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FOLIA ENTOMOLOGICA HUNGARICA ROVARTANI KÖZLEMÉNYEK XUX 1988 p. 123-151 The Scientific Results of Hungarian Soil Zoological Expeditions in New Guinea. Coleoptera, Tenebrionidae: Lagruni By O. MERKL (Received November 1, 1987) Abstract: A total of 37 species collected by Hungarian zoologists in Papua New Guinea is listed. Three new genera (Acutogria, Kaindilagria. Tomogria) and 21 new species (Acutogria falcata, Bothrichara iners, B. intricata, B. iri- descens, B. wau, Casnonidea apicalis, C. baloghi, C. caudata, C. demetrida, C. dobodura. C. flavipes, C. hystrix, C. loksai, C. pallens, C. punctithorax, C. setosa. Kaindilagria forcipata, Lagria amethystina, L. ligulata, L. sapphi- rina. Tomogria perlata) are described. With 49 figures. Hungarian zoologists have visited New Guinea several times. First, S. FENICHEL made fieldwork there, but it ended by his tragic death (1893-95), with an emphasis on birds; he collected also many insects but no lagriine specimens. Following FENICHEL's interrupt- ed work, L. BIRÓ made extensive and meticulous collectings (1895- 1901) in German New Guinea ("Kaiser Wilhelmsland") of that time. In his most valuable material, some speci- mens of Lagriini were also found - these have been written up by PIC (1956). Between 1965 and 1980, Dr. J. BALOGH performed six expeditions to Papua New Gui-j nea (formerly Territory of Papua and New Guinea). Dr. I. LOKSA and Dr. L. MÓCZÁR joint- ed him in one case each. The late Dr. J. J.H. SZENT-IVANY (Adelaide, South Australia) hac) an important share in the organization of these trips and the collecting of material. The Wau Ecology Institute (formerly Rishop Museum Field Station) was mostly the centre and starting place of these expeditions. The present contribution submits the results concerning the tenebrionid tribe Lagriini.| The material comprises 106 specimens belonging to 37 species; of them, 21 are described hereunder as new and 3 of them represent new genera. The two Oreogria species have recent- ly been described in a generic revision (MERKL 1988), while two further species remain un-, identified or undescribed for certain reasons (see List of species). Since important materials of various collectings were available to me during the de- scriptions, the specimens of collectings other than those of J. BALOGH were also taken into! account. These are indicated by four-letter abbreviations in the text; the name of repository; is followed by the name of curator or private collector in the subsequent list: ANIC Australian National Insect Collection, CSIRO, Canberra, ACT, Australia. Dr. J. F. LAWRENCE. ( BMNH British Museum (Natural History), London, United Kingdom. Dr. R.J.W. ALDRIDGE. BPBM Bernice Pauahi Bishop Museum, Honolulu, Hawai'i, U.S.A. Dr. G.A. SAMUELSON.I CASC California Academy of Sciences, Department of Entomology, San Francisco, CAL, U.S.A. Dr. D.H. KAVANAUGH. # - 7th contribution to the knowledge of Lagriini.

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F O L I A E N T O M O L O G I C A H U N G A R I C A R O V A R T A N I K Ö Z L E M É N Y E K

X U X 1988 p. 123-151

The Scientific Results of Hungarian Soil Zoological Expeditions in New Guinea. Coleoptera, Tenebrionidae:

Lagruni

By

O. M E R K L (Received November 1, 1987)

A b s t r a c t : A to ta l of 37 species col lected by Hungarian zoologists in Papua New Guinea is l i s t ed . Three new genera (Acu togr i a , K a i n d i l a g r i a . T o m o g r i a ) and 21 new species (Acutogr ia falcata, B o t h r i c h a r a i n e r s , B . i n t r i c a t a , B . i r i - descens, B . wau, Casnonidea ap ica l i s , C. ba loghi , C. caudata, C. demet r ida , C. dobodura. C. f lavipes , C. h y s t r i x , C. l o k s a i , C. pallens, C. punct i thorax, C. setosa. K a i n d i l a g r i a fo rc ipa ta , L a g r i a amethyst ina , L . l i gu la t a , L . sapphi- r i n a . T o m o g r i a per la ta) are descr ibed . W i t h 49 f igures .

Hungarian zoologists have v i s i t e d New Guinea seve ra l t imes . F i r s t , S. F E N I C H E L made f i e ldwork there , but i t ended by his t r ag ic death (1893-95), w i th an emphasis on b i r d s ; he col lec ted also many insects but no l ag r i ine specimens. Fo l l owing F E N I C H E L ' s i n t e r r u p t ­ed w o r k , L . BIRÓ made extensive and meticulous co l lec t ings (1895- 1901) i n German New Guinea ("Kaiser Wi lhe lms land" ) of that t i m e . In his most valuable m a t e r i a l , some speci­mens of L a g r i i n i were also found - these have been w r i t t e n up by PIC (1956).

Between 1965 and 1980, D r . J . B A L O G H pe r fo rmed s ix expedit ions to Papua New Gui-j nea ( f o r m e r l y T e r r i t o r y of Papua and New Guinea). D r . I . LOKSA and D r . L . MÓCZÁR joint­ed h i m in one case each. The late D r . J . J . H . S Z E N T - I V A N Y (Adelaide, South A u s t r a l i a ) hac) an impor t an t share in the organiza t ion of these t r i p s and the co l lec t ing of m a t e r i a l . The Wau Ecology Ins t i tu te ( f o r m e r l y Rishop Museum F i e l d Station) was most ly the centre and s t a r t i n g place of these expedi t ions .

The present con t r ibu t ion submits the resul t s concerning the t enebr ion id t r ibe L a g r i i n i . | The m a t e r i a l compr i ses 106 specimens belonging to 37 species; of them, 21 are descr ibed hereunder as new and 3 of them represent new genera. The two O r e o g r i a species have recent­ly been descr ibed in a generic r e v i s i o n ( M E R K L 1988), whi le two fur ther species r e m a i n un-, iden t i f i ed or undescr ibed for cer ta in reasons (see L i s t of species).

Since impor t an t ma te r i a l s of va r ious col lec t ings were available to me dur ing the de­s c r i p t i o n s , the specimens of col lec t ings other than those of J . B A L O G H were also taken into! account. These are indicated by f o u r - l e t t e r abbreviat ions in the text; the name of reposi tory; i s fol lowed by the name of cu ra to r or p r iva te co l l ec to r in the subsequent l i s t :

A N I C A u s t r a l i a n Na t iona l Insect Co l l ec t i on , CSIRO, Canber ra , A C T , A u s t r a l i a . D r . J . F . L A W R E N C E . (

B M N H B r i t i s h Museum (Natura l H i s t o r y ) , London, Uni ted Kingdom. D r . R . J . W . A L D R I D G E . B P B M Bern ice Pauahi Bishop Museum, Honolulu , H a w a i ' i , U . S . A . D r . G . A . SAMUELSON.I CASC C a l i f o r n i a Academy of Sciences, Department of Entomology, San F ranc i sco , C A L ,

U . S . A . D r . D . H . K A V A N A U G H .

# -7th con t r ibu t ion to the knowledge of L a g r i i n i .

CIMCI Canadian Nat ional C o l l e c t i o n of Insect , Ottawa, O N T , Canada. D r . A . S M E T A N A . H N H M Hungarian N a t u r a l H i s t o r y Museum, Budapest, Hungary. HPCW Hornabrook ' s pr ivate c o l l e c t i o n . We l l i ng ton , New Zealand. D r . R . W . HORNABROOK

and D r . R . G . ORDISH. I F P E Ins t i tu t für Pflanzenschutzforschung, Ebe r swa lde -F inow, German Democra t i c Re­

publ ic . D r . L . D I E C K M A N N . MCZC Museum of Compara t ive Zoology, H a r v a r d U n i v e r s i t y , Cambr idge , MASS, U . S . A .

D r . A . F . N E W T O N , J r . _ . _ MHNG Muséum d ' H i s t o i r e N a t u r e l l e , Geneva, Swi tzer land . D r . C I . B E S ' U C H E T . SAMA South A u s t r a l i a n Museum, Adela ide , SA, A u s t r a l i a . D r . E . G . M A T T H E W S . SPCB Sedlacek's pr ivate co l l e c t i on , B r o o k f i e l d , Q L D , A u s t r a l i a . M r . J . S E D L A C E K . UQIC U n i v e r s i t y of Queensland, Insect Co l l ec t ion , Br i sbane , Q L D , A u s t r a l i a . M s . M . A .

SCHNEIDER. USNM United States Nat iona l Museum, Smithsonian Ins t i tu t ion , Washington, DC, U . S . A .

D r . G. F . H E V E L . Z M H U Zoologisches Museum, Humbold t Universität, B e r l i n , G e r m a n Democra t i c Republ ic .

D r . F . HLEKE.

In the case of species, at which the Hungar ian m a t e r i a l consis ts of females or s l igh t ly ; damaged males only , I endeavoured to designate holotypes f r o m the m a t e r i a l of the Be rn i ce I Pauahi Bishop Museum, as fa r as i t was possible .

In the desc r ip t ion of species , the abbrev ia t ion E I is applied to the r a t io of eye d iameter and in t e rocu la r distance in d o r s a l v iew. These measurements and the r e l a t ive lengths of an­tennái segments are values measured d i r e c t l y by the ocular m i c r o m e t e r .

The labe l data of holotypes are fu l ly c i ted . The text of one l abe l is in quotation marks and the di f ferent labels are separated by semico lon . Holotype labels are r ed wi th the fo l low­ing text: Holotypus , sex m a r k , generic and species name, M e r k l , 1988.

Data of paratypes are g iven i n the sequence of l o c a l i t y , a l t i tude ( i f poss ib l e ) , t i m e of co l l ec t ing , co l l ec t ing method ( i f possible) , name of co l l ec to r ( s ) . The number of specimens according to sexes and the abbrev ia t ion of r epos i t o ry are in parentheses. The paratype speci­mens are grouped in accordance of provinces of Papua New Guinea. Specimens co l lec ted in the western par t of New Guinea mainland, ( I r i an Jaya) are neglected in this paper and never designated as paratypes. Paratype labels are y e l l o w , w i t h the text as i n the holotype labels (of course, w i t h Paratypus instead of Holotypus) .

To save space, the fo l lowing fu r ther abbreviat ions and condensations seemed to be reasonable. On the one hand, an ove rwhe lming par t of the specimens was col lec ted by the S E D L A C E K ' s - the le t te rs JS, MS and JHS mean J . . M . and J . H . S E D L A C E K . On the other hand, by far the most specimens or ig ina te f r o m Wau (Morobe Prov ince ) and i t i s fel t super­fluous to r ec i t e a l l - often qui te nearby - al t i tudes and days of co l l e c t i ng . T h e r e f o r e , the Wau specimens of the respect ive species are s u m m a r i z e d , wi th an i nd i ca t ion of lowest and h igh­est a l t i tudines , the f i r s t and last dates of co l lec t ing and a l i s t of c o l l e c t o r s .

Acknowledgements . I am grea t ly indebted to a l l persons who provided me h igh ly va lu ­able l ag r i ine m a t e r i a l co l lec ted i n New Guinea and i t i s a pleasure to supply the i r col lec t ions w i t h type specimens. T h e i r names are indicated i n the l i s t of o r i g i n a l r epos i to r ies (see above).. My special thanks are due to M i s s Ágnes BARTQS for p repar ing the excellent habitus d raw- 1

ings .

L I S T O F SPECIES

Codes between square brackets r e f e r to D r . J . B A L O G H ' s f i e ld notes s t i l l i n manu­s c r i p t ; numera l s in parentheses indicate number of specimens. Abbrev ia t ions : JB = J . B A ­LOGH ;.JJSZ = J . J . S Z E N T - I V A N Y .

L a g r i i n a

1. L a g r i a heurni Bo rchmann , 1924 - One specimen: A n g o r a m , 13-16. V I I I . 1969 [ N G A -U . 11. ] , J B .

2. L a g r i a p u p i l l i c o l l i s B o r c h m a n n , 1936 - One specimen: Haus Copper, Wau, M t . M i s -s i m , 22-24. I V . 1965, JB et JJSZ.

3. L a g r i a l i g u l a t a s p . n . - H o l o t y p e a n d 16 p a r a t y p e s : s e e d e s c r i p t i o n . 4. L a g r i a a m e t h y s t i n a s p . n. - O n e p a r a t y p e : s e e d e s c r i p t i o n . 5. L a g r i a s a p p h i r i n a s p . n . - H o l o t y p e and one p a r a t y p e : s e e d e s c r i p t i o n . 6. L a g r i a s p . n . - O n e s p e c i m e n s : M u s g r a v e r i v . v a l l . , c a . 55 k m N N E of P o r t M o ­

r e s b y . 31 . I I I . - 5 . I V . 1965 , J B et J J S Z . It i s a f e m a l e s p e c i m e n without any c h a r a c t e r i s t i c f e a t u r e . I n the l a c k of m a l e , i t w o u l d be an e x a g g e r a t i o n to d e s c r i b e i t .

7. O r e o g r i a l a r v a t a M e r k l , 1988 - 5 s p e c i m e n s : W a u , Mt . K a i n d i , 24 . DC. 1969 [ N G W -M . 2 6 . ] , J B (ho lo type , S and a o p a r a t y p e ) ; i d . [ N G W - U . 3 1 . ] (2 g p a r a t y p e s ) ; M t . K a i n d i , 2400 m , 1 5 - 1 6 . I V . 1965 , J D et J J S Z {Q_ p a r a t y p e ) .

8. O r e o g r i a p l i c a t a M e r k l , 1988 - O n e s p e c i m e n : H a u s C o p p e r , W a u . M t . M i s s i m , 2 2 - 2 4 . I V . 1965 , J B et J J S Z (ho lotype , o j . ,

8. B o t h r i c h a r a m o d e s t a B o r c h m a n n , 1 9 1 5 - O n e s p e c i m e n : A n g o r a m , 1 3 - 1 6 . V I I I . 1 9 6 9 [ N G A - U . 1 1 . ) , J B .

9 . B o t h r i c h a r a p u l c h e l l a ( G u é r i n - M é n e v i l l e , 1830) - 8 s p e c i m e n s : V a n a p a R i v . , 32 m i l e s N of P o r t M o r e s b y , 2 . I V . 1965 , J B et J J S Z (1); B r o w n R i v . , 40 k m N of P o r t M o r e s b y , 6 - 8 . I V . 1965 , J B et J J S Z (1); W a u , M c A d a m P a r k , 1 8 - 2 1 . I V . 1965 , J B et J J S Z (1) ; H a u s C o p p e r . W a u , M t . M i s s i m , 2 2 - 2 4 . I V . 1965 , J B et J J S Z (1); W a u , bea ten f r o m l e a v e s , 8 - 9 . I X . 1968 [ N G W - C . 1 7 . ] , J B (1) ; W a u , K i l o l o C r e e k , 26 . V I I I . 1968 [ N G W - C . 14} , J B (1); B a i y e r R i v e r , S a n c t u a r y , 1 -5 . DC. 1969 [ N G B - U . 2 1 . ] , J B (1); i d . [ N G B - U . 2 3 . ] (1) .

10. B o t h r i c h a r a p a l l i a t a ( M a c l e a y , 1886) - O n e s p e c i m e n : B a i y e r R i v e r , S a n c t u a r y , 1 - 5 . DC. 1969 ( N G B . - U . 2 1 . 3 , J B .

11. B o t h r i c h a r a f u s c o p u b e n s B o r c h m a n n , 1936 - 3 s p e c i m e n s : W a u , B i s h o p M u s e u m F i e l d S t a t i o n , 1 5 - 2 5 . I V . 1965 , J B et J J S Z (1); W a u , M c A d a m P a r k , 1 8 - 2 1 . I V . 1 9 6 5 . J B et J J S Z (1); W a u , beaten f r o m l e a v e s , 8 - 9 . DC. 1968 [ N G W - C . 1 7 ] , J B (1) .

12. B o t h r i c h a r a i n s i g n i s B o r c h m a n n , 1936 - O n e s p e c i m e n : W a u , E d d y C r e e k , 20 50 m , 30 . V I I I . 1968 [ N G W - C . 1 3 . ] , J B .

13 . B o t h r i c h a r a i r i d e s c e n s s p . n . - H o l o t y p e : s e e d e s c r i p t i o n . 14. B o t h r i c h a r a i n e r s s p . n . - H o l o t y p e : s e e d e s c r i p t i o n . 15. B o t h r i c h a r a i n t r i c a t a s p . n . - O n e p a r a t y p e : s e e d e s c r i p t i o n . 16. B o t h r i c h a r a w a u s p . n . - 2 p a r a t y p e s : s e e d e s c r i p t i o n . 17. B o t h r i c h a r a s p . n . - 6 s p e c i m e n s : H a u s C o p p e r , W a u , M t . M i s s i m , 22-24 . I V . 1965 .

J B et J J S Z . T h e s e s i x f e m a l e s c e r t a i n l y r e p r e s e n t a n e w s p e c i e s . M a l e s f r o m o t h e r c o l l e c ­t i o n s m a y be a s s o c i a t e d wi th t h e s e f e m a l e s , t h e s e a r e g r o u p e d in to two s y m p a t r i c s p e c i e s s h a r p l y d i f f e r i n g f r o m e a c h o t h e r . F e m a l e s , h o w e v e r , s e e m to be e n t i r e l y u n i f o r m a n d , at the m o m e n t , I c a n n o t p l a c e t h e m in e i t h e r one of the g r o u p s of m a l e s .

18. T o m o g r i a p e r l a t a gen. et s p . n . - 7 p a r a t y p e s : s e e d e s c r i p t i o n . 19. A c u t o g r i a f a l c a t a gen. et s p . n . - H o l o t y p e and 3 p a r a t y p e s : s e e d e s c r i p t i o n . 20 . K a i n d i l a g r i a f o r c i p a t a gen . et sp. 1 n . - O n e p a r a t y p e : s e e d e s c r i p t i o n .

S t a t i r i n a

2 1 . S o r a b i r o i P i c , 1956 - O n e s p e c i m e n : B u s s u R i v . , L a e , 1 3 . I V . 1965 , J B et J J S Z . 2 2 . C a s n o n i d e a s e m i s u t u r a t a F a i r m a i r e , 1896 - 9 s p e c i m e n s : H a u s C o p p e r , W a u , Mt;

M i s s i m , 2 2 - 2 4 . I V . 1965 , J B et J J S Z (7); L a l o k i , 15 k m N of P o r t M o r e s b y , 30 . I I I . 1965 , J B et J J S Z (1); P o r t M o r e s b y , 2 - 3 . I V . 1965 , J B et J J S Z (1) .

2 3 . C a s n o n i d e a q u a d r i l i n e a t a ( B o r c h m a n n , 1936) - 4 s p e c i m e n s : W a u , 8. DC. 1968 ! N G W -R . 1 9 ! , I . L o k s a (2); i d . , 2 2 - 3 0 . DC. 1969 [ N G W - U . 3 0 . ] , J B (1); W a u , M t . K a i n d i , 1 9 - 2 4 . V I I I . 1969 [ N G W - U . 1 6 ] , J B (1) .

24 . C a s n o n i d e a v a r i a b i l i s ( M a c l e a y , 1 8 8 7 ) - O n e s p e c i m e n : S u n s h i n e P l a n t . , B u l o l o v a i l . , 20 . I V . 1965 , J B et J J S Z .

2 5 . C a s n o n i d e a d i m i d i a f a F a i r m a i r e , ! 1887 - O n e s p e c i m e n : K i u n g a . 2 3 . V I I - 2 . V I I I . 1969 [ N G K - U ( + M ) . 2 . ] . J B .

2 6 . C a s n o n i d e a s e m i v i o l a c e a ( P i c , 1912) T O n e s p e c i m e n : W a u , B i s h o p M u s e u m F i e l d S t a t i o n , 1 5 - 2 5 . I V . 1965 , J B et J J S Z .

2 7 . C a n o n i d e a h y s t r i x s p . n . - H o l o t y p e a n d 7 p a r a t y p e s : s e e d e s c r i p t i o n . 2 8 . C a s n o n i d e a p u n c t i t h o r a x s p . n . - 2 p a r a t y p e s : s e e d e s c r i p t i o n . 2 9 . C a s n o n i d e a a p i c a l i s s p . n . - H o l p t y p e : s e e d e s c r i p t i o n . 30. C a s n o n i d e a f l a v i p e s s p . n . - H o l o t y p e : s e e d e s c r i p t i o n .

31 . Casnonidea baloghi sp. n. - One para type: see desc r ip t ion . 32. Casnonidea loksa i sp. n . - One paratype: see desc r ip t i on . 33. Casnonidea dobodura sp. n . - One paratype: see desc r ip t i on . 34. Casnonidea caudata sp. n . - Ho lo type : see desc r ip t i on . 35. Casnonidea pallens sp. n . - 2 para types: see desc r ip t ion . 36. Casnonidea demet r ida sp. n . - Holotype and 3 paratypes: see desc r ip t ion . 37. Casnonidea setosa sp. n . - Holo type : see desc r ip t i on .

Notes on and r e d e s c r i p t i o n of L a g r i a p u p i l l i c o l l i s B o r c h m a n n . 1936

BORCHMANN (1936:43) descr ibed this species f r o m "Holländisch-Neu-Guinea" on the basis of an unrecorded number of specimens and without reference to the type depos i tory . I t i s v e r y l i k e l y that the types 'were included i n B O R C H M A N N ' s co l lec t ion and were des t roy­ed dur ing W o r l d War I I . I t is c lear f r o m the desc r ip t ion that he knew both sexes and some cha rac t e r i s t i c features leave l i t t l e doubt what species he had. The desc r ip t ion was based on specimens wi th b lu i sh e l y t r a . The e l y t r a l pa t t e rn , however , is ac tual ly much more v a r y i n g , and the un ico loured dark blue e l y t r a represen t m e r e l y an (although frequent) ex t reme of the spec t rum of v a r i a b i l i t y . An in te rmed ia te and c h a r a c t e r i s t i c pat tern is the e l y t r a having a

F igs 1-2. 1= L a g r i a l igu la ta sp. n . , | o \ - j 2= L . p u p i l l i c o l l i s Borchmann , <?

pale spot at the midd la ( F i g . 2) . The spots,! on the one hand, may extend to the overwhelming par t of the e l y t r a and only the base and apex r e m a i n b lu i sh . On the other hand, the spots may be da rke r and obsolete, and this t r end leads to the un ico loured dark e l y t r a .

T w o of the new species descr ibed subsequently ( L a g r i a amethyst ina and L . sapphir ina) appear to be c lose ly a l l i e d to L a g r i a pup i l l i co l l i s 'Borchmann , but the l a t t e r is eas i ly d i s t i n ­guished at the f i r s t glance by the dark green or blue p rono tum as w e l l as dark legs and an­tennae.

Redesc r ip t ion : Head, p rono tum and s c u t e l l u m dark olivaceous green to blue wi th strong me ta l l i c sheen; e l y t r a pu rp l i sh red to blue; w i t h a l a rge , ovoid ochreous spot at the middle ; th is spot leaving su tu ra i m a r g i n dark , n a r r o w e s t at inner side, g radual ly widening toward e l y t r a l m a r g i n , an te r io r border nea r ly h o r i z o n t a l , pos te r io r bo rde r oblique; spot may often be expanded or reduced to disappear; v e n t r a l surface and legs black; antennae dark reddish brown to black. Length: 6 .9 -10 .0 m m .

S. Head l a rge ly impunctate between eyes and on ve r t ex ; w i t h s m a l l pi ts between eyesji ve r tex w i t h 2 blunt , smooth protuberance standing near each other ; temples coarse ly punc-j täte; surface wi th moderate ly long , e rec t h a i r s ; EI= 12:8; length r a t ios of antennái seg­ments : 9: 4: 8: 9: 7: 8: 7: 7: 6;5: 5: 24; no t r a n s v e r s a l or modif ied segments; segment I about as long as ha l f the distance between antennái sockets . P rono tum a l i t t l e t r ansve r se ; widest j u s t a n t e r i o r to middle ; sides weakly arched; punctures coarse , v e r y close, somewhat con­fluent p o s t e r i o r l y ; wi th 2 i n d i s t i n c t , t r a n s v e r s a l impress ions before base; pubescence mod­era te ly long and sparse, e rec t . E l y t r a elongate b road ly , ovoid , convex; punctation moder­ately coarse , near ly u n i f o r m , in terspaces not p l i ca te ; pubescence ra ther shor t , r a the r dense, e rec t . Sterna without modi f i ca t ions . Legs n o r m a l , sparse ly pubescent; t ibiae un­a rmed . Aedeagus: F i g . 7. Habi tus : F i g . ; 2 .

Ç. L a r g e r on average; EI= 13: 13; length r a t i o s of antennái segments V I I I to X I : 7:5:7» 6: 16; smooth area of head less extended; p rono tum more t r ansve r se .

F igs 3-4. 3- L a g r i a amethyst ina sp. n T , 6. - 4= L . sapphi r ina sp. n . , #

Desc r ip t i on of new taxa

L a g r i a l igu la ta sp. n. (Figs 1, 5, 6)

Head, p rono tum and scu t e l l um da rk , shiny, m e t a l l i c green; e l y t r a b rownish w i t h s t rong greenish , pu rp l i sh or v io le t t inge; v e n t r a l surface redd i sh , meso- and metas ternum da rke r ; legs black; antennae l a rge ly y e l l o w i s h , ap ica l 4-5 segments g radua l ly da rke r . Length: 6 .5-10 _1 m m .

8, Head v e r y weakly i m p r e s s e d between eyes; coarse ly and sparse ly punctate; w i t h long , scat tered h a i r s ; eyes conspicuously l a rge , EI= 12: 5;ülength ra t ios of antenna! seg­ments: 7 :4 . 5 : 9 : 8 : 7 . 5: 1 0 : 9 : 9 : 7 . 5:23; segment I about as long as ha l f the distance between antenna! sockets; segment X I somewhat widened, v e r y weakly tw i s t ed , inner surface con­cave (F ig . 5). P rono tum as long as or a l i t t l e longer than broad; widest at the middle , s ides, v e r y weakly arched; punctures coarse , separated by interspaces as broad as puncture d ia­meter ; surface wi th 4 shallow i m p r e s s i o n s ; pubescence modera te ly long, erect . E l y t r a e lon­gate, feebly broadening, weakly convex; punctation coarse , modera te ly dense; interspaces s l i gh t ly ra i sed ; w i t h r a the r long, sparse pubescence cons is t ing of erec t and semierec t h a i r s . Sterna without modi f ica t ions . Legs t y p i c a l ; w i t h sparse , modera te ly long, semierec t h a i r s ; t ib iae una rmed . Aedeagus: F i g . 6. Habi tus : F i g . 1.

g. Considerably l a r g e r and broader on the average; EI= 1 5 : 1 1 ; length ra t ios of anten­nái segments V I I I to X I : 1 0 : 9 : 8 : 19; las t antennái segment s imp le ; p rono tum of same length and width or s l i g h t l y t r ansve r se ; e l y t r a much broader .

Holotype, 6, label led as fo l lows : "NEW GUINEA (NE) B a i y e r R i v e r Sanctuary 1-5. X I . 1969"; " (No. N G B - U . 24. ) leg . D r . J . Ba logh" . I t is deposited i n H N H M .

Paratypes (total 124 specimens) . Wes te rn Highlands P rov ince : label led as holotype | (3 68, 2 go., H N H M ) ; i d . [ N G B - U . 2 2 ^ (1 8, H N H M ) ; i d . [ N G B - U . 2 1 ] (1 8, HNHM); i d . £NGB-R . 8 . ] (1 o, H N H M ) ; i d . , 4000 ' , 6 -11 . V I I . 1974, H . Howden (1 8, 1 o, CNCI) ; J i m m i R i v e r , 4700 f t . . Ï6 . VI I -21 .DC. 1961, W . W . Brand t (1 6, 1 £. A N I C ) ; W u m , Upper J i m i V . , 840 m , 18. V I I . 1955, J . L . G r e s s i t t (3 og, B P B M ) . - Eas te rn Highlands P rov ince : K a r i m u i , 1080 m 8-10. V I I . 1963, JS (1 o, B P B M ) ; Okapa, Agakamatasa, 17.1.1965, R. Hornabrook (1 g, HPCW); Okapa, Okasa, 15. V . 1965, R. Hornabrook (1 g, HPCW); Okapa, Purosa, 7000 f t . , 23. V I . 1964, R. Hornabrook (1 o, HPCW). - Morobe P r o v i n c e : B u l o l o , 680 m , 4. V I . 1969, JS (2 go, B P B M ) ; i d . , 700 m , 18. V I . 1969, JS (1 8, B P B M ) ; i d . , 6. X I . 1969, JS (1 g, B P B M ) ; i d . , 26. X I . 1969, JS, MS (2 66, 3 oo, B P B M ; 1 0, H N H M ex B P B M ) ; 5 k m N Bulo lo , 630 m , 6. V I I I . 1964, JS, MS (1 6, B P B M ) ; Bu lo lo R. , 1Ï30 m , 17. I X . 1969, A . B . M i r z a , JS, MS (2 Z 8 , 1 o, B P B M ) ; F insch Haven, L . Wagner (3 oo, S A M A ) ; Gara ina , 830 m , 13-15.1.1968* JS. MS (2 oo, B P B M ) ; Huon Peninsula , P ind iu , 5ÓÓ-600 m , 19. I V . 1963, JS (1 6, B P B M ) ; i d . , 870-1300 m , 21-22. I V . 1963, JS (1 g. B P B M ) ; Umg. K a i a p i t , 30. I X . 1979, W . G . U l l r i c h (1 g, MHNG); Lae , 4-6 . D i . 1968 [ N G L - C . 1 6 . ] . J . Balogh (1 6, H N H M ) ; Watut R. , 900-1900 m , X . 1969, A . B . M i r z a (2 68, 1 g, B P B M ) ; Wau. 1100- 300 m . 16. X . 1961-3. V I I . 1974, R. Hon-nabrook. H . F . Howden, JS, JHS, MS (9 56, 6 oo, B P B M ; 5 55, C N C I ; 1 6, HPCW; 3 39, 2 Q 2 , H N H M ex B P B M , 1 Ó, H N H M ex CNCI) ; i d . , 22-30. I X . 1969 I N G W - U . 33. !, J . Balogh (3 oo, H N H M ) ; i d . , 2 5 . V I I I . 1968, beaten f r o m decayed leaves [ N G W - C . 10.3 , J . Balogh (1 oi H N H M ) ; Wau, B i g Wau Ck. , 1300 m , X I . 1965, P. Shanahan (1 6, B P B M ) ; Wau, Bu lo lo R. , ' 800-900 m , 3 1 . V I I I . 1965, JS (2 56, B P B M ) ; i d . , 850-950 m , 4. I I . 1966, JS ( l g , B P B M ) ; Wau, Hosp i ta l Ck. , 1150- 1250 m , 9.1.1966,; JS . iMS (1 6, B P B M ) ; Wau, Kaisenik , 1000 m , 1 9 . I V . 1965, JS, MS (1 o, B P B M ) ; Wau, Ka r inga Ck. , 950-1150 m , 3 0 . I X . 1 9 6 4 , MS (2 58. B P B M ; 1 S, H N H M ex B P B M ) ; Wau, KilokV Creek , 26. V I I I . 1968 [ N G W - C . 14.3, J . Balogh I (1 5, 1 o_, H N H M ) ; Wau, M t . K a i n d i , 24-25^ V I I I ; 1968 [ N G W - C . 9 . ] J . Balogh (1 g, H N H M ) ; ' Wau, M t . M i s s i m , 950-1300 m , 8-9.1.1966;, JS, : MS (1 8, 1 g, B P B M ) ; Wau, Wau Ck. , 1200-j 1500 m , 6 . V . 1963, JS (1 8, B P B M ) . - N o r t h e r n P rov ince : Dobodura, I I I - V I I . 1944, D a r l i n g ­ton (2 88, M C Z C ) ; Kokoda, 1200 f t . , V I I I . 1933, L . E . Cheesmann (1 g, B M N H ) ; i d . , DC.1933J (1 Q, B M N H ) ; i d . , 1300 f t . , V I I I . 1933, l ower r a i n f o r e s t , L . E . Cheesmann (2 oo. B M N H ) ; i d ] , 400 m , 1 4 - 1 6 . X I . 1965. JS, MS (1 8, 1 o, B P B M ) ; i d . , 1 8 . X I . 1966, JS (2 88, B P B M ) ; Mana-galase Plateau, V I I . 1972, R. Hornabrook (2, 88,'1 g, HPCW; 1 Ä, 1 p. H N H M ex HPCW); Mt . i Laming ton , 1300-1500 f t . . C . T . M c N a m a r a (6 88, 3 gg, SAMA; 2 88, 1 g, H N H M ex SAMA); ;

Popondetta, 29.1.1965, R. Hornabrook (1 g | HPCW); i d . , 27-28.11.1966, G. Monte i th (2 8 8 } 2 gg, UQIC) ; i d . , 60 m . 1-2. I X . 1963, JS ( l j g , B P B M ) . j

R e m a r k s : Males of L a g r i a l igu la ta may eas i ly be recognized by the s l igh t ly spoon-shap-

ed antennái segment X I . Females Have no d i s t inc t features that would d i s t ingu ish them f r o m the genera l ized females of other (par t ly undescribed) species belonging to L a g r i a and other genera of L a g r i i n a , though the lus t rous green forebody may help in the recogn i t ion .

L a g r i a amethyst ina sp. n . (Figs 3, 8)

Head, p rono tum, scu te l lum, legs and antennae y e l l o w i s h red ; e l y t r a pu rp l i sh v i o l e t ; v e n t r a l surface redd i sh , w i th metas te rnum and abdominal sterna more or less da rke r . Length: 6. 7-8. 2 m m .

3. Head impres sed between eyes; th is i m p r e s s i o n and 2 blunt, i nd i s t inc t p ro tuber ­ances near ly smooth; o therwise , head sparsely punctate w i th long erec t h a i r s ; EI= 12: 10; length ra t ios of antennái segments: 9 : 4 : 9 : 1 2 : 9 : 9 : 9 : 8 : 8 : 7 . 5 : 3 2 ; no t r a n s v e r s a l o r m o d i f i ­ed segments; segment I shor te r than half of distance between antennái sockets . P rono tum t r ansve r se ; widest jus t an te r io r to the midd le ; sides nea r ly p a r a l l e l ; punctures coarse and close, somewhat rugose before base; w i t h 2 to ta l ly i nd i s t i nc t t r ansve r sa l impress ions be­fore base; pubescence ra ther long and sparse , e r ec t . E l y t r a broadly ova l , modera te ly con­vex; punctat ion less coarse , u n i f o r m l y sca t tered , in terspaces not p l ica te ; pubescence shor t , dense, e rec t . Sterna without modi f ica t ions . Legs t y p i c a l , w i th sparse, long pubescence; 1

t ib iae una rmed . Aedeagus: F i g . 8. Habi tus : F i g . 3. o. L a r g e r on average; EI= 12: 15; length r a t ios of antenna! segments V I I I to X I : 8:8:

7: 19; i m p r e s s i o n between eyes weaker: pronotum more t ransverse . Holo type . 6*, label led as fo l lows : " N . Guinea: N E Bulo lo , 700 m 26. X I . 1969"; " J . et

M . Sedlacek Co l l ec to r s BISHOP M U S E U M " . I t is deposited i n B P B M .

F igs 5-12. 5= Las t s ix antennái segments pf L a g r i a l igu la t a sp. n . , S. - 6= Aedeagus of L . | l igu la ta sp. n . - 7 = Aedeagus of L . p u p i l l i c o l l i s Borchmann . - 8= Aedeagus of L . amethyst ina s p . n . - 9= Aedeagus of L , sapphir ina sp. n . ! - 10= Aedeagus of Bo th r i cha ra i r idescens s p . n [• 11= Aedeagus of B . ine r s sp. n . - 12= Aedeagus of B . i n t r i c a t a sp. n.

Paratypes (total 35 specimens). Morobe P rov ince : label led as holotype (2 <j><j>, B P B M , 1 o, H N H M ex B P B M ) ; i d . , JS ( 3 o o , B P B M ; 1 H N H M ex B P B M ) ; i d . , 6. X I . 1969, JS (3 oo, B P B M ; 2 H N H M ex B P B M ) ; k i d . , 690 m , 27. I I I . 1968, P. Colman (2 çç, B P B M ) i d . , 23. I V . 1969, JS (1 o. B P B M ) ; i d . , 4. V I . 1969, JS (1 ff, B P B M ) ; Bu lo lo Gorge , M c A d a m Park , 900 m , 28. X I I . 1977, u . v . l i g h t , R. Hamsen, P. Heber t et J. Ha rmsen (1 S, CNCI ) ; Bulo lo R i v . , 680 m , 23. I V . 1969, JS (1 SPCB); F i n s c h Haven d i s t r . , L . Wagner (2 çç.

SAMA) ; Wau, 1050- 1100 m , 9. D i - 1 1 . X I . 1961, JS, JHS (2 88, B P B M ; 2 So*. 3 gg,- SPCB; 1 g, H N H M ex SPCB); Wau, K i l o l o Creek, 26. V I I I . 1968 [ N G W - C 1 4 . ] , J . Balogh ( l g , HNHM) :

Wau, Mt . M i s s i m , 950- 1300 m , 8-9.1.1966, JS, MS (1 6, B P B M ) ; Zenag-Lae Road. 200 m , 17.1. 1965, JS (1 g, B P B M ) . - Madang Prov ince : Manumbo (1 g, SAMA) . - East Sepik P r o ­v ince : Hauptlager b. M a l u , 31.1.1913, Ka i se r in -Augus ta f luss Expedi t ion , S.G. Bürgers ( l g , Z M H U ) ; Lager a. Topfe r f lu s s , 1 8 . I V . 1913, Ka i se r in -Augus ta f lu s s Expedi t ion , S .G.Bürgers (1 g, Z M H U ) ; M a p r i k , Sepik, 16.11.1965, R. Hornabrook (1 g, HPCW).

R e m a r k s : L a g r i a amethyst ina is s i m i l a r to L . sapphi r ina but the e l y t r a are more broadly oval (s lenderer in sapphi r ina) , always p u r p l i s h v io l e t (bluish or greenish in sapphi­ r i na ) and pronotum is never d a r k l i n g (often so i n sapphi r ina) .

L a g r i a sapphir ina sp. n . (Figs 4, 9)

Head, p rono tum, s c u t e l l u m , legs and antennae pale reddish brown; p rono tum most of­ten (and head somet imes) da rke r , w i t h greenish t inge; last antennái segment somewhat dark­e r ; e l y t r a greenish blue (reddish brown in i m m a t u r e specimens) ; v e n t r a l surface da rke r r e d ­dish b rown. Length : 7 .2 -9 .0 m m .

8. Head impressed between eyes; w i th 2 blunt protuberances behind i m p r e s s i o n ; p ro tuberances and i m p r e s s i o n near ly smooth; o the rwise , head sparse ly punctate; w i t h some long , e rec t h a i r s ; EI= 13: 8; length ra t ios of antennái segments: 9: 4: 10: 10: 6: 8: 8: 7: 7: 6: 30; no t r ansve r sa l or modif ied segments; segment I s h o r t e r than half of distance between anten­nái sockets. P rono tum near ly as long as broad or s l i gh t l y t ransverse ; widest j u s t an t e r io r to the middle ; sides subpara l l e l to weakly arched; punctation coarse and dense, somewhat rugose p o s t e r i o r l y ; surface w i t h 2 or 4 weak i m p r e s s i o n s ; pubescence ra the r long , sparse , e rec t . E l y t r a r a t h e r elongate ova l , moderate ly convex; sides near ly s t ra igh t f r o m humerus to broadest point; punctation coarse , dense; interspaces a l i t t l e p l icate ; pubescence r a the r shor t , dense, e rec t to suberect . Sterna without modi f ica t ions . Legs typ ica l , w i t h sparse, r a t h e r shor t pubescence; t ibiae unarmed. Aedeagus: F i g . 9. Habi tus: F i g . 4.

g. L a r g e r on the average; EI= 14: 12; length ra t ios of antennái segments V I I I to X I : 8 : 8 : 7 : 2 2 ; i m p r e s s i o n between eyes weaker , but protuberances somewhat more p rominen t ; pronotum d i s t i nc t l y t r ansverse .

Holotype, 8, label led as fo l lows : "NEW G U I N E A : Wau, Bishop Museum, F i e l d Station, 15-25. I V . 1965"; " C o l l . D r . J . B A L O G H et D r . J . J . S Z E N T - I V A N Y " . I t i s deposited in H N H M .

Paratypes (total 60 specimens) . Morobe P rov ince : B u l o l o , 26. X I . 1969, JS, MS (1 g, B P B M ) ; Bu lo lo R i v . , 1180 m , 1 7 . I X . 1969, A . B . M i r z a (1 g, B P B M ; 1 g, H N H M ex B P B M ) ; Gara ina , 20. X I - 1 7 . X I I . 1969, A . B . M i r z a (1 g, B P B M ) ; i d . , 28. V I I . 1971, R. Hornabrook (1 8, 1 g, HPCW; 1 g, H N H M ex HPCW); U m g . Gurako r , 1 6 . D i . 1979, W . G . U l l r i c h (1 g, MHNG); M u m i n g , 600 m , 9-10. I I I . 1962, JS (1 g, B P B M ) ; Sat telberg, v . Bennigsen (1 8, I F P E ) ; Simbang, Huon Golf , 8. I I . 1899, L . Biró (1 g, H N H M ) ; Wareo, L . Wagner (1 g, S A M A ) ; Wau, 1050-1300 m , 20. V I I I . 1961-XI . 1978, Deharveng, R. Hornabrook , H . F . How­den, R . E . P a r r o t t , JS, MS, R. Straatman,; (4 88, 6 gg, B P B M ; 3 gg, CNCI ; 1 g HPCW; 1 MHNG; 2 88, 2 gg, SPCB; 1 8, H N H M ex B P B M ; 3 gg, H N H M . 1 each ex C N C I , HPCW, SPCB); Wau, 8-14. X I I . 1976, m e r c u r y vapor l i g h t , G . F . H e v e l e t R . E . Dietz (1 8, 7 gg, USNM; 1 8, 1 g, H N H M ex USNM); Umg. Wau, X I I . 1978, W . G . U l l r i c h (1 g, MHNG); Wau, B i g Wau Creek, 1300 m , X I . 1965, P. Shanahan (1 8, B P B M ) ; Wau, Hospi ta l Creek , 1200 m , 11. 1965, JS, MS (1 g, . B P B M ) ; Wau, M t . M i s s i m , 980 m , 20. V I I . 1969, J . L . G r e s s i t t et V. H i r a s h i m a (1 o, B P B M ) . - Madang Prov ince : F i n i s t e r r e Range, Saidor, Funyende, 1200 m , 24-30. I X . 1958, W . W . Brandt (1 g, B P B M ) ; U m g . Madang, 29. V I . 1979, W . G . U l l r i c h (1 g, MHNG). - Wes te rn Highlands Prov ince : B a i y e r R i v e r , Sanctuary, 1 -5 . IX. 1969 [ N G B - U . 2 3 . ] , J . Balogh (1 g, H N H M ) ; M t . Hagen iVal ley , K e l t i g a , 5600 f t . , 28. I X - 2 5 . X . 1961, W . W . B r a n d t (1 8, A N I C ) . - West Sepik P rov ince : T o r r i c e l l i M t s , Mohai V i l i . , 750 m , 8-15. X I I . 1958, W . W . Brandt (1 6, B P B M ) . - Western P roy ince : D a r u , mouth of F l y R i v e r , V I I I . 1941, R. G; Wind (1 g, CASC); Moorhead, 18 m , 14. V I I . 1964, H . C l i s s o l d (1 g, B P B M ) ; Morehead , 10. X I . 1974, R. Hornabrook (1 8, HPCW). - N o r t h e r n P rov ince : Mt . Lamington , 1300-1500 f t . , i C . T . McNamara (1 g, S A M A ) ; Dobodura, I I I - V I I . 1944, Da r l i ng ton (1 g, M C Z C ) .

R e m a r k s : L a g r i a sapphir ina is c losely a l l i e d to L . amethyst ina but e l y t r a are more elongate (more ovoid in amethyst ina) , g reen i sh -b lu i sh (purp l i sh v io le t i n amethyst ina ), the pronotum and last antenna! segment most ly d a r k l i n g (reddish in amethyst ina) .

B o t h r i c h a r a i r idescens sp. n . (Figs 10, 13)

Head, pronotum and scu te l lum black, densely covered by fine decumbent h a i r s ; e l y t r a l a rge ly p u r p l i s h - b l u i s h , i r idescen t , at places wi th greenish l u s t r e ; pubescence decumbent, incomple te ; pubescent places a l te rna t ing w i t h extensive glabrous par t s ; v e n t r a l surface black; legs dark blackish b rown; antennae reddish brown, a l i t t l e d a r k l i n g toward apex. Pub­escence s i l v e r y whi te . Length: 7. 5-7. 6 m m .

6. Head finely and densely punctate; eyes ra ther convex, t r ansve r se ; EI= 13:8; length r a t ios of antennái segments: 8: 5: 13: 15: 13: 12: 12: 12: 11 : 11: 22; no modif ied or t r a n s v e r s a l segments; segment I about as long as half the distance between antennái sockets. Pronotum very densely and finely punctate; punctuation i nv i s ib l e because of dense pubescence; of near­ly same length and wid th ; widest at base, s l i gh t l y cons t r ic ted ju s t pos t e r io r to the middle-

disc w i th flat impres s ions . E l y t r a elongate, moderate ly convex, s l i gh t l y widening in s t ra igh t l ine , widest in apical 1/4; punctures-coarse, more or less evenly scat tered, w e l l separated, interspaces a lmost f la t , not p l ica te ; basal 1/6 densely pubescent, then fol lowed by a t r ans ­verse glabrous s t r i p of same wid th ; r e s t of e l y t r a wi th l a rge , pa r t ly confluent, round,

F ig s 13-14. 13= B o t h r i c h a r a i r idescens sp. n . , - 14= B . iners sp. n . , <?.

glabrous spots surrounded by pubescent areas; glabrous spots somewhat depressed. Legs ra ther long and slender; f emora wi th scarce , decumbent pubescence; t ibiae wi th pubescence also decumbent, inner side w i t h shor t , s emie rec t h a i r s as w e l l ; fore t ibiae near ly s t r a igh t , unarmed; middle and hind t ib iae s l igh t ly but d i s t i nc t l y arched, inner ma rg in w i t h few s m a l l denticles placed far f r o m each other . Sterna without modi f ica t ions : last abdominal s te rn i te broadly rounded. Aedeagus: F i g . 10. Habi tus : F i g . 13.

o" Unknown. Holotype, 6\ l abe l led as fo l lows : "NEW GUINEA (NE) B a i y e r R i v e r Sanctuary 1-5. D i .

1969."; " (No. N G B - U . 22. ) l eg . D r . J . Balogh" . I t is deposited i n H N H M . Paratype. Western Highlands P rov ince : J i m i V a l l e y , Tabibuga, 23-24. I V . 1970, R.

Hornabrook (1 Í . HPCW). R e m a r k s : B o t h r i c h a r a i r idescens is somewhat s i m i l a r to B . pulchel la (Guérin-Méne­

v i l l e , 1830) but d i s t inc t ly d i f fers f rom i t in the f o l l o w i n g :

ground colour of e l y t r a

t r a n s v e r s a l glabrous s t r i p behind pubescent e l y t r a l base

glabrous spots on e l y t r a

apex of aedeagus

i r idescens sp. n .

b lu i sh - p u r p l i s h , i r idescen t , sh ining

ho r i zon ta l

d i s t i nc t , w e l l depressed

an acute process

pulchel la (Guérin-Méne­v i l l e )

b lu i sh , not i r idescen t , alutaceous

oblique

ind i s t inc t , poor ly depressed

t r i a n g u l a r

B o t h r i c h a r a iners sp. n . (Figs 11 , 14)

Body b lackish b rown; e l y t r a somewhat redd i sh ; su tu ra i ma rg in black. D o r s a l and ven­t r a l surface covered wi th shor t , decumbent, g r ey i sh white pubescence. Length: 9. 1 m m .

<?. Head finely and sparse ly punctate, sparse ly pubescent; eyes ra ther convex, t r ans ­verse ; EI= 13. 5: 10; antennae ra ther stouth; segment length r a t i o s : 9 :4 . 5: 13: 14: 13: 12: 11: 1 0 : 1 1 : 1 1 : 2 1 ; no modif ied o r t r a n s v e r s a l segments; segment I sho r t e r than half the distance between antennái sockets. Pronotum a l i t t l e broader than long; widest jus t an te r io r to the middle , r a the r cons t r i c ted an t e r io r to base; disc w i t h 4 d i s t inc t but flat impress ions ; su r -facely densely and modera te ly coarsely punctate; pubescence r a the r sparse, punctation w e l l v i s i b l e among h a i r s . E l y t r a modera te ly convex, elongate, widest in pos te r io r 1/3: sides arcuate; surface wi th coarse punctures; interspaces not p l i ca te , as broad as or w ide r than puncture d iamete r ; pubescence moderately dense, leaving a few s m a l l spots g labrous; these spots v e r y weakly impres sed . Legs ra ther long and slender, sca rce ly pubescent; t ibiae near­ly s t r a igh t , unarmed. Sterna without modi f ica t ions ; las t abdominal s t e rn i t a rounded. Aedea­gus: F i g . 11 . Habi tus: F i g . 14.

g. Unknown. , Holotype , S, l abe l led as fo l lows : "NEW GUINEA (NE) Wau, M t . K a i n d i , 19-24. V I I I .

1969"; " (No . N G W - U . 12.) l eg . D r . J . Balogh. I t i s deposited i n H N H M . R e m a r k s : Th i s species is somewhat s i m i l a r to B o t h r i c h a r a h e l l e r i Borchmann, 1915,

but the e l y t r a are less elongate, the e l y t r a l punctat ion is coarse r , the antennae are shor t e r and t h i c k e r .

B o t h r i c h a r a i n t r i c a t a sp. n . (Figs 12, 15)

Head, pronotum and scu te l lum black, densely covered by f ine, decumbent h a i r s ; e l y t r a dark redd ish brown wi th r e s t r i c t e d , decumbent pubescence; surface l a rge ly glabrous; ven­t r a l surface, legs and antennae dark reddish brown to black, basal par t of antennae some­what pa le r . Pubescence golden y e l l o w i s h . Length: 9 . 5 - 1 1 . 5 m m .

S. Head f inely and densely punctate; eyes r a the r convex, t r ansve r se ; EI= 12: 13; length ra t ios of antenna! segments: 10: 5: 15: 13: 13: 11: 11 : 11:12: 10: 2 1 ; no modif ied or t r a n s v e r s a l

S e g m e n t s ; segment I about as long as ha l f of distance between antennái sockets. P rono tum ve ry densely and f inely punctate, punctuation almost i n v i s i b l e because of dense pubescence; about as long as broad; widest at base but width measured an te r io r to cons t r i c t ion a lmost equal to basal wid th ; disc wi th 4 i nd i s t i nc t i m p r e s s i o n s . E l y t r a s t rong ly convex and broad, widest in apica l 1/3; surface e x t r e m e l y roughly scu lp tu red ; w i th coarse , sparse punctures and h ighly elevated, i r r e g u l a r , pl icate in t e rva l s making the surface creased; w i t h some barely d i sce rn ib l e , shallow i m p r e s s i o n s ; pubescence sho r t , decumbent, inconspicuous, somewhat denser in basal 1/6 and along su tu ra i m a r g i n , elsewhere sparse r ; impressed spots g labrous . Legs moderate ly long and s lender , w i t h scanty pubescence; fore t ibiae near­ly s t r a igh t , unarmed; middle and hind t ib iae very weakly arched, w i t h a row of s m a l l den­t i c les along inner m a r g i n . Sterna wi thout modi f ica t ions ; last abdominal s te rn i te broadly rounded. Aedeagus: F i g . 12. Habi tus : F i g . 15.

F igs 15-16. 15= B o t h r i c h a r a i n t r i c a t a sp. n . , Ô*. - 16= B . wau sp. n . , <?

p. L a r g e r and broader on the average; eyes s m a l l e r ; EI= 14: 18; length r a t i o s of an­tennái" segments V I I I to X I : 12 :12 :11 :22 ; p rono tum d i s t i nc t l y t r ansverse ; a l l t i b i ae unarmed; e l y t r a s l i gh t ly caudate.

Holotype , 8, l abe l led as fo l lows : " N E W G U I N E A : NE Wau, Morobe D i s t r . 1100 m, 31 . I . 1963"; " J . Sedlacek Co l l ec to r BISHOP". I t is deposited in B P B M .

Paratypes (total 31 specimens) . Morobe P rov ince : Gara ina , 1200 m , 15 .1 . 1968, JS, MS (1 g, B P B M ) ; M t . K a i n d i , 2350 m , 1. I V . 1966, J . L . et M . G r e s s i t t (1 g, B P B M ) ; i d . , 19-24. V I I . 1969, J . L . Gres s i t t (1 g, B P B M ) ; Wau, 1100- 1800 m , P. Colman, R. Hornabrook, R . E . P a r r o t t , JS, MS, P. Shanahan (2 83, 6 gg, B P B M ; 1 g, C N C I ; 1 g, HPCW; 3 gg, SPCB; 1 8, H N H M ) ; Wau, Edie Creek , 2050-2300 m , 17-18. V I I I . 1965, JS (1 g, B P B M ) ; i d . , 2000-2400 m , 12. V I I I . 1963, P. Shanahan. (1 g, B P B M ) ; i d . , 7000 f t . , 2 1 . I X . 1964, M . E . Bacchus (1 g, B M N H ) ; Wau, Haus Copper, M t . M i s s i m , 22-24. I V . 1965, J. Balogh et J . J . Szent-Ivány (1 o, H N H M ) ; M t . M i s s i m , 1600-2000 m , 21-24. I X . 1965, MS (1 o, B P B M ) ; i d . , 1500-1800 m , 6.1.1970, JS, MS (1 g, B P B M ) ; Wau, K u n a i C k . , 1580 m , 22. V . 1965, JS ( l g . S P C B ) ; Wau, K u j e r u , 1500 m , 27. I X . 1969, A . B . M i r z a (1 8, 1 g, B P B M ; 1 6, H N H M ex B P B M ) ; Wau, Wau Creek , 1200- 1500 m , 16-18. I X . 1964, MS (1 8, 1 g, B P B M ; 1 g, H N H M ex BPBM); Watut R. , 900- 1900 m , X . 1969, A . B . M i r z a (1 8, B P B M ) .

R e m a r k s : Here tofore , i t is the only descr ibed species of Bo th r i cha ra having very rough­ly p l ica ted e l y t r a .

B o t h r i c h a r a wau sp. n. (Figs 16-18)

Body black; head, p rono tum, s cu t e l l um and v e n t r a l surface densely covered wi th ve ry shor t , fine decumbent h a i r s ; e l y t r a l ve s t i t u r e uneven: densely pubescent spots a l te rna t ing w i t h places of sparser pubescence. Pubescence s l i v e r y whi te . Length: 9. 1-12. 5 m m .

3. Head f inely and densely punctate; eyes moderate ly convex, t r ansve r se , E I : 16: 11; length ra t ios of antennái segments: 11:6: 15: 15:12: 12: 11: 11: 10: 9: 2 1 ; no modif ied or t rans­v e r s a l segments; segment I about as long as half the distance between antennái sockets. P rono tum densely and finely punctate; punctuation w e l l v i s i b l e , not en t i re ly covered by fine pubescence; about as long as broad; widest jus t a n t e r i o r to the midd le , but base near ly as broad; w i t h 4 ind i s t inc t impre s s ions . E l y t r a modera te ly vconvex, elongate, wides t i n ante­r i o r 1/4; sides widening i n a s t ra igh t l ine to widest point ; punctation coarse and moderately dense; at places, interspaces somewhat broader and s l i gh t l y ra i sed ; pat tern of pubescence c h a r a c t e r i s t i c : denser pubescence cove r ing basal 1/6 (this surface obliquely bordered pos­t e r i o r l y ) , f o rming a p reap ica l spot (reaching side of e l y t r a but not reaching su tu ra i marg in and apex); r emain ing par t of e ly t r a wi th a number of s m a l l i r r e g u l a r spots a r ranged in loose longi tudina l rows ; space amongst densely pubescent spots w i th much sparser v e s t i t u r e ; no def in i te ly glabrous and impressed places. Legs long and s lender , wi th sparse , decumbent pubescence; fore and middle t ibiae nea r ly s t r a igh t , s imp le ; hind t ibiae ve ry weakly arched, w i th a weak but d i s t inc t preapica l swe l l i ng (F ig . 17). Sterna without modi f i ca t ion ; last ab­dominal s te rn i te rounded. Aedeagus: F i g . 18. Habi tus : F i g . 16.

g. Much l a r g e r on the average; eyes s m a l l e r ; EI= 15:20; length ra t ios of antennái seg­ments V I I I to X I : 12 :11 :10 :20 ; pronotum d i s t i n c t l y t r ansverse , impres sed mesa l ly ; e l y t r a l ve s t i t u r e usual ly much more conspicuous; hind t ib iae s i m p l e .

Holotype , 3, l abe l led as fo l lows : "NEW GUINEA (NE) Wau, Morobe D i s t r . 1200 m , 5. V . 1963"; " J . Sedlacek BISHOP". I t is deposited in B P B M .

Paratypes (total 81 specimens) . Morobe P rov ince : K a i n d i - N a m i , 1700 m , 15. V I . 1965, JS (1 g, SPCB); Wau, 1050- 1800 m , 1. I I . 1961-22. I I I . 1969, G. Monte i th . JS, JHS, MS (12 88, 18 gg, B P B M ; 1 8, 4 gg, SPCB; 3 88, 6 gg, H N H M ex B P B M ) ; Wau, 1200 m , 8-14. X I I . 1976, m e r c u r y vapor l igh t , G. F . Hevel et H . E. Die tz (1 g, USNM); Wau, B i g Wau Ck. , 1200-1500 m , I X . 1965, JS, MS (1 o, B P B M ) ; Wau, Haus Copper , M t . M i s s i m , 22-24. I V . 1965, J . Ba ­logh et J . J . Szent-Ivány (2 gg, H N H M ) ; Wau, M t . M i s s i m , 1500-1800 m . 6.1.1970, JS, MS (1 3, 1 g, B P B M ) ; i d . , 1600-2000 m , 21-24. I X . 1964, MS (2 gg, B P B M ) ; W a u . Ku je ru , 1500 m , 27. V . 1969, A . B . M i r z a (1 8, B P B M ) ; Wau, M c A d a m Park , 1200- 1300 m , 7 . V . 1965, JS (1 g, B P B M ) ; Wau, M t . K a i n d i , 1900-2200 m . 19. V I I I . 1965, JS, MS (1 o. B P B M ) ; Wau, Wau. Ck. , 18. IX. 1964, JS (1 g, SPCB); i d . , 1200- 1500 m , 28. I I I . 1963, JS (1 8, B P B M ) ; i d . , 1200-1500 m , 16-18. I X . 1964, MS (1 g, B P B M ) ; 6 k m W of Wau, N a m i Creek, 1700 m , 15. VI.1962 (1 3, 2 gg, B P B M ; 1 g, H N H M ex B P B M ) . - Madang P rov ince : F i n i s t e r r e M t s . , Budenu, cca

4000 f t . , 15-24. X . 1964, M . E. Bacchus (2 88, 2 go, B M N H ) ; F i n i s t e r r e M t s . , M o r o , cca 5550 f t . , 3 0 . X . - 1 5 . X I . 1964, M . E . Bacchus (1 g, B M N H ) . - Eas te rn Highlands P rov ince : Asakamotasa , 6000 f t . , 27.11.1964, R. Hornabrook (2 go, HPCW; 1 g, H N H M ex HPCW); Umg. Kainantu , Onerunka, I V . 1979, W . G . U l l r i c h (1 g, MHNG); i d . , 5 . X . 1979 (1 g, MHNG), i d . , 13 .X . 1979 (1 g, H N H M ex MHNG); Okapa, 23. V . 1965, R. Hornabrook (1 g, H P C W ) ; i d . , cca 5000 f t . , 4-15. I I . 1965, M . E. Bacchus (2 gg, B M N H ) ; Purosa Camp, Okapa area , 15 mis SSE Okapa P a t r o l Post , 1950 m . Camp 10, 2 1 . I X . 1959, Sixth A r c h b o l d Expedi t ion , | L . J . B r a s s (1 g, A M N H ) ; Waisa, N Okapa, c. 5000 f t . , 15.11. 1965, M . E . Bacchus (1 g, B M N H ) . - Western Highlands Prov ince : B a i y e r - R o h i n a , I V . 1979, W . G . U l l r i c h (1 g, MHNG); Mt. Hägen Va l l ey , K e l t i g a , 5600 f t . , 28. I X . - 25. X . 1961, W . W . Brandt (1 g, A N I C ) .

17 18 19 20 21 22

Figs 17-22. 17= Hind t i b i a of Bo th r i cha ra wau sp. n . , S. - 18= Aedeagus of B . wau sp. n . -19-20= Aedeagus of T o m o g r i a per la ta gen. et sp. n . , v e n t r a l view (19) and l a t e r a l view (20). r

21-22= Aedeagus of A c u t o l a g r i a falcata gen, et sp. n . , v e n t r a l view (21) and l a t e r a l v iew (22)'

R e m a r k s : The new species is somewhat s im i l a r to B o t h r i c h a r a kuntzeni Borchmann , 1936 on account of i t s e l y t r a l pubescent spots ar ranged in r o w s , but a large preapical spot is present ( lacking in kuntzeni ) , the e l y t r a l punctation is coarse (fine in kuntzeni) , the i n t e r spaces s l i g h t l y pl icate (not so in kuntzeni) and ftje Hind t ib iae of males are swollen before apex (simple i n kuntzeni) .

T o m o g r i a gen. n.

Diagnos is : Bather b road and convex,- med ium-s i zed ; colour dark; pubescence shor t , semierec t , inconspicuous; head, pronutum and e l y t r a w i t h d i s t inc t punctation.

S. Head wi th i n t e r o c u l a r distance much n a r r o w e r than eye d iameter ; antennae wi thout modified o r t r ansve r sa l segments; segmenf: X I longer than 4 preceding segments combined s imple . P rono tum of same length and wid th , w i th d i s t inc t impre s s ions . E l y t r a ra ther broad, punctural interspaces f o r m i n g i r r e g u l a r , smooth, blunt protuberances; apex s imply round­ed epipleura reaching apex. Legs t yp i ca l ; I middle and hind t ib iae f inely s e r r a t e along inner edge. Sterna s i m p l e ; last abdominal segment s l igh t ly t runca te .

Ç. Much l a r g e r on the average; antennái segment X I shor te r than 3 preceding combin ­ed; eyes s m a l l e r , i n t e rocu la r distance broader ; p rono tum t r ansve r se ; a l l t ib iae unarmed .

Type species: T o m o g r i a pe r la ta sp. n . , by present designation. Remarks : On account of the p e a r l - l i k e tuberc les of the e l y t r a , this genus sharp ly dif­

fers f rom L a g r i a or any other New Guineán genera of L a g r i i n a . The pol ished, rounded pro­tuberances are much more evident when seen by naked eye.

T o m o g r i a per la ta sp. n. (Figs 19, 20, 23)

Head, p rono tum, s cu t e l l um, v e n t r a l surface and legs dark b lackish brown w i t h green­i sh -b ronzy t inge ; e l y t r a dark n u t - b r o w n tinged w i t h bronze; su tu ra i marg in black; antennae da rk brown. Leng th : 7. 1-10. 5 m m .

F igs 23-24. 23= T o m o g r i a pe r l a t a gen. et s p . n . , S*. - 24= A c u t o g r i a falcata gen. et sp .n . , 6*

S. Head coarse ly and densely punctate; w i t h sparse, moderate ly long h a i r s ; EI= 15:6; length ra t ios of antennái segments: 8: 5: 10: 9: 7: 8: 8: 8: 7. 5: 7: 34. P rono tum as long as broad o r ve ry weakly elongate; wides t at base, weakly n a r r o w i n g a n t e r i o r l y ; w i th 2 d i s t inc t , t r ans ­v e r s a l impress ions before base and behind a n t e r i o r m a r g i n ; ouflctation coarse and sparse,

disc w i t h smooth areas at the midd le ; pubescence modera te ly long, sparse , e rec t . E l y t r a wides t in apical 1/4; punctation coarse , unevenly sca t tered; interspaces f o r m i n g pol ished, s t rong ly r a i sed , rounded protuberances s t rewn i r r e g u l a r l y ; pubescence r a the r reduced, cons is t ing of fine s emie rec t to e rec t , scat tered ha i r s . Legs t y p i c a l ; t ibiae s t ra igh t ; middle and hind tibiae wi th a row of s m a l l denticles in apical half . Aedeagus: F igs 19-20. Habi tus: F i g . 23.

g. D i f f e r i ng as stated in the generic descr ip t ion ; EI= 15: 12; length r a t ios of antenna! segments V I I I to X I : 10: 10: 9: 23: 5.

Holotype, 8, l abe l led as fo l lows : "NEW GUINEA: (NE) Wau, Morobe D i s t r . Wau Ck. 1200- 1500 m , 28. I I I . 1963"; " J . Sedlacek Col lec to r BISHOP". I t is deposited in B P B M .

Paratypes (total 101 specimens) . Morobe P rov ince : labe l led as holotype (2 SS, 6 oo, B P B M ; 1 S, 1 a, H N H M ) ; i d . , 3 . I V . 1963, JS (2 gg, B P B M ) ; i d . , 16-18. I X . 1964, MS (To 1 , 2 gg, B P B M ; I S , 1 g, H N H M ex B P B M ) ; Bulo lo , 700 m , 26. X I . 1969, JS, MS (1 S, B P B M ) ; . B u l o l o R i v . , 1180 m , 17. D i . 1969, A . B . M i r z a (2 SS, B P B M ) ; Gara ina , R. Hornabrook (l8, H P C W ) ; U . Watut , SW, 1300-1600 m . I . V . 1963, J . L . G r e s s i t t (1 g, B P B M ) ; Watut R. , X ; 1969, 900-1900 m , A . B . M i r z a (1 g, B P B M ) ; Wau, 1150- 1700 m , 17. V I . 1961-8. V I I I . 1968, J . L . Gre s s i t t , H . F . Howdeh, P. Shanahan, JS, MS (5 SS, 15 oo, B P B M ; 1 S, CNCI ; 1 g, SPCB; 2 SS, 5 gg, H N H M ex B P B M ) ; Wau, B i g Wau Ck . , 1200 m , X I I . 1965, P. Shanahan (1 g, B P B M ) ; i d . , 1200-1500 m , D i . 1965, JS, MS (2 gg, B P B M ) ; Wau, Hosp i t a l Ck. , 1150-1250 m , 9.1.1966, JS, MS (1 g, B P B M ) ; Wau, K u j e r u , 1500 m , 27. D i . 1969, A . B . M i r z a (1 8, 7 og, B P B M ; 1 8, 2 gg, H N H M ex B P B M ) ; Wau, M c A d a m Park , 1200-1300 m . 7. V . 1965, JS (1 g, B P B M ) ; Wau, M t . K a i n d i , • 1500-2200 m , 19. V I I I . 1965. JS, MS (1 g, B P B M ) ; i d . , 1700, 8. V I I I . 1968. JS, MS (1 g, B P B M ) ; i d . , 2000 m , 27. V I I I . 1968 [ N G W - R . l l j , I . Loksa (1 o. H N H M ) ; Wau, M t . K a i n d i , N a m i Creek, 22. V I I I . 1968 [ N G W - C . 4.J , J . Balogh (1 g. H N H M ) ; K a i n d i - N a m i , 22. V I I I . 1968, JS (4 SS, 2 gg, B P B M ; 2 SS, H N H M ex B P B M ) ; Wau, N a m i Creek, 1600-1650 m , 24.11.1963, JS (1 g, B P B M ) ; i d . , 1700 m , 12. V I . 1962, JS (2 SS, 1 g, B P B M ) ; i d . , 1700 m , 15. V I . 1962, JS (2 SS, 2 gg, B P B M ; 1 g, H N H M ex B P B M ) ; i d . , 1700 m , 5. I I . 1963. JS (1 S, 1 g, SPCB); i d . , 1700 m , 22. V I I I . 1963, JS (1 g, B P B M ) ; - Wau, M t . M i s s i m , 22-24. I V . 1965, J . . Balogh et J . J . Szent-Ivány (1 S, 4 g g , H N H M ) ; i d . , 1500-1800 m , 6. 1.1970. JS, MS (2 gg, B P B M ) ; i d . , 1700 m , 7 . I I I . 1963, JS (1 g, B P B M ) ; 24-32 k m SE of Wau, 1500-1900 m , 20. I I I . 1962, JS (1 g, B P B M ) .

A c u t o g r i a gen. n .

Diagnosis : Elongate, modera te ly convex, med ium-s i zed ; co lour da rk ; pubescence s h o r t , semie rec t , sparse; head, p rono tum and e ly t r a w i t h d i s t inc t punctat ion.

8, Head wi th i n t e r o c u l a r distance n a r r o w e r than eye d iameter ; antennái segments I I to X s i m p l e , not t r ansve r se ; segment X I as long as 5 preceeding segments combined, curved , ab rup t ly attenuate in apical 1/3, apex acute. Pronotum s l igh t ly longer than broad, w i th v e r y shal low impress ions . E l y t r a modera te ly widening p o s t e r i o r l y ; surface roughly sculp tured; apex s imp le ; inner m a r g i n of epip leura reaching apex. Legs ra ther s t rong , moderately long; fore femora d i s t i nc t ly longer and s touter than middle and hind ones; fore t ibiae curved and twis t ed i n apica l 1/3, inner m a r g i n w i t h a l a rge , broad, t oo th - l i ke widening jus t before the midd l e ; inner m a r g i n of middle and hind t ibiae finely s e r r a t e subapical ly . Sterna s imp le ; last abdominal s te rn i te rounded at apex.

g. L a r g e r on the average; .antennái segment X I s i m p l e , a l i t t l e longer than 2 preceed-i ing segments combined; i n t e r o c u l a r distance broader; p ronotum t r ansve r se ; fore femora not d i s t i n c t l y longer and s touter than o thers ; fore t ib iae not modi f ied .

Type species: A c u t o g r i a falcata sp. n . , by present designation. R e m a r k s : The most sa l ient cha rac t e r i s t i c s of this genus are the b i l l - shaped , acute

last antennái segment and the uniquely fo rmed fore t ib iae of the males .

A c u t o g r i a falcata sp.; n . (Figs 2 1 , 22, 24)

Head, pronotum and scu te l lum dark b lackish green w i t h me ta l l i c sheen; e l y t r a shiny, da rk b lack ish brown; a l a r g e , common spot jus t behind middle of e l y t r a d a r k e r black (more evident on i m m a t u r e specimens) ; v e n t r a l surface, legs and antennae dark redd ish brown. Length : 8. 0-10. 5 m m .

S. Head coarse ly and densely punctate; w i th sparse, modera te ly long h a i r s ; eyes mod­era te ly convex; EI= 14: 6; antennái segment length r a t i o s : 11 : 5: 10: 8: 7: 7: 7: 7: 7: 6: 34; seg­ment X I , in p r o x i m a l 2 / 3 , nea r ly s t r a igh t , ' evenly wide , subquadrate i n cross sec t ion , then curved and na r rowed , w i th pointed apex. Pronotum s l igh t ly longer than broad, widest just an t e r io r to the middle ; sides evenly arcuate; surface less convex, w i t h 4 v e r y shallow i m ­press ions; punctation ve ry dense and coarse , punctures subcontiguous or even coalescing to t r a n s v e r s a l w r i n k l e s ; pubescence consis t ing of longer , spa r se r , e rec t and sho r t e r , denser decumbent ha i r s . E l y t r a widest i n apical 1/3; punctation coarse , punctures unevenly clus­t e red , leaving i r r e g u l a r interspaces as r a i sed pl icae; pubescence ra ther shor t , sparse, semie rec t , mixed wi th severa l e rec t h a i r s . Legs rather s t rong ; fore femora conspicuously stout; fore t ibiae nea r ly s t r a igh t in p r o x i m a l 2 /3 , then curved and twis ted ; inner ma rg in w i t h a l a rge , a s y m m e t r i c a l , t o o t h - l i k e widening just pos t e r io r to the midd le ; edge of this widening f inely crenula te ; middle t ib iae s l i gh t l y curved , d i s t a l ha l f of inner m a r g i n finely se r ra te ; hind t ib iae nea r ly s t r a igh t , d i s t a l 2/5 of inner m a r g i n w i t h fine dent icula t ion; a l l legs wi th sparse, modera te ly long, s emie rec t h a i r s . Aedeagus: F igs 21-22. Habi tus : F ig .24 .

g. L a r g e r on the average; eyes s m a l l e r , EI= 14: 12; r e l a t i ve lengths of antennái seg­ments V I I I to X I : 9 : 9 : 8 : 2 1 ; segment X I v e r y weakly cu rved , f u s i f o r m , p rono tum s l igh t ly t r ansverse , w i t h a mesal spot much f inely punctate (often obsolete): legs wi thout any modi ­f icat ions , i

Holotype, S, l abe l led as fo l lows : "NEW G U I N E A ; Wau, M c A d a m park, 18 -21 . I V . 1965"; " C o l l . D r . J . B A L O G H et D r . J . J . S Z E N T - I V Á N Y " I t is deposited in H N H M .

Paratypes (total 142 specimens) . Morobe Prov ince : l abe l led as holotype (2 g g . H N H M ) ; i d . , 1200- 1250 m , 20. V I I . 1965, JS (1 g, B P B M ) ; Bu lo lo , 1700 m , 26. X I . 1969, JS, MS (2 6*3 1 Q, B P B M ) ; Gara ina , 20. X I - 1 7 . X I I . 1969, A . B . M i r z a (1 o, B P B M ) ; i d . , R. Hornabrook (1 S, HPCW); Komba, L . Wagner (1 S, 1 g, SAMA) ; Lae, 4-6 . I X . 1968 [ N G L - C . 1 6 ] , J . Ba­logh (1 g , H N H M ) ; Sat telberg (1 p , Z M H U ) ; Watut R. , 900- 1900 m , X . 1969, A . B . M i r z a (2 SS, 3 gg, B P B M ; 1 6, 1 g, H N H M ex B P B M ) ; Wau, 1050- 1800 m , 29. V I I I . 1961-9. 11.1968, G . A . Samuelson, JS, JHS, MS, P. Shanahan (29 SS, 25 gg, B P B M ; 1 g, SPCB; 7 SS, 7 gg, H N H M ex B P B M ) ; Wau, Bu lo lo R. , 850-950 m , 4. I I . 1966, JS (1 Ó, B P B M ) ; i d . , 1130 m , 17. I X . 1969, A . B . M i r z a (3 SS, 3 gg, B P B M ; " 1 S, H N H M ex B P B M ) ; Wau, Hosp i t a l Ck. , 1150- 1250 m , 9. I . 1966, JS, MS (2 SS, B P B M ; 1 S, H N H M ex B P B M ) ; i d . , 1200 m , I . 1965, JS (2 SS, B P B M ) ; Wau, K u j e r u , 1500 m , 27. I X , 1969, A . B . M i r z a (3 SS, 4 gg, B P B M ; 1 S,[ 2 gg, H N H M ex B P B M ) ; Wau, M t . M i s s i m , 950- 1300 m , 8-9.1 .1966, JS, MS (1 g, B P B M ) ; ' i d . , 1120 m , 13.11.1963, JS (1 g, B P B M ) ; i d . , 1500- 1900 m , 6.1.1970, JS, MS (1 g, BPBM); Wau, Wau Ck. , 1200- 1500 m , 28. I I I . 1963, JS (1 g, B P B M ) ; i d . , 1200-1500 m , 3. I V . 1963, JS (1 S, B P B M ) ; i d . , 1200- 1500 m , 16-18. DC. 1964, MS (1 S, 2 go, B P B M ; 1 g . H N H M ex B P B M ) ; 6 k m W of Wau, N a m i Creek , 1700 m , 12. V I . 1962, JS (1 S, B P B M ) . - Madang P r o ­v ince : F i n i s t e r r e Range, Saidor , K i a m b a v i v i l l . , 1-28. V I I I . 1958, W . W . Brand t (1 S, B PB M l -Western Highlands P r o v i n c e : J i m i V a l l e y , Tabibuga, 23-24. I V . 1970, R. Hornabrook (1 <?, HPCW). - Chimbu P rov ince : Chimbu Va l l ey , B i s m a r c k Rge, 5-7500 f t . , X . 1944, Dar l ing ton (1 S, M C Z C ) . - Eas te rn Highlands P rov ince : Asa roka , 20 k m W Goroka , 20. I I I . 1973, K . W . , Stroeder (1 S, CNCI) ; Goroka , 1550 m , 19. V I . 1955, J . L . G r e s s i t t (1 g, B P B M ) ; i d . , 26. V I . 1955, J . L . G r e s s i t t (2 SS, B P B M ) ; i d . , 5400 f t . , 16.1.1956, J . H . B a r r e t t (1 S, B M N H ) ; K a r i m u i , 1080 m , 13. V I I . 1963, JS (2 SS, B P B M ) ; Kassam, 1350 m , 48 k m E of Kainantu, 7. I X . 1959, T . C . Maa (1 S, B P B M ) ; Lapegu, 15. V I . 1977, ex Pinus Patula, H . Roberts (1 ő7, B M N H ) ; Okapa, 3500 f t . , 6. LX_1965, R. Hornabrook (1 S, HPCW); Okapa, Okasa, 1400-1600 m , 17 .1 . 1966, JS (1 S, B P B M ) ; Purosa , 1700 m , 17-25. V . 1966, Gres s i t t et T a w i (2 SS, B P B M ; 1 S, H N H M ex B P B M ) : i d . , 1700-2000 m , 18.1.1966, JS (1 g, B P B M ) . - N o r t h e r n P rov ince : Iongai , 1700- 1900 m . 9. X I . 1965, JS (1 S, B P B M ) ; Kokoda, 400 m , 14-16. X I . 1965, JS (1 S, B P B M ) ; i d . , 18. X I . 1966, JS (1 S, B P B M ) .

i .ündilagria gen. n .

Diagnosis : F a i r l y elongate, i v e r y weakly convex, med ium-s i zed ; colour b rownish ; dor­sal surface wi th duplex pubescence; head, p ronotum and e l y t r a w i th d i s t inc t punctation.

S. Head wi th i n t e r o c u l a r distance much n a r r o w e r than eye d iameter ; eyes convex; an­tennae long and slender; no modif ied or t r ansve r sa l segments; segment X I cu rved , c y l i n d r i ­ca l , somewhat longer than 2 preceding segments combined. P rono tum n a r r o w e r than head.

not t r ansverse . E l y t r a elongate, p r a c t i c a l l y not widening p o s t e r i o r l y ; surface w i t h d i spers ­ed, low protuberances; apex s imp le ; inner m a r g i n of epipleura reaching apex. Legs long and s lender; inner m a r g i n of t ibiae p r a c t i c a l l y una rmed . T h o r a c a l and v i s ib l e abdominal s terna without outgrowths ; last (= fifth) abdominal s t e rn i t e emarginate ap ica l ly ; s t e rn i t e V I s t rong ly modi f ied , f o r m i n g 2 l o b i f o r m and 2 forceps-shaped processes.

<j>. Eyes much s m a l l e r ; i n t e r o c u l a r distance broader ; antennae shor te r ; segment X I shor t e r than 2 preceding segments combined; p rono tum s l igh t ly t ransverse ; abdominal s terna V and V I not modi f i ed .

Type species: K a i n d i l a g r i a forc ipa ta sp. n . , by present designation. R e m a r k s : T h i s genus may eas i ly be d is t inguished f r o m a l l other genera by the pecul ia r

s t ruc tu re of abdominal s te rn i te V I , the s t rongly elongate body and the papil late surface of e l y t r a .

Ka ind i l ag r i a forc ipata sp. n . (Figs 25-28)

Head, p rono tum, scu te l lum and e l y t r a brown w i t h bronzy t inge; head and p rono ta l disc somewhat d a r k e r ; v e n t r a l surface paler reddish b rown; antennae dark b rown, apex and base of segments I to X paler ; legs pale reddish b r o w n , femora somewhat darker subapical-l y ; dorsa l surface w i t h denser, v e n t r a l surface w i t h sparse r pubescence. Length: 7.5-8.5 m m .

25 26 27

Figs 25-27. 25= A b d o m i n a l apex of K a i n d i l a g r i a fo rc ipa ta gen. et sp. n . , S, i n somewhat drawn-out condi t ion. - 26-27= Aedeagus of K . forc ipata gen. et sp. n . , v e n t r a l view (26) and l a t e r a l v iew (27)

6*. Head t r ansve r se , much broader than p rono tum, densely and coarse ly punctate; with, sparse , long h a i r s ; eyes v e r y la rge , p rominen t ly convex; EI= 15: 6; antennae as long as 2/3 of body length; segment length r a t i o s : 16:7; 11: 11: 10: 11: 11.5: 11.5: 12: 11: 28; segment I longer than ha l f the distance between antennái sockets . P rono tum about of same length and w i d t h , widest at base, sides converging in s t r a igh t l ine , weakly cons t r i c ted jus t pos te r io r to the middle ; surface uneven, highest c e n t r a l l y ; punctures coarse , w e l l separated; pubescence

c o n s i s t i n g of a few s e m i e r e c t a n d m a n y l o n g e r , e r e c t h a i r s . E l y t r a v e r y n a r r o w , s u b p a r a l -l e l - s i d e d , f la t ; w i t h c o a r s e p u n c t u r e s s e p a r a t e d by d i s t a n c e s about a s b r o a d a s p u n c t u r e d i a m e t e r ; i n t e r s p a c e s l a r g e l y f lat but s u r f a c e s e t w i t h s t r a g g l i n g , f la t , s m o o t h p r o t u b e r ­a n c e s ; c l o t h e d w i t h s c a t t e r e d , v e r y f ine , m o d e r a t e l y l o n g , s e m i e r e c t p u b e s c e n c e m i x e d w i t h a n u m b e r of m u c h l o n g e r , e r e c t h a i r s . L e g s long and s l e n d e r , wi th long h a i r s ; a l l t i ­b iae n e a r l y s t r a i g h t ; m i d d l e a n d h i n d t i b i a e w i t h i n d i s t i n c t d e n t i c u l a t i o n a long i n n e r edge . A b d o m i n a l s t e r n i t e V w i d e l y n o t c h e d ; apex of m o d i f i e d s t e r n i t e V I and a e d e a g u s u s u a l l y v i s i b l e i n v e n t r a l a s p e c t ( F i g . 2 5 ) . A e d e a g u s : F i g s 2 6 - 2 7 . H a b i t u s : F i g . 28 .

F i g s 2 8 - 2 9 . 28= K a i n d i l a g r i a f o r c i p a t a gen. e t s p . n . , Ô*. - 29= C a s n o n i d e a h y s t r i x s p . n . , ^

g. N o t l a r g e r on the a v e r a g e ; e y e s m u c h s m a l l e r ; E I = 13: 11; a n t e n n á i s e g m e n t s much-s h o r t e r a n d s t o u t e r ; s e g m e n t I not l o n g e r than h a l f the d i s t a n c e between a n t e n n á i s o c k e t s ; r e l a t i v e l e n g t h s of s e g m e n t s V I I I to X I : 9 : 9 : 8 : 13; p r o n o t u m b r o a d e r ; a l l t i b i a e t o t a l l y u n ­a r m e d ; a b d o m i n a l s t e r n i t e V b r o a d l y r o u n d e d at a p e x .

H o l o t y p e , 3, l a b e l l e d a s f o l l o w s : " N E W G U I N E A : N E M t . K a i n d i 2 0 0 0 - 2 3 5 0 m . 1 0 - 1 1 . I . 1965"; " J . et M . S e d l a c e k C o l l e c t o r s B I S H O P " . I t i s d e p o s i t e d in B P B M .

P a r a t y p e s (total 9 s p e c i m e n s ) . M o r o b e P r o v i n c e : L a e , 17. V I I . 1970, H , O h l m u s (1 Ô*, A N I C ) ; M t . K a i n d i . 2350 m , 23 . I I I . 1966 , J . L . G r e s s i t t (1 ç , B P B M ) ; i d . , I I . V . 1 9 6 8 . J S

(1 o, B P B M ) ; i d . , 19.1.1970, JS (1 o, SPCB); M t . K a i n d i near Wau, 7500 f t . , 25. V I . 1974, H . F . Howden (1 S, CNCI) ; Wau, M t . K a i n d i , 24-25. V I I I . 1968 [ N G W - C . 8 . ] , J . Balogh (1 g, H N H M ) ; Wau, 1200 m , 8-14. X I I . 1976, m e r c u r y vapor l i gh t , G. F . Hevei et R. E . D ie tz (2 gg USNM). - Southern Highlands P rov ince : M u l l e r Plateau, Atea-gana-anda, V I I - V I I I . 1978, G. Smi th et a l . (1 g, A N I C ) .

Casnonidea h y s t r i x sp. n . (Figs 29, 32)

Head, p ronotum and scu te l lum y e l l o w i s h r e d , somet imes w i t h olivaceous t inge; e l y t r a reddish brown wi th expressed steely blue to olivaceous t i n t ; p ro - and mesos te rnum y e l l o w ­ish r e d , metas te rnum and abdomen redd ish b rown ; antennae and legs ye l l owi sh r e d ; las t an­tenna! segment and femora a l i t t l e da rke r . Length : 7. 8-9 m m .

S. Head not broader than p rono tum; w i t h some set igerous punctures between eyes; ve r t ex ha rd ly swol len ; EI= 12: 13; antennae r a the r s tout , segment length r a t i o s : 9 : 4 : 9 : 9 : 8 : 9 : 9 : 9 : 9 : 8 : 2 5 . P rono tum s l i gh t l y t r ansve r se , wides t i n an te r io r 1/3, base hard ly na r rower ; ' surface g labrous , copiously punctate; punctures separated by distances at least equal to puncture d i ame te r . E l y t r a w i t h a l l punc tura l rows reaching apex; s t r i a i punctures moder­ately coarse , contiguous; in t e rva l s f la t ; a l l i n t e rva l s w i th numerous (over 30) se t igerous punctures sca t tered throughout whole length; these punctures not a r ranged in r egu la r r o w s ; setae w h i t i s h , moderate ly long, somewhat b r i s t l e - l i k e ; e l y t r a l apex s i m p l y rounded. Legs wi th ha i r s shor t , suberect , f iner than e l y t r a l setae. Last abdominal s te rn i te s i m p l y rounded. Aedeagus: F i g . 32. Habi tus : F i g . 29.

g. Ve r t ex a l i t t l e m o r e swol len , long i tud ina l f u r r o w deeper; EI= 12: 13; r e l a t i ve lengths: of antennái segments V I I I to X I : 9 : 9 : 9 : 1 8 .

Holo type , S, l abe l led as fol lows : "NE W GUINEA : Haus Copper, Wau, M t . M i s s i m , 22-2 4 . I V . 1965"; " C o l l . D r . J . B A L O G H et D r . J . J . S Z E N T - I V Á N Y " . I t i s deposited in H N H M .

Paratypes (total 34 specimens) . Morobe P rov ince : label led as holotype (2 SS, 4 gg, H N H M ) ; Wau, 1100- 1800 m , 7. V I I I . 1961-30. V I I I . 1977, R. Hornabrook, R. P a r r o t t , JS, JHS, MS (10 St, 5 go, B P B M ; 1 S, C N C I ; 10, 1 g, HPCW; 2 SS, 1 g, SPCB; 2 SS, H N H M ex B P B M ) ; Wau. 8-14. X I I . 1976, m e r c u r y vapour l i g h t , G . F . Hevel , e t R . E . Die tz (2 SS, USNM); Wau, Coviak Rdg. , 763 m , 7. X I . 1963, H . C . (1 g, B P B M ) ; Wau, M t . K a i n d i , 4. V . 1971, R . E . P a r r o t t (1 S, CNCI) .

R e m a r k s : The numerous setae on a l l e l y t r a l i n t e r v a l s combined wi th the d i s t i n c t l y punc­tu red but e n t i r e l y glabrous pronotum are diagnostic for th is species.

Casnonidea punct i thorax sp. n . (Figs 30, 33)

Body usua l ly dark r edd i sh brown to black, w i t h head and pronotum and pronotum dark­e r , e l y t r a s l i gh t ly pa ler ; e l y t r a somet imes i r r e g u l a r l y mot t led wi th ochre; i n one case, p r o ­notum and basal 1/3 of e ly t r a pale ochre , res t of e l y t r a mot t led . Length: 9. 2-10. 1 m m .

S. Head not broader than pronotum; w i t h some set igerous punctures between eyes; ver tex moderate ly swol len , w i t h d i s t i nc t longi tudina l f u r r o w ; EI= 14: 12; antennae ra ther s lender , segment length r a t i o s : 10: 5: 11: 11 ; 11 : 13: 13: 14: 14: 14: 23. Pronotum s l igh t ly t rans­verse , widest in an te r io r 1/3, base near ly as wide; surface wi th few scat tered punctures (situated main ly l a t e r a l l y ) bear ing modera te ly long, w h i t i s h , semierec t setae. E l y t r a w i th a l l punc tura l rows reaching apex; s t r i a i punctures modera te ly coarse, contiguous; i n t e r v a l s less convex; a l l i n t e rva l s w i t h many (over 30) set igerous punctures sca t tered throughout whole length; these not a r ranged in two r o w s ; setae w h i t i s h , fine, moderate ly long; e l y t r a l apex s i m p l y rounded. Legs w i t h ha i rs r a t h e r shor t , nea r ly decumbent, fine. Last abdominal s te rn i te s i m p l y rounded. Aedeagus: F i g . 33. Habi tus : F i g . 30.

g. V e r t e x a l i t t l e more swol len ; EI= 15: 13; r e l a t i ve lengths of antenna! segments V I I I to X I : 14: 14: 13: 2 1 .

Holotype , t, label led as fo l lows : " N E W G U I N E A : NE K a r i m u i , 1080 m , 13. V I I . 1963"; " J . Sedlacek BISHOP". I t is deposited i n B P B M .

Paratypes (total 12 specimens) . Eas t e rn Highlands Prov ince : label led as holotype (4 SSt

2 gg, B P B M ; 1 S, H N H M ex B P B M ) ; i d . , 11-12. V I I . 1963 (1 o., B P B M ) ; i d . , 1973, R. H o r n a -

brook i l o, H P C W ) ; Okapa, I . 1968, R. Hornabrook (1 ç, HPCW). - Western Highlands : Ba iye r R i v e r , Sanctuary, 1 - 5 . I X . 1969 [ N G B - U . 2 2 . ] , J . Balogh (1 3, HNHM); i d . NGB-U. 2 i . (1 g, H N H M ) .

Remarks : The most cha rac t e r i s t i c feature of th is species is the sparsely punctate and pubescent p rono tum. Casnonides h y s t r i x also has the punctate p ronotum, but i n that spe­cies , the puntat ion is much denser, the puntures are not bear ing setae, the colour pa t te rn , the shape of body and the r e l a t i ve length of the last antennái segment of male are s t r i k i n g l y different .

F igs 3 0 - 3 1 . 30= Casnonidea punct i thorax sp. n . , 8. - 31= C. apical is sp. n . , 8

Casnonidea apica l is sp. n . (Figs 31 , 34)

Head ye l l ow , somet imes pa r t ly b lack i sh ; p ronotum ye l low; e l y t r a l a rge ly ye l low wi th apex black to a va r i ab le extent (usually f r o m l eve l of abdominal s te rn i te I V ) ; ep ip leura black­ish in pos t e r io r par t , a n t e r i o r l y to the l eve l of the middle of metas te rnum (somet imes this co loura t ion reaches e l y t r a l base); v e n t r a l surface ye l low o r meso- and metas te rnum black­i sh ; antennae black, segments I and I I y e l l o w i s h ; legs ye l low to b lackish . Length: 7. 8-8. 2 m m .

6*. Head not broader than p rono tum; w i t h two flat impress ions between eyes; ve r tex weakly swol len; EI= 12: 14; antennae r a t h e r stout, r e l a t ive length r a t ios of segments: 8:4: 9 : 9 : 9 : 1 0 : 1 0 : 1 0 : 1 0 : 9 : 2 0 . P rono tum t r ansve r se , widest in an te r io r 1/4, base much nar­r o w e r : surface impuncta te , g labrous . E l y t r a w i th a l l punc tura l rows reaching apex; s t r i a i punctures coarse , separated by distances shor te r than puncture d iamete r ; i n t e rva l s convex; pa i red in t e rva l s en t i r e l y glabrous; unpa i red in te rva l s w i t h set igerous punctures scat tered throughout whole length; i n t e r v a l I w i t h 17, I I I wi th 18, V w i t h 17, V I I w i t h 18, IX wi th 18, l a t e r a l ma rg in w i t h 12 set igerous punctures ; setae b rowni sh , modera te ly long; apex s imp ly rounded. Legs wi th ha i r s moderate ly long, suberect. Las t abdominal s t e rn i t e s i m p l y round­ed. Aedeagus: F i g . 34. Habi tus : F i g . 31 .

37

U 38 39 40

Figs 32-40. 32= Aedeagus of Casnonidea h y s t r i x sp. n . - 33= Aedeagus of C. punct i thorax sp. n . - 34= Aedeagus of C. apica l is sp. n. - 35= Aedeagus of C. flavipes sp. n . - 36 = Aedeagus of C. baloghi sp. n . - 37= Aedeagus of C. loksa i sp. n . - 38= Aedeagus of C. do-bodura sp. n . - 39= Aedeagus of C. pallens sp. n . - 40= Aedeagus of C. demet r ida sp. n.

g. Ve r t ex a l i t t l e more swol len ; EI= 12: 15; r e l a t ive lengths of antennái segments V I I I to X I : 10 :10 :10 :17 .

Holotype , Ô*, labe l led as fo l lows : " N E W GUINEA (NE) Wau, Mt . Ka ind i 24-25. V I I I . 1968"; " (No . N G W - C . 6. ) leg. D r . J . Ba logh" . I t is deposited in H N H M .

Paratypes (total 11 specimens) . Morobe Prov ince : B u l o l o - V a t u t , 700 m , 15-21 . V I . 1969. JS (1 S, B P B M ) ; Wau, 1050-1150 m , 13. X . 1961-7.1.1963, JS, JHS (4 Stf. 1 <j>, B P B M ; 1 6, 2 22> SPCB; 1 5, H N H M ex B P B M ; 1 3, H N H M ex SPCB).

R e m a r k s : Th i s species is s i m i l a r to Casnonidea v a r i a b i l i s (Macleay, 1887), but the colour pat tern of the e l y t r a c l ea r l y dis t inguishes i t .

Casnonidea f lavipes sp n . (Figs 35, 41)

Head, p ronotum, scu te l lum and v e n t r a l surface dark b lackish brown to black; e l y t r a b lackish b r o w n , r a r e l y un ico lo rous , usua l ly w i t h a va r i ab le number of i r r e g u l a r pale spots spots may p a r t l y be confluent and more numerous on humera l par t ; antennae brown wi th 2 basal segments somewhat redd i sh ; legs pale y e l l o w , often semi t rans lucen t . Length: 9. 5-11,2 m m .

S. Head not broader than pronotum; w i t h 2 shal low impress ions between eyes; EI= 15:16; antennae ra the r s lender ; segment length r a t i o s : 1 1 : 5 . 5 : 1 3 : 1 3 : 1 4 : 1 4 : 1 5 : 1 6 : 1 6 : 1 5 : 2 2 . Pronotum d i s t i nc t l y t r ansve r se , base and a n t e r i o r 1/4 of about same wid th ; surface gla­brous , impuncta te , shining, often wi th 2 ind i s t inc t l a t e r a l depress ions . E l y t r a w i t h a l l punc­t u r a l rows reaching apex; s t r i a i punctures moderate ly coarse , separated by distances at least equal to puncture d i ame te r ; in te rva l s weakly convex; pa i red in t e rva l s en t i r e ly gla­brous; unpai red in t e rva l s w i t h a few scat tered set igerous punctures; i n t e r v a l I w i th 1 sub-basal, I I I w i t h 6, V wi th 2, V I I w i t h 4, IX w i t h 5, l a t e r a l m a r g i n wi thout set igerous punc­tu res ; setae da rk , f ine, inconspicuous; e l y t r a l apex s imp ly rounded. Legs w i t h pubescence reduced; femora glabrous; t ib iae w i t h a few decumbent ha i r s along inner edge. Last ab­dominal s te rn i te s i m p l y rounded. Aedeagus: F i g . 35. Habi tus : F i g . 4 1 .

F igs 41-42. 41= Casnonidea flavipes sp. n . , Ô*. - 42= C. baloghi sp. n . , Z3

<j>. No shallow i m p r e s s i o n s between eyes but ve r tex m o r e swol len , longi tudinal fu r row more d i s t i n c t ; EI= 14: 17; r e l a t i ve lengths of antennái segments V I I I to X I : 16: 16: 15:20.

Holotype , 3, l abe l led as fo l lows : "NEW GUINEA (NE) Wau, 22. I X . -30 . I X . 1969. " ; "(No. N G W - U . 33. ) l eg . D r . J . Ba logh" . I t : i s deposited i n H N H M .

Paratypes (total 17 specimens) . Morobe P rov ince : Wau, 1200-1268 m , I . 1966-6. 1. 1971, B . B . L o w e r y , JS (1 g, A N I C ; 2 88, SPCB); Wau, B i g Wau Ck. , 1200-1500 m , I X . 1965., JS, MS (1 o_, B P B M ) ; Wau, Wau Ck. , 17. I X . 1964, JS (1 p, SPCB); 6 k m W of Wau, N a m i Creek , 1700 m , 12. V I . 1962, JS (1 8, B P B M ) ; i d . , 15. V I . 1962, JS (1 8, B P B M ) ; M t . K a i n d i , 1500-2200 m , 19. V I I I . 1965, JS, MS (1 p, B P B M ) ; M t . M i s s i m , 2000 m , JS (1 g, H N H M ex SPCB); Teptep, Huon Peninsula, 10. V I I . 1975, R . W . Hornabrook (1 o., HPCW). - Madang P r o v i n c e : F i n i s t e r r e M t s . , M o r o c. 5550 f t . , 30. X . -15 . X I . 1964, M . E . Bacchus (1 6, B M N H ) . - West­e r n Highlands Prov ince : M t . Hägen area , Ba iye r R i v e r , Sanctuary, 4 0 0 0 ' , 6 -11 . V I I . 1974, H . Howden (1 8, CNCI) . - Eastern Highlands P rov ince : Okapa, 17. X . 1964, at l i gh t , R. H o r ­nabrook (1 p, HPCW); i d . , 1. 1968, R. Hornabrook (1 8, H N H M ex H P C W ) ; i d . . 1800 m . 25. V I L , JS (1 3, SPCB); Purosa , 1700 m , 17-25. V . 1966, Gres s i t t et T a w i (1 g, B P B M ) . - Cen­t r a l P r o v i n c e : Owen Stanley Range, G o i l a l a , Borne, 1950 m , 16-30. I V . 1954, W . W . B r a n d t (1 o_, B P B M ) .

R e m a r k s : The most cha rac t e r i s t i c feature of this species is the pale yel low co lour which is i n sharp contrast to the b lack i sh res t of body. Specimens w i t h mot t led e l y t r a some­what resemble Casnonidea punct i thorax , but the p rono tum is glabrous and smooth (hairy and punctate i n punct i thorax) , only unpaired in t e rva l s have a s m a l l number of set igerous punc­tures ( a l l i n t e rva l s have a lo t of them in punct i thorax) and legs ye l low (dark brown i n punc­ t i t h o r a x ) .

Casnonidea baloghi sp. n . (Figs 36, 42)

Body dark blackish b rown . Length: 10 .5 -12 .2 m m . 3. Head not broader than p rono tum; w i t h 2 or 3 flat depressions between eyes; EI=

16: 16; antennae ra ther s lender ; segment length r a t i o s : 12: 6: 14: 12: 14: 14: 15: 15: 14: 13: 20. P r o n o t u m as long as broad o r a l i t t l e t r ansve r se ; surface impunctate , g labrous , sh in ing . E l y t r a w i t h most punctura l rows reaching apex but IV to IX d i s in tegra t ing p reap ica l ly ; most rows becoming f u r r o w - l i k e i n pos te r io r 1/3; s t r i a i punctures coarse , contiguous, i n l a t e r a l rows wide ly separated; i n t e rva l s weakly convex; pa i red in te rva l s e n t i r e l y glabrous; unpa i r ­ed i n t e r v a l s w i t h a few set igerous punctures ; i n t e r v a l I w i th 6, I I I w i t h 6, V wi th 7, V I I w i t h 4, IX., w i t h 5, l a t e r a l m a r g i n wi th 3 (preapical) set igerous punctures; setae pale, shor t ; e l y t r a l apex s i m p l y rounded. Legs w i t h few semierec t h a i r s . Las t abdominal s te rn i te s i m p l y rounded. Aedeagus: F i g . 36; Habi tus: F i g . 42.

o. Depressions between eyes v e r y s m a l l , punc tu re - l i ke ; EI= 18: 18; length ra t ios of antennái segments V I I I to X I : 16: 16: 14:22; p ronota l sides w i t h an ovoid , rugulosely punctate area.

Holo type , S, label led as fo l lows : " N E W GUINEA: NE Wau, 1150-1600 m , 9. I I . 1968"; " J . Sedlacek Co l l ec to r BISHOP". I t i s deposited in B P B M .

Paratypes (total 17 specimens) . Morobe Prov ince : labe l led as holotype (1 8, H N H M ex B P B M ) ; Wau, 1150-1550 m , 6. I I . 1963-27. DC. 1969. JS, MS, R. Hornabrook (2 3 8 , 1 p , B P B M ; 1 8 , HPCW; 4 8 8 , 2 oo., SPCB; 1 8, 1 o., H N H M ex SPCB); Wau, Haus Copper, M t . M i s s i m , 22-24 . I V . 1965, J . Balogh et J . J . Szent-Ivány (1 p, H N H M ) ; Wau, K u j e r u , 1500 m j 27. I X . 1969, A . B . M i r z a (1 p. B P B M ) ; Wau. Wau Ck. , 29. I I I . 1963, JS (1 3 , SPCB); i d . , I 1200-1500 m , 6. V . 1963, JS (1 Q_, B P B M ) .

R e m a r k s : The most c h a r a c t e r i s t i c feature of this species is the loose ly si tuated s t r i a i i punctures , ma in ly in the l a t e r a l r o w s . The rugulose p ronota l side of female is found in Cas-! nonidea loksa i as w e l l , but the combinat ion of unicolorous blackish body, the reduced e l y t r a l ves t i tu re and the fo rmat ion of the p reap ica l par t of e l y t r a l s t r iae r ead i ly d is t inguish C. ba- j l ogh i .

The species is dedicated to D r . J . B A L O G H (Zoosys temat ica l and Eco log i ca l InstituteT Eötvös Loránd U n i v e r s i t y , Budapest).

Casnonidea loksa i sp. n . (Figs 37, 43)

Head dark reddish b rown; pronotum and scu te l lum pale reddish b r o w n ; e l y t r a b lack i sh brown; v e n t r a l surface reddish brown; antennae and legs b rown, f emora l base and 2 basal segments of antennae somewhat paler . Length : 10 .2-10. 5 m m .

S. Head not broader than p rono tum; wi th 2 shallow depressions between eyes ; EI= 14: 14; ve r tex weakly swol len , longi tudinal f u r r o w faintly i m p r e s s e d ; antennae ra ther s lender ; segment length ra t ios : 1 1 : 5 : 1 3 : 1 2 : 1 3 : 1 2 : 1 4 : 1 4 : 1 5 : 1 4 : 2 1 . Pronotum a l i t t l e t rans ve r se ; widest in a n t e r i o r 1/3; surface glabrous, sh in ing , w i th v e r y ind is t inc t punctures on sides around a n t e r i o r angles. E l y t r a wi th a l l punc tu ra l rows reaching apex; rows s t rongly impres sed ; s t r i a ! punctures modera te ly coarse, contiguous; i n t e rva l s convex; pa i red inter-

Figs 43-44. 43= Casnonidea loksai sp. n . , S. - 44= C. pallens sp. n . , 5*

vais g labrous , except i n t e r v a l I I which bears 4 subbasal setae; unpai red in t e rva l s w i th sev­e r a l set igerous punctures sca t te red throughout whole length; these punctures ar ranged in 2 loose ser ies i n inner unpaired i n t e r v a l s ; i n t e r v a l I w i th 26, I I I w i t h 39, V w i t h 30, V I I wi th 20, IX wi th 25, l a t e r a l m a r g i n w i t h 32 set igerous punctures; setae pale, r a ther long; e l y t r a l apex s imply rounded. Legs w i t h moderately long , semie rec t h a i r s . Last abdominal s te rn i te s imply rounded. Aedeagus: F i g . 37. Habitus: F i g . 43.

<j>. Depress ions between eyes ve ry s m a l l ; ve r t ex s t rongly swol len , longi tud ina l fur row

deeply impressed ; EI= 15: 15; length r a t ios of antenna! segments V I I I to X I : 14: 15: 14: 19. 5; p rono tum coarse ly rugose l a t e r a l l y .

Holotype , 8, l abe l led as fo l lows : "NEW G U I N E A : NE Wau, 12- 1600 m 9 - I I 1968"; " J . Sedlacek C o l l e c t o r BISHOP". I t is deposited in B P B M .

Paratypes ( total 3 specimens): Morobe P rov ince : Wau, 8. I X . 1968 [ N G W - R . 1 9 ] , I . Loksa (1 o., H N H M ) ; i d . , 4000 f t . , 1 - 3. V I I . 1974, H . F . Howden (1 8, CNCI) . - Madang P r o ­v ince : F i n i s t e r r e M t s . , Damant i , 3500 f t . , 2 - 1 1 , X . 1964, M . E . Bacchus (1 c-, B M N H ) .

R e m a r k s : On account of the rugose pronota l sides of female, this species is somewhat re la ted to C. baloghi . The pronotum and v e n t r a l surface of C. loksai are reddish (black in baloghi) , setae of i n t e r v a l s are numerous and long (few and short in baloghi) , punc tura l rows n o r m a l (d is in tegra t ing before apex in baloghi) .

T h i s species is dedicated to D r . I . LOKSA (Zoosys temat ica l and E c o l o g i c a l Ins t i tu te , Eötvös Loránd U n i v e r s i t y , Budapest).

Casnonidea dobodura sp. n . (Figs 38, 45)

Body dark b lack ish brown; antennae, palps, t ib iae and t a r s i somewhat paler . Length: 10. 0- 10. 2 m m .

45 46 47

Figs 45-47. 45= Head of Casnonidea dobodura sp. n . , 8. - 46= E l y t r a l apex of C. caudata sp. n . , o.. - 47= A b d o m i n a l apex of C. caudata sp. n . , <j>, v e n t r o l a t e r a l v iew

8. Head (F ig . 45) not broader than p rono tum, l a r g e l y impunctate ; ve r tex moderate ly swol len , w i th long i tud ina l fu r row d i s t inc t ; EI= 18:7; antennae ra ther s lender; segment length r a t i o s : 1 0 : 5 : 1 1 : 1 1 : 1 2 : 1 3 : 1 2 : 1 2 : 1 3 : 1 2 : 2 3 . P rono tum near ly as long as broad, wides t in a n t e r i o r 1/3; base a l i t t l e n a r r o w e r ; surface g labrous , impunctate . E l y t r a w i t h a l l punc­t u r a l rows reaching apex; s t r i a i punctures r a the r coarse , contiguous; i n t e rva l s fa in t ly con­vex; pa i red in t e rva l s e n t i r e l y glabrous; unpai red in t e rva l s w i t h setigerous punctures scat­t e red throughout whole length; i n t e r v a l I w i t h 21 , I I I w i t h 19, V wi th 20, V I I w i t h 17, IX w i t h 16. l a t e r a l side w i t h 19 set igerous punctures; setae black, moderate ly long; e l y t r a l apex s i m p l y rounded. Legs w i t h ha i r s shor t , suberect , a l i t t l e f i ne r than e l y t r a l setae. Las t ab­dominal s te rn i te s i m p l y rounded. Aedeagus: F i g . 38.

g. EI= 18:8 (antenna! segments p a r t l y lost, on the s ingle female).

Holotype, Ó1, label led as fo l lows : "Dobodura , Papua, N . G . M a r - J u l y , 1944 Dar l ing ton" . I t is deposited i n M C Z C . i

Paratype. Western P rov ince : Kiunga, 28-30. V I I . 1969 [ N G K - U . 3 . ] , J . Balogh (1 o_, H N H M ) .

R e m a r k s : The ex t r eme ly n a r r o w in t e rocu l a r distance makes this species eas i ly d i s ­t inguishable . The male holotype f rom Dobodura (Nor thern Prov ince) has a l l body par ts i n ­tac t , though the e l y t r a l ve s t i t u r e has l a r g e l y been w o r n down. The female paratype lacks the left middle and hind legs as w e l l as a l l but left basal 4 antennái segments, wh i l e the e l y t r a l ve s t i t u r e is a lmost in tac t .

Casnonidea caudata sp. n . (Figs 46, 47)

Head, p rono tum, scu te l lum and v e n t r a l surface reddish ; e l y t r a black w i t h me ta l l i c g reen i sh -b lu i sh sheen; antennae and legs black, segments I and I I of antennae as w e l l as femora l base redd i sh . Length: 11.2 m m .

Ç. Head a l i t t l e n a r r o w e r than p rono tum; w i t h 2 s m a l l punctures between eyes; EI= 17: 17; ve r t ex s t rong ly swol len , w i th wel l -deve loped long i tud ina l fu r row; antennae ra the r stout; segment length r a t i o s : 1 1 : 5 : 1 1 : 1 3 : 1 3 : 1 3 : 1 4 : 1 4 : 1 5 : 1 3 : 2 2 . Pronotum weakly t r ans ­ve r se , widest at base; surface g labrous , impuncta te , sh in ing. E l y t r a wi th only punctura l rows I and X reaching apex, so e l y t r a w i t h a smooth p reap ica l area; rows v e r y weakly i m ­pressed; s t r i a i punctures minute , subcontiguous; i n t e rva l s broad and f la t ; p a i r e d in t e rva l s e n t i r e l y g labrous ; unpaired in t e rva l s w i t h a few scat tered set igerous punctures ; i n t e r v a l I w i t h 8, I I I w i th 5, V wi th 6, V I I w i t h 6, IX w i t h 6, l a t e r a l m a r g i n wi th 11 se t igerous punc­tu res ; setae black, shor t ; e l y t r a l apex bicuspidate , i . e . e l y t r a having an outer subrectan-gular and an inne r rounded angle ( F i g . 46). Legs w i t h ha i r s shor t , sparse, s e m i e r e c t . Las t abdominal s t e rn i t e w i th a process d i r ec t ed upwards (F ig . 47).

S. Unknown. Holotype, p , label led as fo l lows : " N E W GUINEA (NE) B a i y e r R i v e r Sanctuary 1-5. DC.

1969"; " (No. N G B - U . 24J leg. D r . J . Ba logh" . I t i s deposited in H N H M . R e m a r k s : Though known by the sole female specimen, Casnonidea caudata i s c l e a r l y

dis t inguished f r o m a l l other descr ibed congeners by i t s bicuspidate e l y t r a l apex and by hav­ing a process on the last abdominal s t e rn i t e . In other features i t much resembles C. s e m i - violacea P ic .

Casnonidea pallens sp. n . (Figs 39, 44)

Body pale chestnut in co lour ; e l y t r a somet imes wi th weak violaceous sheen, main ly i n basal ha l f or in humera l par t . Length: 8 .2 -10 .0 m m .

<?. Head not broader than p rono tum; wi th 2 flat punctures between eyes (sometimes confluent and f o r m i n g a t r ansve r sa l f u r r o w ) ; EI= 15: 15; antennae ra ther s lender ; segment length r a t i o s : 10: 5: 10: 10: 11 : 10: 10: 10 :9 :9 : 29. P rono tum as long as or a l i t t l e longer than broad, widest in an te r io r 1/4, base of same wid th ; surface en t i r e ly g labrous , impuncta te , shining. E l y t r a w i t h only punctura l rows I and X reaching apex, so e l y t r a w i t h a smooth preapica l area; s t r i a i punctures modera te ly coarse , contiguous; in t e rva l s f la t ; pa i red i n ­te rva l s e n t i r e l y glabrous; unpai red in t e rva l s w i t h ves t i tu re ex t remely reduced; i n t e r v a l I w i t h 1 subbasal, I I I w i th 1 subbasal and 2 p reap ica l , V wi thout , V I I wi th 2 subbasal , IX w i t h 2 p reap ica l , l a t e r a l m a r g i n without se t igerous punctures; setae dark, f ine, inconspicuous; e l y t r a l apex s i m p l y rounded; weakly caudate in do r so l a t e r a l aspect. Legs w i t h pubescence reduced; femora glabrous; t ibiae wi th f ine, decumbent ha i r s along inner edge. Las t abdom­ina l s te rn i te s i m p l y rounded. Aedeagus: F i g . 39. Habi tus : F i g . 44.

o.. V e r t e x a l i t t l e more swol len ; EI= 13: 15; length r a t ios of antennái segments V I I I to X I : 9 : 1 1 : 10: 22.

Holotype , S, l abe l led as fo l lows : " N . | Guinea: NE B u l o l o , 700 m 26. X I . 1969"; " J . et M . Sedlacek Co l l ec to r s BISHOP M U S E U M " , I t i s deposited in B P B M .

Paratypes (total 60 specimens) , Morobe P rov ince : labe l led as holotype (4 #<?, 4 PP, B P B M ; 1 Î , 1 p . H N H M ex B P B M ) ; B u l o l o , 700 m (1 S, SBCB); i d . , 6. X I . 1969, JS (3 &S, . 1 o_, B P B M ; 1 S, H N H M ex B P B M ) ; i d . , 1130 m , 17. I X . 1969, J . et M . Sedlacek (1 S, BPBM);

Pind iu , 20. I V . 1963, JS (1 3, B P B M ) ; i d . , 870-1300 m . 21-22. I V . 1963, JS (1 g, B P B M ) ; Watut R. , 900-1900 m . X . 1969, A . B . M I R Z A (1 8, B P B M ) ; Wau, 1050- 1300 m , 2. I X . 1 9 6 1 -X I I . 1968, H . Oh lmus , JS, JHS, MS (1 g, A N I C ; 9 88, 8 gg, B P B M ; 1 8, 2 gg, SPCB; 1 8, 3 gg, H N H M ex B P B M ) ; id . , 1200 m , 8-14. X I I . 1976, m e r c u r y vapor l i gh t , G. F . Heve i et i R. E . Die tz (4 Í3, USNM; 1 3, H N H M ex USNM); Wau, K i l o l o Creek . 26. V I I I . 1968 [ N G W - C j 14 . J , J . Balogh (1 g, H N H M ) ; Wau, M t . M i s s i m , 950-1300 m . 8-9 .1 .1966, J . et M . Sedla­cek (1 S, B P B M ) . - Madang P rov ince : Madang, 25. I X . 1970, R. Hornabrook (1 ô*. H P C W ) . -Eas te rn Highlands P rov ince : K a r i m u i , I I I . 1973, R. Hornabrook (1 S, HPCW); Okapa, 3500 f t . , 6 . I X . 1965, R. Hornabrook (1 8, HPCW). - Wes t e rn Highlands P rov ince : B a i y e r R i v e r . Sanctuary, 1 -5 . IX . 1969 [ N G B - U . 2 3 . ] , J . Balogh (1 g, H N H M ) . - C e n t r a l P rov ince : B i s i a -numu, E of Po r t Moresby , 500~m. 22. I X . 1955, C r o t a l a r i a f l o w e r s , J . L . Gre s s i t t (1 8, B P B M ) ; i d . , 2 3 . I X . 1955 (1 3, B P B M ) , - Western P r o v i n c e : O r i o m o Govt . Sta., 2 6 - 2 8 . X . 1960, shelf fungus, J . L . G r e s s i t t (1 g, B P B M ) . - N o r t h e r n P r o v i n c e : Dobodura, I I I - V I I , 1944. Da r l i ng ton (1 6, M C Z C ) .

R e m a r k s : Th i s species is r e a d i l y d is t inguishable f r o m the descr ibed congeners by the fo l lowing combina t ion of cha rac te r s : pale b rownish co lou r , smooth p reap ica l area of e l y t r a and the v e r y reduced e l y t r a l v e s t i t u r e .

Casnonidea demet r ida sp. n . (Figs 40, 48)

Dorsa l surface reddish b r o w n , often w i t h h u m e r a l par t of e l y t r a b lack ish ; this dark coloura t ion extending to middle of e l y t r a ; v e n t r a l surface reddish b r o w n , metas te rnum somet imes b lackish ; antennae b rowni sh , somewhat d a r k e r t oward apex; legs reddish b rown; d i s t a l half of femora and p r o x i m a l hal f of t ib iae b lack i sh (much more expressed on fore and middle legs) . Length: 7 . 5 - 9 . 0 m m .

3. Head a l i t t l e broader than pronotum; w i t h 3 s m a l l round pi ts between eyes; EI= 11 : 16; ve r t ex weakly swol l en , long i tud ina l fu r row i n d i s t i n c t ; antennae ra the r s lender; seg­ment length r a t i o s : 9: 4: 8: 9: 10: 10: 11 : 11 : 11: 11 : 17. P rono tum a l i t t l e longer than b road , widest in an t e r io r 1/4; surface impuncta te , sh in ing , l a rge ly glabrous but w i t h a few setae in an t e r io r angles. E l y t r a d i s t i n c t l y flattened; only punc tu ra l rows I and X reaching apex, so e l y t r a w i t h a smooth p reap ica l area; s t r i a i punctures modera te ly coarse , contiguous; in-j t e rva l s f la t ; pa i red in t e rva l s e n t i r e l y glabrous; unpai red in t e rva l s w i t h set igerous punctures scat tered throughout whole length; i n t e r v a l I w i t h 14, I I I w i t h 16, V w i t h 17, V I I w i t h 12, IX w i t h 13, l a t e r a l m a r g i n w i t h 19 set igerous punctures; setae black, f ine, ra ther long; e l y t r a l apex s i m p l y rounded. Legs w i t h pubescence f ine , r a t h e r long, semie rec t . Las t ab­domina l s t e rn i t e s imply rounded. Aedeagus: F i g . 40. Habi tus : F i g . 48.

g. Pi ts between eyes much s m a l l e r ; middle p i t t o t a l l y i n d i s t i n c t ; EI= 10: 14; length ra t ios of antennái segments V I I I to X I : 10: 11: 10: 15.

Holo type , 8, label led as fo l lows : Papua New Guinea, Wau, 19. I X . 1972., Malaise t r a p , leg . I - , Móczár". I t i s deposited in H N H M .

Paratypes (total 13 specimens) . Morobe P rov ince : B u l o l o R. , 1130 m , 17. DC. 1969, A . B . M i r z a (1 3. B P B M ) ; M t . K a i n d i , 1500-2200 m , 19. V I I I . 1965, J . et M . Sedlacek (1 o, B P B M ) ; Wau, 1100- 1400 m , 27. V I I I . 1961-4. I I . 1968, R. Hornabrook , JS, MS (2 33, 4 gg, B P B M ; 1 3, HPCW); Wau, M c A d a m park , 18-21 . I V . 1965, J . Balogh et J . J . Szent-Ivány ( 1 Í , H N H M ) ; Wau, Haus Copper , M t . M i s e i m , 22-24. I V . 1965, J . Balogh et J . J . Szent-Ivány (1 8, H N H M ) ; Wau, M t . M i s s i m , 1200 m,: 28. V I I I . 1968 [ N G W - C . l l ] , J . Balogh (1 g , H N H M ) ; 6 k m W of Wau, N a m i Creek , 1200 m , 15. V I . 1962, JS (1 g, B P B M ) . ,

R e m a r k s : Th i s species may be recognized by the fo l lowing combinat ion of charac te r s large but not v e r y prominent eyes, by which the head is broader than p rono tum; flat tened e l y t r a wi th smooth p reap ica l area; a number of soft, fine ha i r s i n unpai red e l y t r a l i n t e r ­va ls ; r a the r long, f rag i le legs w i t h modera te , s emie rec t pubescence; an t e r io r angles of pronotum w i t h few h a i r s .

Casnonidea setosa sp. n . (F ig . 49)

Body reddish b rown, metas te rnum and basal ha l f of e l y t r a a l i t t l e da rke r . Length: 8. 7 m m .

g. Head d i s t i nc t l y broader than pronotum; w i t h 2 confluent pits between eyes; eyes

conspicuously prominent ; EI= 13: 15; ver tex weakly swol len , long i tud ina l fu r row Ind i s t inc t ; antennae ra the r slender; segment length r a t i o s : 1 0 : 4 : 9 : 1 0 : 1 1 : 1 1 : 1 2 : 1 1 : 1 2 : 1 1 : 1 7 . P r o ­notum longer than broad, widest i n an t e r io r 1/3; surface impuncta te , sh in ing , w i t h a few setae in an t e r io r angles. E l y t r a d i s t i n c t l y f la t tened; a l l punc tura l rows reaching apex; s t r i a i punctures modera te ly coarse , not contiguous; i n t e r v a l s fa int ly convex; pa i red in t e rva l s en-t i r e l y a g l a b r o u s ; unpaired i n t e r v a l s w i th se t igerous punctures sca t te red throughout whole length; i n t e r v a l I w i th 12, I I I w i t h 2 1 , V w i t h 17, V I I wi th 18, I X w i t h 15, l a t e r a l m a r g i n w i t h 25 set igerous punctures; setae dark , f ine , v e r y long; e l y t r a l apex s imply rounded. Legs wi th conspicuously long, s e m i e r e c t to e rec t ha i r s . Last abdominal segment s i m p l y rounded. Habi tus : F i g . 49.

F ig s 48-49. 48= Casnonidea demet r ida sp. n . , <3. - 49= C. setosa sp. n . , <j>

S. Unknown. Holotype , o, label led as fo l lows : " N E W GUINEA (NE) B a i y e r R i v e r Sanctuary 1-5. D i .

1969"; " (No. NGB-U.24.) l eg . D r . J . Ba logh" . I t is deposited in H N H M . R e m a r k s : This species i s v e r y s i m i l a r to Casnonidea demet r ida but a l l the punctura l

rows of e l y t r a reach the apex (only I and X j i n demet r ida) . M o r e o v e r , the eyes are more prominent , the ha i rs on the e l y t r a and p a r t i c u l a r l y on the legs are v e r y long.

REFERENCES

B O R C H M A N N , F r . (1936): Coleoptera F a m . L a g r i i d a e . - I n : W Y T S M A N , P. (ed.): Genera Insec to rum, fasc. 204. Louis D e s m e t - V e r t e n e u i l , B r u x e l l e s , 561* pp.

M E R K L , O. (1988): O r e o g r i a gen. n . f r o m New Guina (Coleoptera, Tenebr ionidae : L a g r i i n i ) . Ac ta zool . hung. 34 (2-3) : 247-271. I

P IC, M . (1956): Nouveaux Coléoptères de d iverses f a m i l l e s . - Annls h i s t . - n a t . Mus . natn . hung. (ser. n.) 7: 71-92.

A u t h o r ' s address: D r . O. M E R K L Zoo log ica l Depar tment Hungar ian N a t u r a l H i s t o r y Museum Baross u . 13 H - 1088 Budapest HUNGARY