAIN SCIENCES

EDITOR-IN-CHIEF: Irwin B. Levitan (Philadelphia, PA, USA)

DEPUTY EDITORS-IN-CHIEF: Leslie C. Griffi th (Waltham, MA, USA) Gary S. Aston-Jones (Charleston, SC, USA)

SENIOR EDITORS

Gary S. Aston-Jones (Charleston, SC, USA)Francesco P. Battaglia (Nijmegen, The Netherlands)Fen-Biao Gao (Worcester, MA, USA)Leslie C. Griffi th (Waltham, MA, USA)Mary Heinricher (Portland, OR, USA)Lorraine Iacovitti (Philadelphia, PA, USA)Leonard K. Kaczmarek (New Haven, CT, USA)Jae Young Koh (Seoul, Korea)Diane Lipscombe (Providence, RI, USA)Brian A. MacVicar (Vancouver, Canada)David J. Perkel (Seattle, WA, USA)Peter H. Reinhart (Boston, MA, USA)Erich Schröger (Leipzig, Germany)Michael E. Selzer (Philadelphia, PA, USA)J. Paul Taylor (Memphis, TN, USA)Flora M. Vaccarino (New Haven, CT, USA)Barry Waterhouse (Philadelphia, PA, USA)Christina L. Williams (Durham, NC, USA)

VOLUME 1509 ( 2013 )

AMSTERDAM – BOSTON – LONDON – NEW YORK – OXFORD – PARIS – PHILADELPHIA –SAN DIEGO – ST LOUIS