growth and development of trees - college of agriculture ...growth and development of trees...
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Growth and development of Trees
Objectives:
1. To study the morphological and
physiological processes that occur for a
temperate deciduous tree during the annual
cycle, and the whole life cycle.
2. To use this information to make informed
management decisions.
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Life and Annual Cycle
Life
Germination
Juvenility/Maturity
Old age
Annual
Dormancy
Growth
Vegetative
Reproductive
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Example of Oak Life Cycle
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Juvenility Plants are non-reproductive for several years,
even if given correct environmental conditions
Length varies with species (1-45 yrs)
Also may have different characteristics:
a. Leaf shape (lobed vs. not lobed)
b. Ease of rooting
c. Stem anatomy (cambial growth)
d. Presence of thorns (ex. Honey locust)
e. Production of pigment
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Example: English Ivy
Juvenile:
grows along ground
lobed leaves
Mature:
grows upright
leaves are not lobed
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Juvenility
Length of time related to vegetative stature
Longer juvenile period generally results in a larger
plant
Transition zone
Zone between juvenile and adult tissue
Rejuvenation
Cut back and let re-grow
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Juvenile and Adult Tissues can be grafted together
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There is no Transition Zone between Juvenile and Adult Tissue on a Grafted Plant
seedling grafted
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Seasonal Cycles
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CHERRY ANNUAL CYCLE Calendar date basis
MORPHOLOGY Leaves
Leaf abscission
Roots
Trunk
Fruit
Cell division
Pit hardening
Final swell
Abscission
Harvest
FBI
F-Development
Physiology Dormancy
Summer
Rest
Spring
JAN FEB MAR APR MAY JUNE JULY AUG SEPT OCT NOV DEC
A H
Temp, low and high temp, light, water, nutrition, biological temp, nutrition
160-180 days
58-65 days
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Types of Dormancy
Ecodormancy
Environmental factors
Paradormancy (ectodormancy)
Physiological factors outside the plant
Endodormancy
Physiological factors within the plant (shoot apex)
The transition into and out of a dormant state
is gradual (quiescence or predormancy)
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Dormancy Definitions
Ecto-dormancy
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Example of how a Plant may Experience Different Types of
Dormancy throughout a Season
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CHERRY ANNUAL CYCLE MORPHOLOGY Leaves
Leaf abscission
Roots
Trunk
Fruit
Cell division
Pit hardening
Final swell
Abscission
Harvest
FBI
F-Development
Physiology Dormancy
Summer
Rest
Spring
JAN FEB MAR APR MAY JUNE JULY AUG SEPT OCT NOV DEC
A H
Temp, low and high temp, light, water, nutrition, biological temp, nutrition
160-180 days
58-65 days
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Cold Hardiness in Montmorency
Jul Aug Sept Oct Nov Dec Jan Feb Mar Apr May June
Temp, low and high temp, light, water, nutrition, biological temp, nutrition
30
20
10
0
-10
-20
-30
LT5
0 o
F
Acclimation
Deep winter hardiness
De-acclimation
Killing frost
Short days
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Breaking dormancy! Chilling Requirement
Definition:
# of hours of temp between 0-10°C required to
break endodormancy (regulated by physiological
factors inside the affected structure)
How? Inhibitors, promotors?
Advantages?
Disadvantages?
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Chilling Requirements
Low-chilling + Longer productive
season
- more susceptible to
late frost damage
High-chilling
+ protected against
late frost
- Shorter growing
season
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Then in Spring… Heat Units: quantitative term used to describe
the accumulation of heat used to break ectodormancy (also called growing degree days)
Traditional method of calculation using max, min.
Degree Days calculation:
(Max+Min)/2 - base temp = degree days
example: (70+50)/2 - 50 = 10 degree days
used to: predict bud break, harvest date, bloom date,
vegetative growth, insect and disease growth (IPM)
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Degree days
Other methods of calculation
Baskerville emin method. Uses sine function
for change in temperature.
Different DD bases: examples
40, 42, 45, 50 degrees F.
Can vary with the stage of development
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Phenology
Study of timing of natural events
Ex. bud break in spring
Related to GDD
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# roots
# leaves
Environmental factors:
fruit
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Stages are not
At equal DD
intervals
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Developmental stages
Calendar date is not predictible, because
development is driven by temperature.
Therefore prediction is best based on
temperature dates as a currency.
Stages of development are not equally spaced
based on either calendar date or temperature
units.
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Different Parts of Plants Require Different Number of Growing Degree Days
KNOW THIS
GRAPH
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Vegetative growth: Roots
Governed by temperature, water, and oxygen
4-6°C min; 20-25 optimum; 30-35 max
Temperature threshold is lower than for shoots
In competition with shoots for resources
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The plant only has a
limited amount of
resources to put
towards growth
Within a growing
season, the roots,
shoots, and fruits do
not all grow at the
same time or rate
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Do Roots grow in the winter
Do roots under go endormancy
No, do not need a chilling requirement.
Do roots grow in the winter
Yes but they need heat and
Oxygen
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Shoot Growth Division of cells of the apical meristem
Elongation**
Differentiation
Maturation
Rudimentary leaf
leaf
apical meristem
lateral bud
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REPRODUCTIVE GROWTH
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Flower bud initiation
Timing: varies with species and variety.
Species time of initiation Peach late June-late July
cherry, sweet Early July
cherry, tart mid-July
Apple mid June-early Aug
Pear early July-early Aug
Blackberry late Aug
Strawberry (June) Sept
Strawberry (DN) Early summer
Walnut, pecan Early spring
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CHERRY ANNUAL CYCLE Calendar date basis
MORPHOLOGY Leaves
Leaf abscission
Roots
Trunk
Fruit
Cell division
Pit hardening
Final swell
Abscission
Harvest
FBI
F-Development
Physiology Dormancy
Summer
Rest
Spring
JAN FEB MAR APR MAY JUNE JULY AUG SEPT OCT NOV DEC
A H
Temp, low and high temp, light, water, nutrition, biological temp, nutrition
160-180 days
58-65 days