fig. 1 most abundant apiformes species calculated as a proportion
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Fig. 1 Most abundant Apiformes species calculated as a proportion of the total abundance of Apiformes in the collection period. Pinon Canyon Maneuver Site, 2008.
70.096
60.096
50.096
40.096
30.096
20.096
10.096
0.0%
Fig. 2 Least abundant Apiformes species calculated as a proportion of the total abundance of Apiformes in the collection period. Pinon Canyon Maneuver Site, 2008.7
04% 1
j 0.391>
0.2%
0.1%
0.0%
Fig. 3 Most abundant Carabidae species calculated as a proportion of the total abundance of Carabidae in the collection period. Pinon Canyon Maneuver Site, 2008.
Fig. 4 Least abundant Carabidae species calculated as a proportion of the total abundance of Carabidae in the collection period. Pinon Canyon Maneuver Site, 2008.
Fig. 5 Asilidae species abundance calculated as a proportion of the total abundace of Asilidae in the collection period. Pinon Canyon Maneuver Site, 2008.
Fig. 6 Butterfly species abundance calculated as a proportion of the total abundance of butterflies in the collection period. Pinon Canyon Maneuver Site, 2008.
0.0%
5.0%
10.0%
15.0%
20.0%
25.0%
30.0%
Fig. 7 Most abundant Orthoptera species calculated as a proportion of the total abundance of Orthoptera in the collection period. Pinon Canyon Maneuver Site, 2008.
Fig. 8 Moderately abundant Orthoptera species calculated as a proportion of the total abundance of Orthoptera in the collection period. Pinon Canyon Maneuver Site, 2008.
Fig. 9 Least abundant Orthoptera species calculated as a proportion of the total abundance of Orthoptera in the collection period. Pinon Canyon Maneuver Site, 2008.
Fig. 10 Apiformes dominant species comparison between 2007 and 2008. Each species as a percentage of the entire taxa. Pinon Canyon Maneuver Site, 2008.
Fig. 11 Carabidae dominant species comparison between 2007 and 2008. Each species as a percentage of the entire taxa. Pinon Canyon Maneuver Site, 2008.
Fig. 12 Asilidae dominant species comparison between 2007 and 2008. Each species as a percentage of the entire taxa. Pinon Canyon Maneuver Site, 2008.
Fig. 13 Butterfly dominant species comparison between 2007, 2008 and 2009. Each species as a percentage of the entire taxa. Pinon Canyon Maneuver Site, 2009
Pyrgus communis
Euptoieta claudia
Colias philodice
Brephidium exile
Speyeria coronis
Cercyonis pegala
Pontia protodice
Nathalis iole
Phyciodes picta
Hesperia uncas
Hesperia pahaska
Callophrys gryneus
Other
2009 Butterflies
Fig. 14 Orthoptera dominant species comparison between 2007 and 2008. Each species as a percentage of the entire taxa. Pinon Canyon Maneuver Site, 2008.
Fig. 15 Acrididae dominant species comparison between 2007 and 2008. Each species as a percentage of the entire taxa. Pinon Canyon Maneuver Site, 2008.
The only species of concern observed in 2007, Euphilotes rita (Mattoni, [1966]) was not observed in 2008. In an effort to document E. rita in
2008, we searched for host plants, larvae and adults of this species while carrying out other collection work, in additon to the butterfly survey
technique that was used to document the species in 2007. We collected two individuals of a moth, Alypia octomaculata (Fabricius), in the
Cottonwood habitat. The occurrence of the moth at Pinon Canyon will be a range extension if a persistent population can be demonstrated. We
will look for A. octomaculata larva and its plant hosts in 2009 to verify a persistant population. We collected 86 new species this year. The biology
and behavor of the individual species will determine a species presence in a habitat based on yearly environmental variations. An updated species
list can be found in Appendix 1. Species added to the list in 2008 are in bold.
Habitat Results
Fig. 16 Sand sagebrush habitat in June of 2007 on the left and June of 2008 on the right to demonstrate differences in habitat quality. Pinon Canyon Maneuver Site, 2008.
Habitat quality was relatively poor in 2008 when compared with 2007, based on visual assessment of the habitat by the presence of green,
flowering plants or moisture and the overall lack of rain in 2008 (Fig. 16). The habitat resources were scarce in most plant communities we studied
throughout the collection period with the exception of cottonwood habitat that is associated with a continuous water source. There was a general
trend for species to be found in a more restricted set of habitats in 2008 than in 2007 and more individuals were collected near permanent water
sources. Several species, like Amara quenseli, Nathalis iole, and Melanoplus discolor that were present in multiple habitats in 2007 were only
observed in a single or few habitats in 2008 (Appendix 2). In contrast, some species, like Anthophora sp., Efferia sp., Psilocurus camposi, and Amara
carinata that were found in few habitats in 2007 were found in additional habitats in 2008 (Appendix 2). The presence of a species like Arphia
conspersa, Melanoplus flavidus, or Philonicus limpidipennis in different habitats from 2007 demonstraites either an increase the amount of time
actively searching for food and mates or that species’ ability to outcompete other species that are less drought tolerant.
Table 2: Total specimens, number of species, and diversity indices for collections from nine sites at Pinon Canyon Maneuver Site, 2008.
While diversity was relativley high in 2008, evenness was very low. High diversity with a low evenness indicated that there was still a high
number of species in the habitat and that they were relativley abundant, but the likelyhood of encountering any particular species at a single
Habitat Shannon's Index Shannon's Evenness Modified Simpson's Index Total Individuals
collected Species
JUMO/BOER 3.32 0.05
0.93 372
64
BOGR/HIJA 2.34 0.04 0.80 980 57 ARFI/YUGL2 2.68 0.04 0.86 766 60 ATCA/SPAI 2.78 0.05 0.88 490 56 ARFI/YUGL1 2.80 0.05 0.88 588 60 PODE/BRJA 2.50 0.03 0.81 1,262 80 OPIM/BOGR 2.03 0.04 0.78 746 46 GLME/FRJA 1.97 0.05 0.68 453 40 TA spp. 1.46 0.10 0.69 513 14
collection event or time period was low. The low evenness indicates the insect population was unpredicable and unstable. The juniper habitat had
the highest diversity and had a more stable community of insects than any other habitat. Most of the habitats were comparable in diversity and
evenness, except the Tamarix sp. site. It had a low diversity and a very low evenness. This indicates that most species are not consistantly using this
habitat and are most likely vagrant individuals who are just passing through. The highest number of individuals were collected at the cottonwood
habitat, where water was available throughout the year. Species present in the habitats were much lower when compared to 2007.
Sampling Results
Pitfall sampling was the most productive collection technique for 2008, yielding the most individuals and species. Although the diversity
was slightly less than the other techniques, Shannon’s diversity (2.96-2.87) and evenness of Shannon’s diverstiy (0.078-0.026) were very similar for
all collection techniques (Table 3). Evenness for all categories was less than 0.10 (low), with samples being very variable (Table 3). Surveys were still
important techniques although the Asilid and Apiformes surveys were relatively unsucessful (Table 3). The low collection success was likely caused
by the climatic conditions and little persistance of suitable food for these groups in any one location. The current collection techniques appear to
be satisfactory and we will conduct the 2009 surveys in the same manner.
Table 3: Total specimens, number of species, and diversity indices for three collection methods at Pinon Canyon Maneuver Site, 2008.
Collection method
Total individuals collected
Species Shannon's
Index Shannon's Evenness
Modified Simpson's Index
Pitfalls Trap 5,515 112 2.87 0.026 0.89
Surveys 348 63 2.96 0.047 0.88
Conclusions
The biodiversity project at Pinon Canyon Manuever Site is continuing to add species to the species list as well as gaining valuable insight into
habitat presence of a species as a preliminary indication of habitat use. Diversity remained similar between years independent of climate
differences, but the ability to document all species in an area was reduced as indicated by the low evenness of Shannon’s diversity and the low
abundance of species that were collected in 2008. Habitat presence of species will shift with changing climatic events. The difference in climate
from year to year strongly influences the collection and analysis of results. Climatic differences confound habitat usage. Without multiple years of
data collected in similar climatic years, general habitat use is difficult to determine. The third summer of collection will allow for better
comparisons to be made with more accurate conclusions about a species use of a habitat.
Malaise Trap 323 39 2.93 0.075 0.91
Appendix 1: Species list with 2008 additions in bold. Pinon Canyon Maneuver Site 2008. Orthoptera (grasshoppers, crickets, katydids)
Family Scientific Name Common Name
Acrididae Acrolophitus hirtipes Green Fool Grasshopper
Acrididae Aeoloplides turnbulli Russian Thistle Grasshopper
Acrididae Ageneotettix deorum White-whiskered Grasshopper
Acrididae Amphitornus coloradus Striped Slant-faced Grasshopper
Acrididae Arphia pseudonietana Northwestern Red-Winged Grasshopper
Acrididae Arphia conspersa Speckle-winged Rangeland Grasshopper
Acrididae Arphia simplex Plains Yellow-winged Grasshopper
Acrididae Aulocara elliotti Big Headed Grasshopper
Acrididae Aulocara femoratum White Cross Grasshopper
Acrididae Boopedon gracile Graceful Range Grasshopper
Acrididae Boopedon nubilum Ebony Grasshopper
Acrididae Camnula pellucida Clear-winged Grasshopper
Acrididae Chloealtis abdominalis Thomas's Broad-Winged Grasshopper
Acrididae Chloealtis conspersa Sprinkled Broad-Winged Grasshopper
Acrididae Chorthippus curtipennis Marsh Meadow Grasshopper
Acrididae Chortophaga viridifasciata Northern Green-striped Grasshopper
Acrididae Circotettix rabula Wrangler Grasshopper
Acrididae Cordillacris crenulata Crenulated Grasshopper
Acrididae Cordillacris occipitalis Spotted Winged Grasshopper
Acrididae Dactylotum bicolor Pictured Grasshopper
Acrididae Derotmema haydeni Hayden’s Grasshopper
Acrididae Eritettix simplex Velvet Striped Grasshopper
Acrididae Hadrotettix magnificus Magnificent Grasshopper
Acrididae Hesperotettix speciosus Western Grass-Green Grasshopper
Acrididae Hadrotettix trifasciatus Three Banded Range Grasshopper
Acrididae Hesperotettix viridis Snakeweed Grasshopper
Acrididae Hippiscus ocelote Wrinkled Grasshopper
Acrididae Hippopedon capito Apache Grasshopper
Acrididae Hypochlora alba Cudweed Grasshopper
Acrididae Leprus cyaneus Blue-Winged Grasshopper
Acrididae Leprus robustus
Acrididae Melanoplus spp Spurthroated Grasshoppers
Acrididae Melanoplus angustipennis Narrow-winged Sand Grasshopper
Acrididae Melanoplus arizonae Arizona Spur-throated Grasshopper
Acrididae Melanoplus bivittatus Two-Striped Grasshopper
Acrididae Melanoplus borealis Northern Grasshopper
Acrididae Melanoplus bowditchi Sagebrush Grasshopper
Acrididae Melanoplus confusus Little Pasture Spurthroated Grasshopper
Acrididae Melanoplus differentialis Differential Grasshopper
Acrididae Melanoplus discolor Contrasting Spurthroated Grasshopper
Acrididae Melanoplus dodgei Dodge's Spurthroated Grasshopper
Acrididae Melanoplus fasciatus Huckleberry Spur-Throat Grasshopper
Orthoptera (grasshoppers, crickets, katydids)
Acrididae Melanoplus flavidus Yellowish Spurthroated Grasshopper
Acrididae Melanoplus femurrubrum Red-legged Grasshopper
Acrididae Melanoplus foedus Striped Sand Grasshopper
Acrididae Melanoplus gladstoni Gladston’s Spurthroated Grasshopper
Acrididae Melanoplus glaucipes Glaucous-Legged Spurthroated Grasshopper
Acrididae Melanoplus lakinus Lakin Grasshopper
Acrididae Melanoplus occidentalis Flabellate Grasshopper
Acrididae Melanoplus packardii Packard Grasshopper
Acrididae Melanoplus ponderosus Ponderous Spurthroated Grasshopper
Acrididae Melanoplus regalis Regal Spurthroated Grasshopper
Acrididae Melanoplus sanguinipes Migratory Grasshopper
Acrididae Melanoplus splendidus Splendid Grasshopper
Acrididae Melanoplus tristis
Acrididae Melanoplus yarrowii Yarrow's Spurthroated Grasshopper
Acrididae Mermiria bivittata Two-striped Mermiria Grasshopper
Acrididae Mermiria picta Painted Slantfaced Grasshopper
Acrididae Mestobregma plattei Platt Range Grasshopper
Acrididae Metator pardalinus Pard Grasshopper
Acrididae Opeia obscura Obscure Grasshopper
Acrididae Orphulella speciosa Pasture Grasshopper
Acrididae Pardalophora apiculata Coral-winged Grasshopper
Acrididae Paropomala wyomingensis Wyoming Toothpick Grasshopper
Acrididae Phlibostroma quadrimaculatum Four-spotted Grasshopper
Acrididae Phoetaliotes nebrascensis Large-headed Grasshopper
Acrididae Psoloessa delicatula Brown-Spotted Range Grasshopper
Acrididae Psoloessa texana Texas Spotted Range Grasshopper
Acrididae Pseudopomala brachyptera Big Slender Grasshopper
Acrididae Schistocerca obscura Obscure Bird Grasshopper
Acrididae Spharagemon collare Mottled Sand Grasshopper
Acrididae Spharagemon equale Orange-legged Grasshopper
Acrididae Stethophyma gracile Graceful Sedge Grasshopper
Acrididae Syrbula admirabilis Admirable Grasshopper
Acrididae Syrbula montezuma Slantfaced Grasshopper
Acrididae Trachyrhachys coronata Crowned Grasshopper
Acrididae Trachyrhachys kiowa Kiowa range Grasshopper
Acrididae Trimerotropis sp Slender Grasshopper
Acrididae Trimerotropis gracilis Thomas' Slender Grasshopper
Acrididae Trimerotropis latifasciata Broad Banded Grasshopper
Acrididae Trimerotropis melanoptera Black-winged Grasshopper
Acrididae Trimerotropis pallidipennis Pallid-winged Grasshopper
Acrididae Tropidolophus formosus Great Crested Grasshopper
Acrididae Xanthippus corallipes Red Shanked Grasshopper
Gryllidae Acheta domesticus House Cricket
Gryllidae Allonemobius fasciatus Striped Ground Cricket
Gryllidae Cycloptilum sp
Gryllidae Eunemobius sp Carolina Ground Cricket
Gryllidae Gryllus pennsylvanicus Fall Field Cricket (Black field cricket)
Oedipodinae Encoptolophus costalis Dusky Grasshopper
Rhaphidophoridae Ceuthophilus spp Camel cricket
Orthoptera (grasshoppers, crickets, katydids)
Romaleidae Brachystola magna Plains Lubber Grasshopper
Tetrigidae Tetrix ornata Ornate Pygmy Grasshopper
Tettigoniidae Microcentrum rhomhbifolium Angle-Winged Katydid
Tettigoniidae Pseudophyllinae sp Katydid
Tettigoniidae Scudderia furcata Fork-tailed Bush Katydid
Odonata (dragonflies and damselflies)
Family Scientific Name Common Name
Calopterygidae
Broad-Winged Damselflies
Coenagrionidae
Pond Damselflies
Cordulidae
Emeralds
Gomphidae
Clubtail Dragonflies
Libellulidae Libellula pulchella Twelve Spotted Skimmer
Libellulidae Tramea lacerata Black Saddlebags
Neuroptera
Family Scientific Name Common Name
Chrysopidae
Lacewing
Mantispidae
Mantis Fly
Myrmeleontidae
Ant lion
Coleoptera (beetles)
Family Scientific Name Common Name
Anthicidae Notoxus sp Ant-like Flower Beetle
Buprestidae
Metallic Wood Boring Beetles
Cantharidae Cantharis sp Soldier Beetles
Carabidae Agonum cyclifer Ground Beetle
Carabidae Agonum decorum Ground Beetle
Carabidae Agonum extensicolle Ground Beetle
Carabidae Agonum placidum Ground Beetle
Carabidae Agonum punctiforme Ground Beetle
Carabidae Amara sp Ground Beetle
Carabidae Amara carinata Ground Beetle
Carabidae Amara deparca Ground Beetle
Carabidae Amara quenseli Ground Beetle
Carabidae Amara thoracica Ground Beetle
Carabidae Amblycheila cylindriformis Great Plains Giant Tiger Beetle
Carabidae Badister obtusus Ground Beetle
Carabidae Brachinus sp Ground Beetle
Carabidae Brachinus phaeocerus Ground Beetle
Carabidae Calathus sp Ground Beetle
Carabidae Calathus opaculus Ground Beetle
Carabidae Calosoma sp Ground Beetle
Carabidae Calosoma affine Ground Beetle
Carabidae Calosoma marginake Ground Beetle
Coleoptera (beetles)
Carabidae Calosoma peregrinator Ground Beetle
Carabidae Calosoma scrutator Ground Beetle
Carabidae Chlaenius tricolor Vivid metallic ground beetles
Carabidae Chlaenius s sericeus Ground Beetle
Carabidae Chlaenius t tomentosus Ground Beetle
Carabidae Cicindela sp Ground Beetle
Carabidae Cicindela belfragei Loamy-ground Tiger Beetle
Carabidae Cicindela formosa Big Sand Tiger Beetle
Carabidae Cicindela n nigrocoerulea Ground Beetle
Carabidae Cicindela o obsoleta Ground Beetle
Carabidae Cicindela p punctulata Ground Beetle
Carabidae Cicindela s scutellaris Festive Tiger Beetle
Carabidae Cratacanthus dubius Ground Beetle
Carabidae Cyclotrachelus constrictus Ground Beetle
Carabidae Cyclotrachelus substriatus Ground Beetle
Carabidae Cymindis interior Ground Beetle
Carabidae Cymindis planipennis Ground Beetle
Carabidae Cymindis punctiger Ground Beetle
Carabidae Dicaelus l laevipennis Ground Beetle
Carabidae Diplocheila obtusa Ground Beetle
Carabidae Euryderus sp Ground Beetle
Carabidae Euryderus grossus Ground Beetle
Carabidae Galerita sp Ground Beetle
Carabidae Galerita janus Ground Beetle
Carabidae Harpalus sp Ground Beetle
Carabidae Harpalus caliginosus Ground Beetle
Carabidae Harpalus fraternus Ground Beetle
Carabidae Harpalus paratus Ground Beetle
Carabidae Harpalus pensylvanicus Ground Beetle
Carabidae Hartonymus alternatus Ground Beetle
Carabidae Lebia viridis Ground Beetle
Carabidae Micrixys distincta Ground Beetle
Carabidae Pasimachus californicus Ground Beetle
Carabidae Pasimachus elongatus Ground Beetle
Carabidae Piosoma setosum Ground Beetle
Carabidae Poecilus sp Ground Beetle
Carabidae Poecilus chalcites Ground Beetle
Carabidae Poecilus cyanicolor Ground Beetle
Carabidae Poecilus l lucublandus Ground Beetle
Carabidae Poecilus scitulus Ground Beetle
Carabidae Pterostichus commutabilis Ground Beetle
Carabidae Rhadine dissecta sp Ground Beetle
Carabidae Scarites subterraneus Big Headed Ground Beetle
Carabidae Scarites quadriceps Ground Beetle
Cerambycidae Oberea sp Long-horned Wood-boring Beetles
Cerambycidae Prionus sp Long-horned Wood-boring Beetles
Chrysomelidae Calligrapha sp Leaf Beetles
Chrysomelidae Diabrotica sp Leaf Beetles
Coleoptera (beetles)
Cleridae
Checkered Flower Beetles
Coccinellidae Anatis lecontei Large Ladybeetle
Coccinellidae Coccinella septempunctata Seven-spotted Ladybeetle
Coccinellidae Hippodamia parenthesis
Coccinellidae Hippodamia convergens Convergent Ladybeetle
Coccinellidae Myzia interrupta
Curculiondiae
Weevils
Dytiscidae
Predacious Diving Beetles
Elateridae
Click Beetles
Histeridae
Clown Beetles
Lampyridae
Fireflies
Meloidae Epicauta sp Blister Beetles
Nitidulidae Capophilus sp Sap Beetles
Scarabidae Aphodices sp
Scarabidae Canthon sp Dung Beetle
Scarabidae Eurphoria inda
Scarabidae Melanocanthon sp Dung Beetle
Sciritidae
Marsh Beetles
Silphidae Nicrophorus sp Burying Beetles
Silphidae Silpha sp Carrion Beetles
Staphylinidae Aleochara sp Rove Beetles
Tenebrionidae Diaperis sp Darkling Beetles
Tenebrionidae Eleodes sp Darkling Beetles
Trogidae
Hide Beetles
Lepidoptera (butterflies and moths)
Family Scientific Name Common Name
Arctiidae Grammia sp Tiger Moths
Arctiidae Hyproprea sp
Geometridae
Geometrid Moths
Hesperiidae Amblyscirtes vialis Common Roadside Skipper
Hesperiidae Hesperia pahaska Pahaska Skipper
Hesperiidae Hesperia uncas Uncas Skipper
Hesperiidae Hesperopsis alpheus Saltbush Sootywing
Hesperiidae Megathymus streckeri Strecker's Giant Skipper
Hesperiidae Pholisora catullus Common Sootywing
Hesperiidae Pyrgus communis Common Checkered Skipper
Hesperiidae Pyrgus scriptura Small Checkered Skipper
Hesperiidae Thorybes pylades Northern Cloudywing
Lycaenidae Brephidium exile Western Pygmy Blue
Lycaenidae Callophrys gryneas Juniper Hairstreak
Lycaenidae Euphilotes rita Rita Blue
Lycaenidae Hemiargus isola Reakirt’s Blue
Lycaenidae Leptotes marina Marine Blue
Lycaenidae Plebejus acmon Acmon Blue
Lycaenidae Plebejus saepiolus Greenish Blue
Lepidoptera (butterflies and moths)
Lycaenidae Strymon melinus Grey Hairstreak
Noctuidae
Millers or Owlet Moths
Noctuidae Alypia octomaculata Eight-spotted Forester
Nymphalidae Anaea andria Goatweed Leafwing
Nymphalidae Cercyonis oetus Small Wood Nymph
Nymphalidae Cercyonis pegala Common Wood Nymph
Nymphalidae Danaus plexippus Monarch
Nymphalidae Euptoieta claudia Variegated Fritillary
Nymphalidae Junonia coenia Common Buckeye
Nymphalidae Limenitis archippus Viceroy
Nymphalidae Nymphalis antiopa Mourning Cloak
Nymphalidae Phyciodes campestris Field Crescent
Nymphalidae Phyciodes mylitta Mylitta Crescent
Nymphalidae Phyciodes pallida Pale Crescent
Nymphalidae Phyciodes picta Painted Crescent
Nymphalidae Phyciodes tharos Pearl Crescent
Nymphalidae Polygonia interrogationis Question Mark
Nymphalidae Speyeria coronis Coronis Fritillary
Nymphalidae Speyeria edwardsii Edward's Fritillary
Nymphalidae Thessalia fulvia Fulvia Checkerspot
Nymphalidae Vanessa atalanta Red Admiral
Nymphalidae Vanessa cardui Painted Lady
Nymphalidae Vanessa virginiensis American Lady
Papilionidae Papilio mulitcaudata Two-tailed Swallowtail
Papilionidae Papilio polyxenes Black Swallowtail
Papilionidae Papilio rutulus Western Tiger Swallowtail
Pieridae Colias cesonia Southern Dogface
Pieridae Colias eurytheme Orange Sulphur
Pieridae Colias philodice Clouded Sulphur
Pieridae Eurema nicippe Sleepy Orange
Pieridae Nathalis iole Dainty Sulphur
Pieridae Pieris rapae Cabbage White
Pieridae Pontia occidentalis Western White
Pieridae Pontia protodice Checkered White
Pyralidae
Pyralid Moth
Sphingidae Hyles lineata White Lined Sphinx Moth
Sphingidae Proserpinus juanita Juanita Sphinx Moth
Diptera (flies)
Family Scientific Name Common Name
Asilidae Albibarbefferia sp Robber Flies
Asilidae Albibarbefferia albibarbis Robber Flies
Asilidae Albibarbefferia bicolor Robber Flies
Asilidae Albibarbefferia leucocoma Robber Flies
Asilidae Aridefferia prattii Robber Flies
Asilidae Aridefferia subarida Robber Flies
Asilidae Asilus auriannulatus Robber Flies
Diptera (flies)
Asilidae Asilus sp Robber Flies
Asilidae Atomosia sp Robber Flies
Asilidae Atomosia melanopogon Robber Flies
Asilidae Atomosia nuda Robber Flies
Asilidae Atomosia puella Robber Flies
Asilidae Carinefferia costalis Robber Flies
Asilidae Diogmites sp Robber Flies
Asilidae Diogmites neoternatus Robber Flies
Asilidae "Efferia" spp Robber Flies
Asilidae Efferia anomala Robber Flies
Asilidae Efferia apache Robber Flies
Asilidae Efferia bryanti Robber Flies
Asilidae Efferia duncani Robber Flies
Asilidae Laphystia varipes Robber Flies
Asilidae Lasiopogon sp Robber Flies
Asilidae Lasiopogon quadrivittatus Robber Flies
Asilidae Leptogaster sp Robber Flies
Asilidae Leptogaster brevicornis Robber Flies
Asilidae Leptogaster weslacensis Robber Flies
Asilidae Nerax beameri Robber Flies
Asilidae Ospriocerus sp Robber Flies
Asilidae Ospriocerus abdominalis Robber Flies
Asilidae Philonicus sp Robber Flies
Asilidae Philonicus arizonensis Robber Flies
Asilidae Philonicus limpidipennis Robber Flies
Asilidae Pogoniefferia sp Robber Flies
Asilidae Pogoniefferia bicaudata Robber Flies
Asilidae Pogoniefferia helenae Robber Flies
Asilidae Pogoniefferia texana Robber Flies
Asilidae Pogonosoma sp Robber Flies
Asilidae Proctacanthella exquisita Robber Flies
Asilidae Proctacanthus sp Robber Flies
Asilidae Promachus sp Robber Flies
Asilidae Psilocurus sp Robber Flies
Asilidae Psilocurus arizonensis Robber Flies
Asilidae Psilocurus birdi Robber Flies
Asilidae Psilocurus camposi Robber Flies
Asilidae Stenopogon sp Robber Flies
Asilidae Stichopogon sp Robber Flies
Culicidae
Mosquitoes
Muscidae Musca domestica House Fly
Mydidae Mydas sp Mydas Fly
Scorpiones
Family Scientific Name Common Name
Buthidae Centruroides vittatus Scorpion
Hymenoptera (bees)
Family Scientific Name Common Name
Andrenidae Andrena confederata
Andrenidae Andrena helianthi
Andrenidae Andrena merriami
Andrenidae Perdita spp
Andrenidae Protandrena spp
Apidae Anthophora spp
Apidae Bombus spp Bumblebee
Apidae Diadasia diminuta
Apidae Diadasia enavata
Apidae Diadasia olivacea
Apidae Diadasia rinconis rinconis
Apidae Eucera spp
Apidae Holcopasites tegularis
Apidae Melecta sp
Apidae Melissodes intorta
Apidae Melissodes spp
Apidae Neolarra verbesinae
Apidae Nomada sp
Apidae Svastra obliqua
Apidae Tetralphiella sp
Apidae Triepeolus sp
Apidae Xenoglossa sp
Colletidae Colletes spp Digger Bee
Halictidae Agapostemon angelicus
Halictidae Agapostemon angelicus or texanus
Halictidae Agapostemon sp
Halictidae Augochlora sp
Halictidae Augochloropsis sp
Halictidae Dieunomia sp
Halictidae Halictus ligatus
Halictidae Halictus virgatellus or provancheri
Halictidae Lasioglossum spp
Halictidae Nomia sp Alkali Bee
Halictidae Sphecodes sp
Megachilidae Athidium sp
Megachilidae Dianthidium spp
Megachilidae Megachile amica
Megachilidae Megachile brevis
Megachilidae Megachile brevis onobrychidis
Megachilidae Osmia spp
Braconidae
Formicidae
Ants
Ichneumonidae
Ichneumonid Wasps
Mutilidae Dasymatina sp Velvet Ants
Pompilidae Pepsis sp Spider Wasps
Sphecidae
Thread-wasted Wasp
Vespidae Polistes sp Paper Wasp
Vespidae Vespula sp
Appendix 2: Presence or absence of a species for each year. Black indicates the 2007 occurrence and 2008 is indicated with grey. Pinon Canyon Maneuver Site 2008.
Carabidae
Species ARFI/YUGL
ATCA/SPAI
BOGR/HIJA
GLME/FRJA
JUMO/BOER
OPIM/BOGR
PIPO/SONU
PODE/BRJA TA sp.
Agonum cyclifer
Agonum decorum
Agonum extensicolle
Agonum placidum
Amara carinata
Amara deparca
Amara quenseli
Amara thoracica
Amblycheila cylindriformis
Badister obtusus
Brachinus phaeocerus
Carabidae
Species ARFI/YUGL
ATCA/SPAI
BOGR/HIJA
GLME/FRJA
JUMO/BOER
OPIM/BOGR
PIPO/SONU
PODE/BRJA TA sp.
Calathus opaculus
Calosoma affine
Calosoma scrutator
Chlaenius s. sericeus
Chlaenius t. tomentosus
Chlaenius tricolor
Cicindela n. nigrocoerulea
Cicindela o. obsoleta
Cicindela obsoleta
Cicindela p. punctulata
Cicindela s. scutellaris
Cicindela scutellaris
Cratacanthus dubius
Cyclotrachelus constrictus
Cyclotrachelus substriatus
Cymindis interior
Cymindis planipennis
Cymindis punctiger
Dicaelus l. laevipennis
Diplocheila obtusa
Euryderus grossus
Galerita janus
Harpalus caliginosus
Harpalus fraternus
Harpalus paratus
Harpalus pensylvanicus
Lebia viridis
Micrixys distincta
Pasimachus californicus
Pasimachus elongatus
Piosoma setosum
Poecilus chalcites
Carabidae
Species ARFI/YUGL
ATCA/SPAI
BOGR/HIJA
GLME/FRJA
JUMO/BOER
OPIM/BOGR
PIPO/SONU
PODE/BRJA TA sp.
Poecilus cyanicolor
Poecilus l. lucublandus
Poecilus lucublandus
Poecilus scitulus
Pterostichus commutabilis
Rhadine dissecta sp.
Scarites subterraneus
Lepidoptera
Species ARFI/YUGL
ATCA/SPAI
BOGR/HIJA
GLME/FRJA
JUMO/BOER
OPIM/BOGR
PIPO/SONU
PODE/BRJA TA sp.
Amblyscirtes vialis
Anaea andria
Brephidium exile
Callophrys gryneas
Cercyonis oetus
Cercyonis pegala
Colias cesonia
Colias eurytheme
Colias philodice
Danaus plexippus
Euphilotes rita
Euptoieta claudia
Eurema nicippe
Hemiargus isola
Hesperia pahaska
Hesperia uncas
Hesperopsis alpheus
Leptotes marina
Limenitis archippus
Lycaeides melissa
Lepidoptera
Species ARFI/YUGL
ATCA/SPAI
BOGR/HIJA
GLME/FRJA
JUMO/BOER
OPIM/BOGR
PIPO/SONU
PODE/BRJA TA sp.
Megathymus streckeri
Nathalis iole
Nymphalis antiopa
Papilio multicaudata
Papilio polyxenes
Papilio rutulus
Pholisora catullus
Phyciodes campestris
Phyciodes mylitta
Phyciodes pallida
Phyciodes picta
Phyciodes tharos
Pieris rapae
Plebejus acmon
Plebejus saepiolus
Polygonia interrogationis
Pontia occidentalis
Pontia protodice
Pyrgus communis
Pyrgus scriptura
Speyeria coronis
Speyeria edwardsii
Strymon melinus
Thessalia fulvia
Thorybes pylades
Vanessa cardui
Vanessa virginiensis
Orthoptera
Scientific Names ARFI/YUGL
ATCA/SPAI
BOGR/HIJA
GLME/FRJA
JUMO/BOER
OPIM/BOGR
PIPO/SONU
PODE/BRJA TA sp.
Orthoptera
Scientific Names ARFI/YUGL
ATCA/SPAI
BOGR/HIJA
GLME/FRJA
JUMO/BOER
OPIM/BOGR
PIPO/SONU
PODE/BRJA TA sp.
Acheta domesticus
Acrolophitus hirtipes
Aeoloplides turnbulli
Ageneotettix deorum
Amphitornus coloradus
Arphia conspersa
Arphia pseudonietana
Arphia simplex
Aulocara elliotti
Aulocara femoratum
Boopedon gracile
Boopedon nubilum
Brachystola magna
Cactylotum bicolor
Camnula pellucida
Ceuthophilus sp.
Chloeatis abdominalis
Choealtis conspersa
Chorthippus curtipennis
Chortophaga viridifasciata
Circotettix rabula
Cordillacris crenulata
Cordillacris occipitalis
Cycloptilum sp.
Dactylotum bicolor
Derotmema haydeni
Encoptolophus costalis
Eritettix simplex
Eunemobius sp.
Gryllus pennsylvanicus
Hadrotettix magnificus
Hadrotettix trifasciatus
Orthoptera
Scientific Names ARFI/YUGL
ATCA/SPAI
BOGR/HIJA
GLME/FRJA
JUMO/BOER
OPIM/BOGR
PIPO/SONU
PODE/BRJA TA sp.
Hesperotettix speciosus
Hesperotettix viridis
Hesperotettix wyomingensis
Hippiscus ocelote
Hippopedon capito
Hypochlora alba
Melanoplus angustipennis
Melanoplus arizonae
Melanoplus bivittatus
Melanoplus borealis
Melanoplus bowditchi
Melanoplus confusus
Melanoplus differentialis
Melanoplus discolor
Melanoplus dodgei
Melanoplus fasciatus
Melanoplus femurrubrum
Melanoplus flavidus
Melanoplus foedus
Melanoplus gladstoni
Melanoplus glaucipes
Melanoplus lakinus
Melanoplus occidentalis
Melanoplus packardii
Melanoplus ponderosus
Melanoplus regalis
Melanoplus sanguinipes
Melanoplus sp.
Melanoplus splendidus
Melanoplus tristis
Melanoplus yarrowii
Mermiria bivittata
Orthoptera
Scientific Names ARFI/YUGL
ATCA/SPAI
BOGR/HIJA
GLME/FRJA
JUMO/BOER
OPIM/BOGR
PIPO/SONU
PODE/BRJA TA sp.
Mermiria picta
Mestobregma plattei
Metator pardalinus
Microentrum sp.
Opeia obscura
Orphulella speciosa
Pardalophora apiculata
Paropomala wyomingensis
Phlibostroma quadrimaculatum
Phoetaliotes nebrascensis
Pseudophyllinae sp.
Pseudopomala brachyptera
Psoloessa delicatula
Psoloessa texana
Rhaphidophoridae sp.
Schistocerca obscura
Scudderia furcata
Spharagemon collare
Spharagemon equale
Stethophyma gracile
Syrbula admirabilis
Syrbula montezuma
Tetrix ornata
Trachyrhachys coronata
Trachyrhachys kiowa
Trimerotropis gracilis
Trimerotropis latifasciata
Trimerotropis melanoptera
Trimerotropis pallidipennis
Trimerotropis sp.
Tropidolophus formosus
Orthoptera
Scientific Names ARFI/YUGL
ATCA/SPAI
BOGR/HIJA
GLME/FRJA
JUMO/BOER
OPIM/BOGR
PIPO/SONU
PODE/BRJA TA sp.
Xanthippus corallipes
Hymenoptera
Species ARFI/YUGL
ATCA/SPAI
BOGR/HIJA
GLME/FRJA
JUMO/BOER
OPIM/BOGR
PIPO/SONU
PODE/BRJA TA sp.
Agapostemon angelicus
Agapostemon angelicus
or texanus
Agapostemon sp.
Andrena confederata
Andrena helianthi
Andrena merriami
Anthophora sp. 1
Anthophora sp. 2
Anthophora sp. 3
Athidium sp.
Augochlora sp.
Augochloropsis sp.
Bombus sp. 1
Bombus spp.
Colletes spp.
Diadasia diminuta
Diadasia enavata
Diadasia olivacea
Diadasia rinconis rinconis
Dianthidium sp. 1
Dianthidium sp. 2
Dieunomia sp.
Eucera sp.
Eucera sp. 1
Eucera sp. 2
Hymenoptera
Species ARFI/YUGL
ATCA/SPAI
BOGR/HIJA
GLME/FRJA
JUMO/BOER
OPIM/BOGR
PIPO/SONU
PODE/BRJA TA sp.
Halictus ligatus
Halictus virgatellus or
provancheri
Holcopasites tegularis
Lasioglossum spp. 1
Lasioglossum spp. 2
Megachile amica
Megachile brevis
Megachile brevis
onobrychidis
Melecta sp.
Melissodes intorta
Melissodes sp. 1
Melissodes sp. 2
Neolarra verbesinae
Nomia sp.
Osmia spp.
Perdita spp.
Protandrena sp. 1
Protandrena sp. 2
Sphecodes sp.
Svastra obliqua
Tetralphiella sp.
Triepeolus sp.
Xenoglossa sp.
Asilidae
Species ARFI/ YUGL ATCA/SPAI BOGR/HIJA GLME/FRJA JUMO/BOER OPIM/BOGR
PIPO/SONU PODE/BRJA
"Efferia" spp.
Albibarbefferia albibarbis
Albibarbefferia bicolor
Albibarbefferia leucocoma
Albibarbefferia spp.
Aridefferia prattii
Aridefferia subarida
1
Asilus auriannulatus
Atomosia melanopogon
Atomosia nuda
4
Atomosia puella
Carinefferia costalis
Diogmites neoternatus
Efferia candida
Efferia spp.
Haplopogon sp.
Laphystia varipes
Lasiopogon quadrivittatus
Leptogaster brevicornis
Leptogaster weslacensis
Nerax beameri
Ospriocerus abdominalis
Philonicus arizonensis
1
Philonicus limpidipennis
6
Pogoniefferia bicaudata
Pogoniefferia helenae
Asilidae
Species ARFI/ YUGL ATCA/SPAI BOGR/HIJA GLME/FRJA
JUMO/BOER OPIM/BOGR
PIPO/SONU
PODE/ BRJA
Pogoniefferia texana
Proctacanthella exquisita
Psilocurus birdi
Psilocurus camposi
Scleropogon arizonensis
Stenopogon albibasis
Stichopogon sp. 1