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<ul><li><p>F O L I A E N T O M O L O G I C A H U N G A R I C A R O V A R T A N I K Z L E M N Y E K </p><p>X U X 1988 p. 123-151 </p><p>The Scientific Results of Hungarian Soil Zoological Expeditions in New Guinea. Coleoptera, Tenebrionidae: </p><p>Lagruni </p><p>By </p><p>O. M E R K L (Received November 1, 1987) </p><p>A b s t r a c t : A to ta l of 37 species col lected by Hungarian zoologists in Papua New Guinea is l i s t ed . Three new genera (Acu togr i a , K a i n d i l a g r i a . T o m o g r i a ) and 21 new species (Acutogr ia falcata, B o t h r i c h a r a i n e r s , B . i n t r i c a t a , B . i r i - descens, B . wau, Casnonidea ap ica l i s , C. ba loghi , C. caudata, C. demet r ida , C. dobodura. C. f lavipes , C. h y s t r i x , C. l o k s a i , C. pallens, C. punct i thorax, C. setosa. K a i n d i l a g r i a fo rc ipa ta , L a g r i a amethyst ina , L . l i gu la t a , L . sapphi- r i n a . T o m o g r i a per la ta) are descr ibed . W i t h 49 f igures . </p><p>Hungarian zoologists have v i s i t e d New Guinea seve ra l t imes . F i r s t , S. F E N I C H E L made f i e ldwork there , but i t ended by his t r ag ic death (1893-95), w i th an emphasis on b i r d s ; he col lec ted also many insects but no l ag r i ine specimens. Fo l l owing F E N I C H E L ' s i n t e r r u p t ed w o r k , L . BIR made extensive and meticulous co l lec t ings (1895- 1901) i n German New Guinea ("Kaiser Wi lhe lms land" ) of that t i m e . In his most valuable m a t e r i a l , some specimens of L a g r i i n i were also found - these have been w r i t t e n up by PIC (1956). </p><p>Between 1965 and 1980, D r . J . B A L O G H pe r fo rmed s ix expedit ions to Papua New Gui-j nea ( f o r m e r l y T e r r i t o r y of Papua and New Guinea). D r . I . LOKSA and D r . L . MCZR jointed h i m in one case each. The late D r . J . J . H . S Z E N T - I V A N Y (Adelaide, South A u s t r a l i a ) hac) an impor t an t share in the organiza t ion of these t r i p s and the co l lec t ing of m a t e r i a l . The Wau Ecology Ins t i tu te ( f o r m e r l y Rishop Museum F i e l d Station) was most ly the centre and s t a r t i n g place of these expedi t ions . </p><p>The present con t r ibu t ion submits the resul t s concerning the t enebr ion id t r ibe L a g r i i n i . | The m a t e r i a l compr i ses 106 specimens belonging to 37 species; of them, 21 are descr ibed hereunder as new and 3 of them represent new genera. The two O r e o g r i a species have recently been descr ibed in a generic r e v i s i o n ( M E R K L 1988), whi le two fur ther species r e m a i n un-, iden t i f i ed or undescr ibed for cer ta in reasons (see L i s t of species). </p><p>Since impor t an t ma te r i a l s of va r ious col lec t ings were available to me dur ing the des c r i p t i o n s , the specimens of col lec t ings other than those of J . B A L O G H were also taken into! account. These are indicated by f o u r - l e t t e r abbreviat ions in the text; the name of reposi tory; i s fol lowed by the name of cu ra to r or p r iva te co l l ec to r in the subsequent l i s t : </p><p>A N I C A u s t r a l i a n Na t iona l Insect Co l l ec t i on , CSIRO, Canber ra , A C T , A u s t r a l i a . D r . J . F . L A W R E N C E . ( </p><p>B M N H B r i t i s h Museum (Natura l H i s t o r y ) , London, Uni ted Kingdom. D r . R . J . W . A L D R I D G E . B P B M Bern ice Pauahi Bishop Museum, Honolulu , H a w a i ' i , U . S . A . D r . G . A . SAMUELSON.I CASC C a l i f o r n i a Academy of Sciences, Department of Entomology, San F ranc i sco , C A L , </p><p>U . S . A . D r . D . H . K A V A N A U G H . </p><p># -7th con t r ibu t ion to the knowledge of L a g r i i n i . </p></li><li><p>CIMCI Canadian Nat ional C o l l e c t i o n of Insect , Ottawa, O N T , Canada. D r . A . S M E T A N A . H N H M Hungarian N a t u r a l H i s t o r y Museum, Budapest, Hungary. HPCW Hornabrook ' s pr ivate c o l l e c t i o n . We l l i ng ton , New Zealand. D r . R . W . HORNABROOK </p><p>and D r . R . G . ORDISH. I F P E Ins t i tu t fr Pflanzenschutzforschung, Ebe r swa lde -F inow, German Democra t i c Re</p><p>publ ic . D r . L . D I E C K M A N N . MCZC Museum of Compara t ive Zoology, H a r v a r d U n i v e r s i t y , Cambr idge , MASS, U . S . A . </p><p>D r . A . F . N E W T O N , J r . _ . _ MHNG Musum d ' H i s t o i r e N a t u r e l l e , Geneva, Swi tzer land . D r . C I . B E S ' U C H E T . SAMA South A u s t r a l i a n Museum, Adela ide , SA, A u s t r a l i a . D r . E . G . M A T T H E W S . SPCB Sedlacek's pr ivate co l l e c t i on , B r o o k f i e l d , Q L D , A u s t r a l i a . M r . J . S E D L A C E K . UQIC U n i v e r s i t y of Queensland, Insect Co l l ec t ion , Br i sbane , Q L D , A u s t r a l i a . M s . M . A . </p><p>SCHNEIDER. USNM United States Nat iona l Museum, Smithsonian Ins t i tu t ion , Washington, DC, U . S . A . </p><p>D r . G. F . H E V E L . Z M H U Zoologisches Museum, Humbold t Universitt, B e r l i n , G e r m a n Democra t i c Republ ic . </p><p>D r . F . HLEKE. </p><p>In the case of species, at which the Hungar ian m a t e r i a l consis ts of females or s l igh t ly ; damaged males only , I endeavoured to designate holotypes f r o m the m a t e r i a l of the Be rn i ce I Pauahi Bishop Museum, as fa r as i t was possible . </p><p>In the desc r ip t ion of species , the abbrev ia t ion E I is applied to the r a t io of eye d iameter and in t e rocu la r distance in d o r s a l v iew. These measurements and the r e l a t ive lengths of antenni segments are values measured d i r e c t l y by the ocular m i c r o m e t e r . </p><p>The labe l data of holotypes are fu l ly c i ted . The text of one l abe l is in quotation marks and the di f ferent labels are separated by semico lon . Holotype labels are r ed wi th the fo l lowing text: Holotypus , sex m a r k , generic and species name, M e r k l , 1988. </p><p>Data of paratypes are g iven i n the sequence of l o c a l i t y , a l t i tude ( i f poss ib l e ) , t i m e of co l l ec t ing , co l l ec t ing method ( i f possible) , name of co l l ec to r ( s ) . The number of specimens according to sexes and the abbrev ia t ion of r epos i t o ry are in parentheses. The paratype specimens are grouped in accordance of provinces of Papua New Guinea. Specimens co l lec ted in the western par t of New Guinea mainland, ( I r i an Jaya) are neglected in this paper and never designated as paratypes. Paratype labels are y e l l o w , w i t h the text as i n the holotype labels (of course, w i t h Paratypus instead of Holotypus) . </p><p>To save space, the fo l lowing fu r ther abbreviat ions and condensations seemed to be reasonable. On the one hand, an ove rwhe lming par t of the specimens was col lec ted by the S E D L A C E K ' s - the le t te rs JS, MS and JHS mean J . . M . and J . H . S E D L A C E K . On the other hand, by far the most specimens or ig ina te f r o m Wau (Morobe Prov ince ) and i t i s fel t superfluous to r ec i t e a l l - often qui te nearby - al t i tudes and days of co l l e c t i ng . T h e r e f o r e , the Wau specimens of the respect ive species are s u m m a r i z e d , wi th an i nd i ca t ion of lowest and h ighest a l t i tudines , the f i r s t and last dates of co l lec t ing and a l i s t of c o l l e c t o r s . </p><p>Acknowledgements . I am grea t ly indebted to a l l persons who provided me h igh ly va lu able l ag r i ine m a t e r i a l co l lec ted i n New Guinea and i t i s a pleasure to supply the i r col lec t ions w i t h type specimens. T h e i r names are indicated i n the l i s t of o r i g i n a l r epos i to r ies (see above).. My special thanks are due to M i s s gnes BARTQS for p repar ing the excellent habitus d raw- 1 </p><p>ings . </p><p>L I S T O F SPECIES </p><p>Codes between square brackets r e f e r to D r . J . B A L O G H ' s f i e ld notes s t i l l i n manus c r i p t ; numera l s in parentheses indicate number of specimens. Abbrev ia t ions : JB = J . B A LOGH ;.JJSZ = J . J . S Z E N T - I V A N Y . </p><p>L a g r i i n a </p><p>1. L a g r i a heurni Bo rchmann , 1924 - One specimen: A n g o r a m , 13-16. V I I I . 1969 [ N G A -U . 11. ] , J B . </p><p>2. L a g r i a p u p i l l i c o l l i s B o r c h m a n n , 1936 - One specimen: Haus Copper, Wau, M t . M i s -s i m , 22-24. I V . 1965, JB et JJSZ. </p></li><li><p>3. L a g r i a l i g u l a t a s p . n . - H o l o t y p e a n d 16 p a r a t y p e s : s e e d e s c r i p t i o n . 4. L a g r i a a m e t h y s t i n a s p . n. - O n e p a r a t y p e : s e e d e s c r i p t i o n . 5. L a g r i a s a p p h i r i n a s p . n . - H o l o t y p e and one p a r a t y p e : s e e d e s c r i p t i o n . 6. L a g r i a s p . n . - O n e s p e c i m e n s : M u s g r a v e r i v . v a l l . , c a . 55 k m N N E of P o r t M o </p><p>r e s b y . 31 . I I I . - 5 . I V . 1965 , J B et J J S Z . It i s a f e m a l e s p e c i m e n without any c h a r a c t e r i s t i c f e a t u r e . I n the l a c k of m a l e , i t w o u l d be an e x a g g e r a t i o n to d e s c r i b e i t . </p><p>7. O r e o g r i a l a r v a t a M e r k l , 1988 - 5 s p e c i m e n s : W a u , Mt . K a i n d i , 24 . DC. 1969 [ N G W -M . 2 6 . ] , J B (ho lo type , S and a o p a r a t y p e ) ; i d . [ N G W - U . 3 1 . ] (2 g p a r a t y p e s ) ; M t . K a i n d i , 2400 m , 1 5 - 1 6 . I V . 1965 , J D et J J S Z {Q_ p a r a t y p e ) . </p><p>8. O r e o g r i a p l i c a t a M e r k l , 1988 - O n e s p e c i m e n : H a u s C o p p e r , W a u . M t . M i s s i m , 2 2 - 2 4 . I V . 1965 , J B et J J S Z (ho lotype , o j . , </p><p>8. B o t h r i c h a r a m o d e s t a B o r c h m a n n , 1 9 1 5 - O n e s p e c i m e n : A n g o r a m , 1 3 - 1 6 . V I I I . 1 9 6 9 [ N G A - U . 1 1 . ) , J B . </p><p>9 . B o t h r i c h a r a p u l c h e l l a ( G u r i n - M n e v i l l e , 1830) - 8 s p e c i m e n s : V a n a p a R i v . , 32 m i l e s N of P o r t M o r e s b y , 2 . I V . 1965 , J B et J J S Z (1); B r o w n R i v . , 40 k m N of P o r t M o r e s b y , 6 - 8 . I V . 1965 , J B et J J S Z (1); W a u , M c A d a m P a r k , 1 8 - 2 1 . I V . 1965 , J B et J J S Z (1) ; H a u s C o p p e r . W a u , M t . M i s s i m , 2 2 - 2 4 . I V . 1965 , J B et J J S Z (1); W a u , bea ten f r o m l e a v e s , 8 - 9 . I X . 1968 [ N G W - C . 1 7 . ] , J B (1) ; W a u , K i l o l o C r e e k , 26 . V I I I . 1968 [ N G W - C . 14} , J B (1); B a i y e r R i v e r , S a n c t u a r y , 1 -5 . DC. 1969 [ N G B - U . 2 1 . ] , J B (1); i d . [ N G B - U . 2 3 . ] (1) . </p><p>10. B o t h r i c h a r a p a l l i a t a ( M a c l e a y , 1886) - O n e s p e c i m e n : B a i y e r R i v e r , S a n c t u a r y , 1 - 5 . DC. 1969 ( N G B . - U . 2 1 . 3 , J B . </p><p>11. B o t h r i c h a r a f u s c o p u b e n s B o r c h m a n n , 1936 - 3 s p e c i m e n s : W a u , B i s h o p M u s e u m F i e l d S t a t i o n , 1 5 - 2 5 . I V . 1965 , J B et J J S Z (1); W a u , M c A d a m P a r k , 1 8 - 2 1 . I V . 1 9 6 5 . J B et J J S Z (1); W a u , beaten f r o m l e a v e s , 8 - 9 . DC. 1968 [ N G W - C . 1 7 ] , J B (1) . </p><p>12. B o t h r i c h a r a i n s i g n i s B o r c h m a n n , 1936 - O n e s p e c i m e n : W a u , E d d y C r e e k , 20 50 m , 30 . V I I I . 1968 [ N G W - C . 1 3 . ] , J B . </p><p>13 . B o t h r i c h a r a i r i d e s c e n s s p . n . - H o l o t y p e : s e e d e s c r i p t i o n . 14. B o t h r i c h a r a i n e r s s p . n . - H o l o t y p e : s e e d e s c r i p t i o n . 15. B o t h r i c h a r a i n t r i c a t a s p . n . - O n e p a r a t y p e : s e e d e s c r i p t i o n . 16. B o t h r i c h a r a w a u s p . n . - 2 p a r a t y p e s : s e e d e s c r i p t i o n . 17. B o t h r i c h a r a s p . n . - 6 s p e c i m e n s : H a u s C o p p e r , W a u , M t . M i s s i m , 22-24 . I V . 1965 . </p><p>J B et J J S Z . T h e s e s i x f e m a l e s c e r t a i n l y r e p r e s e n t a n e w s p e c i e s . M a l e s f r o m o t h e r c o l l e c t i o n s m a y be a s s o c i a t e d wi th t h e s e f e m a l e s , t h e s e a r e g r o u p e d in to two s y m p a t r i c s p e c i e s s h a r p l y d i f f e r i n g f r o m e a c h o t h e r . F e m a l e s , h o w e v e r , s e e m to be e n t i r e l y u n i f o r m a n d , at the m o m e n t , I c a n n o t p l a c e t h e m in e i t h e r one of the g r o u p s of m a l e s . </p><p>18. T o m o g r i a p e r l a t a gen. et s p . n . - 7 p a r a t y p e s : s e e d e s c r i p t i o n . 19. A c u t o g r i a f a l c a t a gen. et s p . n . - H o l o t y p e and 3 p a r a t y p e s : s e e d e s c r i p t i o n . 20 . K a i n d i l a g r i a f o r c i p a t a gen . et sp. 1 n . - O n e p a r a t y p e : s e e d e s c r i p t i o n . </p><p>S t a t i r i n a </p><p>2 1 . S o r a b i r o i P i c , 1956 - O n e s p e c i m e n : B u s s u R i v . , L a e , 1 3 . I V . 1965 , J B et J J S Z . 2 2 . C a s n o n i d e a s e m i s u t u r a t a F a i r m a i r e , 1896 - 9 s p e c i m e n s : H a u s C o p p e r , W a u , Mt; </p><p>M i s s i m , 2 2 - 2 4 . I V . 1965 , J B et J J S Z (7); L a l o k i , 15 k m N of P o r t M o r e s b y , 30 . I I I . 1965 , J B et J J S Z (1); P o r t M o r e s b y , 2 - 3 . I V . 1965 , J B et J J S Z (1) . </p><p>2 3 . C a s n o n i d e a q u a d r i l i n e a t a ( B o r c h m a n n , 1936) - 4 s p e c i m e n s : W a u , 8. DC. 1968 ! N G W -R . 1 9 ! , I . L o k s a (2); i d . , 2 2 - 3 0 . DC. 1969 [ N G W - U . 3 0 . ] , J B (1); W a u , M t . K a i n d i , 1 9 - 2 4 . V I I I . 1969 [ N G W - U . 1 6 ] , J B (1) . </p><p>24 . C a s n o n i d e a v a r i a b i l i s ( M a c l e a y , 1 8 8 7 ) - O n e s p e c i m e n : S u n s h i n e P l a n t . , B u l o l o v a i l . , 20 . I V . 1965 , J B et J J S Z . </p><p>2 5 . C a s n o n i d e a d i m i d i a f a F a i r m a i r e , ! 1887 - O n e s p e c i m e n : K i u n g a . 2 3 . V I I - 2 . V I I I . 1969 [ N G K - U ( + M ) . 2 . ] . J B . </p><p>2 6 . C a s n o n i d e a s e m i v i o l a c e a ( P i c , 1912) T O n e s p e c i m e n : W a u , B i s h o p M u s e u m F i e l d S t a t i o n , 1 5 - 2 5 . I V . 1965 , J B et J J S Z . </p><p>2 7 . C a n o n i d e a h y s t r i x s p . n . - H o l o t y p e a n d 7 p a r a t y p e s : s e e d e s c r i p t i o n . 2 8 . C a s n o n i d e a p u n c t i t h o r a x s p . n . - 2 p a r a t y p e s : s e e d e s c r i p t i o n . 2 9 . C a s n o n i d e a a p i c a l i s s p . n . - H o l p t y p e : s e e d e s c r i p t i o n . 30. C a s n o n i d e a f l a v i p e s s p . n . - H o l o t y p e : s e e d e s c r i p t i o n . </p></li><li><p>31 . Casnonidea baloghi sp. n. - One para type: see desc r ip t ion . 32. Casnonidea loksa i sp. n . - One paratype: see desc r ip t i on . 33. Casnonidea dobodura sp. n . - One paratype: see desc r ip t i on . 34. Casnonidea caudata sp. n . - Ho lo type : see desc r ip t i on . 35. Casnonidea pallens sp. n . - 2 para types: see desc r ip t ion . 36. Casnonidea demet r ida sp. n . - Holotype and 3 paratypes: see desc r ip t ion . 37. Casnonidea setosa sp. n . - Holo type : see desc r ip t i on . </p><p>Notes on and r e d e s c r i p t i o n of L a g r i a p u p i l l i c o l l i s B o r c h m a n n . 1936 </p><p>BORCHMANN (1936:43) descr ibed this...</p></li></ul>