Invasive plant species on Kosrae, Federated States ofMicronesia

James C Space, Barbara Waterhouse, Julie S. Denslow, Duane Nelson andErick E. Waguk

U.S.D.A. Forest ServicePacific Southwest Research StationInstitute of Pacific Islands Forestry

Honolulu, Hawai'i, USA

22 December 2000

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22 December 2000

Invasive plant species on Kosrae, Federated States of Micronesia

James C Space, Barbara Waterhouse, Julie S. Denslow, Duane Nelson and Erick E. Waguk1

This is a continuation of the survey of islands in Micronesia for invasive plant species requestedby the Pacific Islands Committee, Council of Western State Foresters. A survey of otherMicronesian islands was conducted in 1998 and was discussed in a previous report2. This reportis based on a survey of the island of Kosrae, Federated States of Micronesia, from 24 to 28March 2000. The objectives, as with the previous survey, were three-fold: (1) To identify plantspecies on the islands that are presently causing problems to natural and semi-naturalecosystems; (2) to identify species that, even though they are not presently a major problem,could spread more widely or spread to other islands where they are not present, potentiallycausing problems; and (3) to confirm the absence of species that are a problem elsewhere and, ifintroduced to Kosrae, could be a threat there.

During our visit local experts showed us sites of known infestations. We also had availablecopies of various botanical and weed surveys conducted in the past (see Appendix 1,References). A visit of such short duration does not permit an exhaustive survey of the weedflora of the island. However, the intent was to conduct an overall survey. Additional surveys of

1 Former Director, Pacific Southwest Research Station, USDA Forest Service (now retired); Botanist, AustralianQuarantine and Inspection Service; Research Ecologist/Team Leader, Institute of Pacific Islands Forestry, USDAForest Service; Forest Health Coordinator, Institute of Pacific Islands Forestry, USDA Forest Service and StateForester, Kosrae State, Federated States of Micronesia, respectively.

2 Space, James C. and Marjorie Falanruw (1999). Observations on invasive plant species in Micronesia. Reportprepared for the meeting of the Pacific Islands Committee, Council of Western State Foresters, Majuro, Republic ofthe Marshall Islands, February 22-26, 1999.

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individual species or sensitive areas can and should be conducted as needed. This reportsummarizes our findings and makes some suggestions for further action.

For convenience and to be consistent with the Micronesia report, invasive species occurring in orof threat to Kosrae have been grouped into four categories:

1. Species that are invasive elsewhere in similar ecosystems but were not seen on our visitand are not reported in the literature as being present on Kosrae (352 species).

2. Species that are invasive elsewhere and are also invasive or potentially invasive onKosrae (12 species).

3. Species that are invasive or weedy elsewhere and are cultivated, common or weedy onKosrae (48 species).

4. Native species (or early introductions) that exhibit aggressive behavior (9 species).

These species are listed in Appendix 2. Additional information about each species is located ona World Wide Web site, http://www.hear.org/pier, and on the PIER-CD.

There are a number of other species that are invasive weeds in gardens, fields, and pastures andalong roadsides but don’t seem to pose a particular threat to wildland ecosystems. These speciesare not included.

1. Dangerous species not known to be on Kosrae

Perhaps because of its location “off the beaten path”, many invasive species have yet to reachKosrae. These are listed in Appendix 2, Table 1. The worst of these include the following:

Antigonon leptopus (chain of hearts). This climbing vine has become a widespread pest onGuam and there are isolated plants in cultivation on Pohnpei. Traffic between Guam, Pohnpeiand Kosrae and the fact that this is a pretty flower that people like to plant in their gardens makesthis species a potential threat to Kosrae.

Cecropia obtusifolia and C. peltata, invasive tree species that are a problem in Hawai'i andFrench Polynesia, respectively.

Cinnamomum verum (cinnamon), found to be very invasive in American Samoa and elsewhere.It is present on Pohnpei.

Clidemia hirta (Koster’s curse), a serious problem species in Hawai’i and in other locations. InMicronesia it is only present in Palau, but would be a serious problem if introduced as it growswell in the shade of closed forests.

Coccinia grandis (ivy or scarlet gourd), a smothering vine that is showing potential for seriousdamage to the forests of Saipan. The vines climb over trees and form such dense cover that theforest underneath is completely shaded out and destroyed. It is also invasive in Guam andHawai'i and a horticultural variety is present on Pohnpei. The proximity of the plant on Pohnpeiand Guam poses a threat to Kosrae.

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Two rubber trees, Funtumia elastica (African rubber tree) and Castilla elastica (Panama rubbertree), species that have proven very invasive in Samoa. While not known to be present inMicronesia (Castilla elastica is present in Hawai'i) and thus posing a lesser threat, their invasivenature makes these species prime candidates for prompt eradication if found.

Eichhornia crassipes (water hyacinth), present on Guam and Pohnpei and a threat to Kosrae’srivers and wetlands.

The Hyptis species, H. capitata, H. pectinata and H. suaveolens. Kosrae is presently free of theseinvasive plants. H. capitata is present on Pohnpei, though, and could pose a threat to Kosrae.

Miconia calvescens (the purple plague), which has caused serious damage to the ecosystem ofTahiti and had spread to several other islands in French Polynesia. It has also escaped in Hawaiiand is the subject of an intensive and costly eradication effort there. Recently it was discoveredin Queensland, Australia. It is an attractive garden plant and might be introduced this way or astiny seeds on shoes or used equipment.

A number of grass species, including:

• Melinis minutiflora (molasses grass), a species that is both invasive and causes a serious firehazard. It has fostered the establishment of fire regimes on many islands where it has beenintroduced. It is present on Guam, and is thus a threat to other Micronesian islands.

• Pennisetum setaceum (fountain grass), another grass that is a major problem in Hawai’i. It isalso present on Guam.

• Imperata cylindrica (cogon grass), known as a very invasive species and reported to bepresent on Guam.

• A number of other grass species of various degrees of invasiveness, including Andropogongayanus, Andropogon glomeratus, Andropogon virginicus, Brachiaria mutica, Brachiariasubquadripara, Cenchrus brownii, Cenchrus ciliaris, Chloris barbata, Cynodon dactylon,Digitaria insularis, Digitaria violascens, Echinochloa polystachya, Hymenachne amplex-icaulis, Hyparrhenia rufa, Ischaemum rugosum, Ischaemum timorense, Panicum maximum,Panicum repens, Paspalum dilatatum, Paspalum fimbriatum, Paspalum paniculatum,Paspalum urvillei, Pennisetum clandestinum, Rhynchelytrum [Melinis] repens,Schizachyrium condensatum, Setaria pallide-fusca, Setaria palmifolia and Sorghumhalepense. Grasses are easily introduced as contaminants in imported seed, imported sandand gravel or on used machinery.

Mimosa invisa (giant sensitive plant) and Mimosa pudica (sensitive plant) are both absent fromKosrae but present on Pohnpei and Guam. Mimosa invisa is a particularly nasty plant as it iscovered with thorns and forms dense tangles that are difficult to walk through. Mimosa pudica, asmaller plant with only small prickles, can form dense mats. Kosrae is at high risk for invasionby both these plants.

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Momordica charantia (bitter-melon), a member of the cucumber family, is a climbing vine andits fruit can be the host for fruit flies. It is present on Pohnpei, Chuuk and Guam.

Passiflora foetida is present on Kosrae, but several invasive members of this invasive genus areabsent, including:

• Passiflora mollissima (banana poka), a smothering vine that is a problem in Hawai'i and NewZealand.

• Passiflora suberosa (wild passionfruit, devil's pumpkin or indigo berry).

• Passiflora laurifolia, Passiflora ligularis, Passiflora maliformis, Passiflora quadrangularisand Passiflora rubra. Passiflora quadrangularis is present on Pohnpei.

Piper auritum (eared pepper, locally called “false sakau”), recently introduced to Pohnpei. Thisspecies suckers profusely and produces many small seeds that are spread by birds, rodents andbats. An attempt is being made to eradicate it on Pohnpei. It was introduced into Pohnpei as anew form of sakau, but is worthless in this regard.

Pithecellobium dulce (Madras thorn), on all the other major Micronesian islands but apparentlynot yet present on Kosrae.

Psidium cattleianum (strawberry guava), a small tree that forms dense thickets, is a majorproblem species in Hawai'i, Tahiti and elsewhere. It is present on Pohnpei and thus is a seriousthreat to Kosrae.

Spathodea campanulata (African tulip tree), a commonly planted ornamental present onPohnpei, Chuuk and Guam. This tree has become a major problem in Fiji, the Hawaiian Islandsand some other places. It was not seen on Kosrae and it would be good to exclude it from theisland.

Tibouchina herbacea (glorybush or cane ti), another species that is a major problem in Hawai'i.

The best indicator that a species might be invasive is the fact that it is invasive elsewhere.However, each island ecosystem is unique, and invasiveness cannot be predicted with certainty.A good strategy is to be extremely cautious and exclude these and other species known to beinvasive or weedy elsewhere (although an even better strategy, now being adopted by countriessuch as Australia and New Zealand, is to exclude all species not shown by risk analysis to be ofnegligible risk). Known problem species that have the potential to cause problems in tropicalisland ecosystems and are not yet present on Kosrae are listed in Appendix 2, Table 1. Thesespecies should be excluded through plant quarantine and, if establishment is detected, promptlyevaluated for eradication. In addition, species that are present in Guam, Pohnpei and Hawai'i butare not present on Kosrae are listed in Appendix 3. There is high risk of introduction of thesespecies because of air and ship traffic between these points and Kosrae.

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2. Species that are invasive elsewhere and are likewise invasive or have the potential tobecome so on Kosrae

Some known invasive plants that are causing trouble in similar ecosystems have been introducedinto Kosrae (Appendix 2, Table 2). Some of them are already causing problems while others arenot. Some are cultivated plants that have not (yet) escaped and their potential for causingdamage is so far unknown. However, one of the best predictors of invasiveness is the behaviorof the species elsewhere, and these are known troublemakers.

Adenanthera pavonina (the coral bean tree), invasive in secondary forests and alreadywidespread on Kosrae.

Chromolaena odorata (Siam weed), a highly invasive pan-tropical weed. It will likely becomeeven more widespread over time. Biological controls are available that are effective in openareas, less so in shaded stands. On Kosrae it is likely to follow new developments such as roads.

Clerodendrum quadriloculare is suspicious because it appears to have the ability to invade intactor relatively intact native forests. This species is notorious for being a prolific producer of rootsuckers and in fact the plant is easily propagated by means of root cuttings. A dense, mono-specific understory of this species was seen growing in full shade beneath the forest canopy onPohnpei, making it a likely candidate to invade intact or only slightly disturbed native foreststands. Specimens observed on Kosrae were cultivated ones in yards, but this species should bemonitored for possible naturalization. If so, naturalized populations should be promptlyeradicated.

Dieffenbachia maculata (spotted Dieffenbachia or dumb cane) appears to be mostly anornamental at present on Kosrae. However, this is potentially a very bothersome species, as itreproduces vegetatively and can thrive in the dense shade of an intact native forest canopy.

Hedychium coronarium (white ginger) can be an invader of swampy areas and wet forests andappears to have escaped from cultivation on Kosrae.

Mikania micrantha (mile-a-minute weed) is apparently a fairly recent introduction, as it is notyet widespread. It is very invasive in some locations in the Pacific and should be evaluated forpossible eradication or control.

Occasional trees of Paraserianthes falcataria (Moluccca albizia) were seen. This species has thepotential, over time, of spreading widely through the forest (as can be seen on Pohnpei). Furtherplanting should be discouraged. If this is not a desired species, emphasis could be given tocutting the existing trees.

Syngonium podophyllum, a climbing aroid, is probably another escape from cultivation. LikeDieffenbachia maculata, it has the ability to spread in the deep shade of intact forests. It isdifficult to eradicate by hand as it is able to reproduce from a single node and bits and pieces ofthe stems or roots are easily overlooked.

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3. Species that are mentioned or listed as weedy or invasive elsewhere and arecultivated, common or weedy on Kosrae

A large number of other cultivated, common or weedy introduced species were noted. Many ofthese species, which might best be termed aggressive weeds, are mostly prevalent alongroadsides or on disturbed sites, although some species, particularly alien trees, can graduallyspread into forested ecosystems. In the case of vines and plants that form dense ground cover,the regeneration of native species can be inhibited.

Some of these species could become a problem in the future, since there is often a long lag timebetween introduction and when a species begins to cause serious impacts. These species (listedin Appendix 2, Table 3) should be monitored for spread and possible control, if necessary.

Potentially invasive tree species include Acacia auriculiformis (earleaf acacia), Bauhiniamonandra (orchid tree), Ceiba pentandra (kapok), Moringa oleifera (horseradish tree), Samaneasaman (monkeypod), and Tecoma stans (yellow bells).

Dissotis rotundifolia (dissotis) is quite prevalent in shaded areas and waterways.

A number of introduced grasses have become established, including Bothriochloa bladhii,Cenchrus echinatus (bur grass); Chloris radiata (plush-grass, radiate fingergrass), Digitariaciliaris (fingergrass, smooth crabgrass); Eleusine indica (goose grass), Paspalum conjugatum(Hilo grass), Pennisetum polystachyon (mission grass) and Pennisetum purpureum (elephant ornapier grass). Pennisetum polystachyon may be a new introduction as it was only seen at theairport and near the Catholic church.

Other widespread weedy species include Blechum brownei (blackweed) and Desmanthusvirgatus, possibly a recent introduction.

Passiflora foetida (love-in-a-mist) is common but not overly aggressive.

Desmodium tortuosum (Spanish clover or Florida beggar weed) was noted at four sites.

Stachytarpheta jamaicensis was widespread but Stachytarpheta urticifolia (blue rat's tail) hasapparently not yet arrived—be on the lookout for it.

Wedelia trilobata (Singapore daisy) is a widespread pest, as on most Pacific islands.

4. Native species (or early introductions) exhibiting aggressive behavior

Some native species (or early introductions) exhibit aggressive characteristics. These species arelisted in Appendix 2, Table 4.

Merremia peltata (pul) is quite invasive any place there is disturbance.

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Rubus moluccanus (kohkihl) has a very bad reputation for invasiveness on tropical islands(Mauritius, La Reunion) but is considered native on Kosrae (Fosberg, 1979). It is widespreadbut does not seem to be causing major problems.

Strategies for dealing with invasive species

The first line of defense against invasive species is to keep them out. Control at ports of entry isessential, and land management officials should work closely with plant protection and quarantineofficials to make them aware of known and potential invasive plant species. Plant quarantineofficers are familiar with most agricultural pests, but they may not be aware of some of the peststhat threaten wildland ecosystems. In cooperation with the plant quarantine organization, a list ofnoxious species to be excluded should be developed and exclusion of these species should bebacked by the force of law and regulation. Better yet is to go to the approach recently adopted byAustralia and under serious consideration by a number of other countries. That is to exclude allalien species not shown to be of negligible risk. Risk assessment and management techniques canbe used to assess the likelihood and effects of possible introductions and to develop exclusion anderadication strategies.

Education of the public about the danger of introductions and encouraging the use of nativespecies can be helpful. People need to understand why they should follow the quarantineregulations, why they shouldn’t dump garden cuttings in the woods, and why they should reportsuspicious plants. What may just be a pretty flower to be planted in a yard or garden can turn outto be an invasive species. Education of schoolchildren is especially important, as this is the mostimpressionable age. Children can also have a notable effect on the actions of their parents.

The public also needs to understand that the immediate eradication of a small area of a problemspecies, even if it involves the use of herbicides, may be better than living with a problemspecies forever. There are many instances where you hear people say "I wish we'd taken actionwhen this pest was first noticed". It may even be worthwhile to take people to a place wherethey can be shown the full extent of a problem if the infestation is allowed to spread so that theywill accept and support eradication. For example, anyone visiting Tahiti would very likely comeaway convinced that Miconia calvescens is an ecological disaster and that it should be preventedfrom becoming established on other Pacific islands.

The public should also be encouraged to report suspicious plants. Funding can be requested toprepare information folders or "wanted" posters. Prompt follow-up to public reports is essentialto maintain credibility.

It is particularly important to work with any local plant importers, as these are often sources ofnew introductions. A positive approach is to work together to develop a “white list” of bothnative and non-native species that the public can be encouraged to plant.

Foresters, conservation officers, extension agents and others that spend time in the field shouldbe alert to new species that exhibit invasive behavior. Often, these species first show up in urbanor farm areas and tend to first become established in gardens and disturbed areas. Suspiciousplant species should be promptly reported. Periodically scheduled surveys can also be conductedfor new or expanding infestations. An evaluation should be conducted for any new species that

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appears to be invasive or is known to be invasive elsewhere. Assistance by an expert who isfamiliar with the species and methods for its eradication or control should be requested if needed.Prompt action is essential, since once a species becomes widespread, control or eradication canbe extremely costly or impossible. Assistance is also available on-line from experts through thePacific Pestnet and Aliens list-servers

Laws and regulations should be reviewed to make sure they are adequate to deal with newintroductions. This includes the ability of government to require the control of noxious specieson private lands or to take action on private lands if the landowner cannot be located or does nottake prompt action. Provision for emergency funds to deal with immediate problems should alsobe in place. Model laws and regulations can be obtained from states and countries that haveimplemented them.

The Federated States of Micronesia and Kosrae State governments are encouraged to takeadvantage of the Federal assistance programs in dealing with invasive plant species. Cost-sharefunding is available to provide locally available expertise in forest health protection. Experts arealso on call from the Forest Service’s Forest Health Protection staff, or they can arrange forexpert consultation. Finally, cost-share funding can be requested to deal with forest healthproblems, including plant pests, under the Cooperative Forestry and Hawaii Tropical ForestryRecovery acts. Funding is subject to recommendations resulting from a professional evaluationof the problem and the overall availability of funds. Funding also tends to be prioritized basedon an economic evaluation of cost-effectiveness, so rapid eradication of a pest species while it isstill confined to a small area would undoubtedly take precedence over the chronic problem of aspecies that has escaped control.

Recommendations

In addition to the above strategies, we offer the following specific recommendations:

• Make every effort to keep out all the species listed in Appendix 2, Table 1, with specialemphasis on those listed in Appendix 3 (those species present in Guam, Pohnpei and Hawai’ibut not yet on Kosrae). All of these are known invasive species elsewhere, and there is nosense in running the risk that they will act the same on Kosrae. As we are able to gathermore information on other species that might threaten tropical island ecosystems we will addthem to the list. For those with Internet access, additional information and new listings canbe found at http://www.hear.org/pier. The information is also available on CD for local use.Assistance from experts in identifying and managing invasive species can also be obtainedby subscribing to and posting inquiries on the Aliens and Pacific Pestnet list servers.

• Take special measures to keep Antigonon leptopus, Castilla elastica, Cecropiaobtusifolia/peltata, Cinnamomum verum, Clidemia hirta, Coccinia grandis, Eichhorniacrassipes, Funtumia elastica, Miconia calvescens, Psidium cattleianum and Spathodeacampanulata out, to monitor for their occurrence, and to eradicate them immediately iffound. These are all well-documented problem species that have had a major impact onnatural ecosystems elsewhere. The risk from these species is very high.

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Since there is so much traffic to Kosrae from Guam and Hawai'i, where many of thesespecies are present, it would be highly desirable to institute special measures to preventintroduction of invasive species from these sources. At a minimum, quarantine officersshould be alert to people who might have been in the woods or rural areas in Guam orHawai'i, especially on the Big Island of Hawai'i, and inspect their shoes or boots for seeds.Any trucks or equipment coming from Hawai'i that have been used in rural areas must bepower washed or steam cleaned. Warning posters should be produced to alert the public andencourage reporting of any introductions. Any infestations picked up from public reportingor scouting should be promptly eradicated before the plants set seed.

• Closely monitor the situation in regard to Piper auritum, the recently introduced pepper(“false sakau”) on Pohnpei. If this species is not eradicated, it will definitely be a threat tothe ecosystems of adjacent islands. It should not be planted on Chuuk and promptlyeradicated if found.

• Species listed in Appendix 2, Table 3 are pests and, although they may not be presentlycausing serious damage to wildland ecosystems, they are certainly not desirable species.They should not be more widely distributed.

• A number of species used in forestry and ornamental plantings are, at least to some degree,invasive. While many of these species have desirable ornamental or physical characteristics,planting exotics as opposed to native species is a policy question that needs to be carefullyconsidered. Most species that naturalize to any extent will gradually spread throughoutavailable and suitable habitat, given enough time.

• Require steam cleaning or power washing of all used equipment coming into Kosrae. This isdesirable not only from the standpoint of excluding invasive plants, but also insects anddiseases. Especially suspect are roadbuilding machinery, military equipment and off-roadvehicles. Equipment that has been in Hawai'i is of especially high risk because of the dangerof introducing Miconia calvescens and some other especially noxious species prevalent there.Guam, Saipan and Tinian are of only slightly lower risk, as many invasive species are presentthere that are not on Kosrae.

• Change the customs arrival form to have people indicate if they have been in forests as wellas farms. This would then alert quarantine officers to inspect boots and other clothing forsoil and seeds. New Zealand uses this language on its entry forms.

• Intact native forests are the most resistant to invasion. Any measures that limit the amount ofdisturbance will help keep invasive species out.

• A risk-rating scheme should be developed and the various known and potential invasivespecies evaluated for their risk of introduction, spread and potential damage. The Institute ofPacific Islands Forestry is looking at adapting the Australian risk assessment system to theneeds of the Pacific.

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• A joint training session/workshop should be conducted for foresters, land managers,extension agents, quarantine officers and other interested individuals and organizations in therecognition, exclusion, eradication, and control of invasive species. The goal of the trainingand workshop should be to foster cooperation and joint action against invasive species. Weare looking into cooperating with the Secretariat of the Pacific Community to provide suchtraining through their quarantine assistance program.

• The purpose of this survey was to give an overall assessment of the situation. Individualspecies of concern should be more fully evaluated as needed as to extent, invasiveness, andthe possibility for control or eradication. Technical assistance should be requested, if needed,to evaluate individual species.

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Appendix 1.

Background material and references:

Cole, T. G., C. D. Whitesell, W. A. Whistler, N. McKay and A. H. Ambacher. 1988. Vegetation survey and forestinventory, Kosrae. Pac. SW Forest and Range Expt. Stn. Resource Bull. PSW-25. USDA Forest Service. 14 pp. plusmap.

Cronk, Q. C. B., and J. L. Fuller. 1995. Plant invaders. Chapman and Hall. 241 pp.

Csurhes, S. and R. Edwards. 1998. Potential environmental weeds in Australia: Candidate species for preventativecontrol. Canberra, Australia. Biodiversity Group, Environment Australia. 208 pp.

D’Antonio, C. M., and P. M. Vitousek. 1992. Biological invasions by exotic grasses, the grass-fire cycle, and globalchange. Ann. Rev. Ecol. And System. 23:63-87.

Falanurw, M. C., T. G. Cole and A. H. Ambacher. 1989. Vegetation survey of Rota, Tinian, and Saipan,Commonwealth of the Northern Mariana Islands. Pac. SW Forest and Range Expt. Stn. Resource Bulletin PSW-27.11 pp. plus map.

Falanruw, M. C., T. G. Cole, A. H. Ambacher, K. E. McDuffie and J. E. Maka. 1987. Vegetation survey of Moen,Dublon, Fefan and Esten, State of Truk, Federated States of Micronesia. Pac. SW Forest and Range Expt. Stn.Resource Bull. PSW-20. USDA Forest Service. 11 pp.

Falanuruw, M. C., J. E. Maka, T. G. Cole and C. D. Whitesell. 1990. Common and scientific names of trees andshrubs of Mariana, Caroline, and Marshall Islands. Pac. SW Forest and Range Expt. Stn. Resource Bulletin PSW-67. 91 pp.

Fosberg, F. R., M. V. C. Falanruw, and M.-H. Sachet. 1975. Vascular flora of the northern Mariana Islands.Smithsonian Contrib. Bot. 22:1-45.

Fosberg, F. R., M. V. C. Falanruw, and M.-H. Sachet. 1977. Additional records of vascular plants from thenorthern Mariana Islands. Micronesica 13: 27-31.

Fosberg, F. R., M. V. C. Falanruw, and M.-H. Sachet. 1980. Additional records of vascular plants from thenorthern Mariana Islands. Micronesica 16: 211-214.

Fosberg, F. R., and M.-H. Sachet. 1975. Flora of Micronesia. Part 1. Gymnospermae. Smithsonian Contrib. Bot.20: 1-15.

Fosberg, F. R., and M.-H. Sachet. 1975. Flora of Micronesia. Part 2. Casuarinaceae, Piperaceae, and Myricaceae.Smithsonian Contrib. Bot. 24: 1-28.

Fosberg, F. R., and M.-H. Sachet. 1977. Flora of Micronesia. Part 3. Convolvulaceae. Smithsonian Contrib. Bot.36: 1-34.

Fosberg, F. R., and M.-H. Sachet. 1980. Flora of Micronesia. Part 4. Caprifoliaceae, Compositae. SmithsonianContrib. Bot. 45: 1-40.

Fosberg, F. R., M.-H. Sachet and R. L. Oliver. 1993. Flora of Micronesia. Part 5. Bignoniaceae--Rubiaceae.Smithsonian Contrib. Bot. 81: 1-135.

Fosberg, F. R., M.-H. Sachet and R. L. Oliver. 1979. A geographical checklist of the Micronesian dicotyledonae.Micronesica 15:1-295.

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Fosberg, F. R., M.-H. Sachet and R. L. Oliver. 1987. A geographical checklist of the Micronesianmonocotyledonae. Micronesica 20:1-126.

Hafliger, E. and H. Scholz. 1980. Grass Weeds. CIBA-GEIBY Ltd., Basle, Switzerland. Two volumes.

Hafliger, E. 1980. Monocot Weeds. CIBA-GEIBY Ltd., Basle, Switzerland. 132 pp. plus plates.

Holm, L. G., D. L. Plucknett, J.V. Pancho, and J.P. Herberger. 1977. The world’s worst weeds: Distribution andEcology. East-West Center/University Press of Hawaii. 609 pp.

McLean, C. D., C. D. Whitesell, T. G. Cole, and K. E. McDuffie. 1988. Timber resources of Kosrae, Pohnpei,Truk, and Yap, Federated States of Micronesia. Pac. SW Forest & Range Expt. Sta. Resource Bull. PSW-24.USDA Forest Service. 8 pp.

Merlin, M., and J. Juvik. 1996. Plants and their environments in Chuuk (Ira me neeniier non Chuuk). Honolulu,Hawaii: East-West Center. 121 pp.

Merlin, M., R. Taulung, and J. Juvik. 1993. Plants and environments of Kosrae (Sahk kap ac kain in can Kosrae).Honolulu, Hawaii: East-West Center. 113 pp.

Meyer, J-Y. 2000. Preliminary review of the invasive plants in the Pacific islands (SPREP Member Countries) in:Sherley, G. (tech. ed.). Invasive species in the Pacific: A technical review and draft regional strategy. South PacificRegional Environment Programme, Samoa. pp. 85-114.

Mueller-Dombois, D., and F. R. Fosberg. 1998. Vegetation of the tropical Pacific Islands. New York, Springer-Verlag. 733 pp.

Neal, M. C. 1965. In Gardens of Hawaii. Bernice P. Bishop Museum Special Publication 50, Bishop Museum Press.924 pp.

Parsons, W. T. and E. G. Cuthbertson. 1992. Noxious weeds of Australia. Inkata Press, Melbourne/Sydney. 692 pp.

Raulerson, L. and A. Rinehart. 1991. Trees and shrubs of the Northern Mariana Islands. Commonwealth of theNorthern Mariana Islands. 120 pp.

Smith, A. C. 1979-1991. Flora Vitiensis Nova: A New Flora of Fiji. Lawai, Kauai, Hawaii. National TropicalBotanical Garden. Five Volumes.

Stone, B. C. 1970. The Flora of Guam. Micronesica 6:1-659.

Stone, C.P. , C. W. Smith and J. T. Tunison (eds) 1992. Alien Plant Invasions in Native Ecosystems on Hawaii:Management and Research. University of Hawaii, Cooperative National Park Research Studies Unit, Honolulu.University of Hawaii Press.

Swarbrick, J. T. 1997. Weeds of the Pacific Islands. Technical paper No. 209. South Pacific Commission, Noumea,New Caledonia. 124 pp.

Wagner, W. L., D. R. Herbst, and S. H. Sohmer. 1990. Manaual of the flowering plants of Hawai'i. University ofHawaii Press & Bishop Museum Press, Honolulu. 1854 pp., 2 vols.

Waterhouse, B. M., and A. A. Mitchell. 1998. Northern Australia quarantine strategy weeds target list. AustralianQuarantine & Inspection Service, Miscellaneous Publication No. 6/98. 110 pp.

Waterhouse, D. F. 1993. Biological control: Pacific prospects. Supplement 2. Australian Centre for InternationalAgricultural Research, Canberra. 138 pp.

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Waterhouse, D. F. 1994. Biological control of weeds: Southeast Asian prospects. Australian Centre for InternationalAgricultural Research, Canberra. 302 pp.

Waterhouse, D. F. 1997. The major invertebrate pests and weeds of agriculture and plantation forestry in theSouthern and Western Pacific. The Australian Centre for International Agricultural Research, Canberra.. 69 pp.

Waterhouse, D. F. and K. R. Norris. 1987. Biological control: Pacific prospects. Inkata Press, Melbourne. 454 pp.

Whistler, W. A. 1995. Wayside Plants of the Islands. Isle Botanica, Honolulu. 202 pp.

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Appendix 2

Table 1. Species that are invasive elsewhere in similar ecosystems but are not known to bepresent in Kosrae

Scientific Name Common Names Family Habit

Acacia confusaEnglish: Formosa koa, Formosa acacia; Other: sosigi, shoshigi, sosugi,boiffuring (Guam and CNMI–Chamorro); serepa, soschghi (CNMI–Carolinian); pilampwoia (Pohnpei); ianángi, yanangi (Palau)

Fabaceae tree

Acacia farnesianaEnglish: Ellington curse, klu, sweet acacia; French: cassie; Other:aroma, popinac, kandaroma (Guam, CNMI); kolu (Hawai'i); vaivai vaka-vatona, vaivai vakavotona, ban baburi, oki (Fiji); te kaibakoa (Kiribati);debena (Nauru) (Swarbrick, 1997)

Fabaceae shrub

Acacia longifolia English: Sydney golden wattle, golden wattle, long-leaved wattle, long-leaved acacia, sallow wattle, coast wattle, golden rods Fabaceae tree

Acacia mangium English: mangium; Other: tuhkehn pwelmwahu (Pohnpei) Fabaceae treeAcacia mearnsii English: black wattle Fabaceae treeAcaciamelanoxylon English: Australian blackwood, blackwood acacia Fabaceae tree

Acacia nilotica English: prickly acacia, Egyptian thorn, Nile acacia, algaroba, babul,gum arabic tree; Other: tiare (New Caledonia) Fabaceae tree

Acacia spp. English: acacia Fabaceae shrub/treeAdenantheraabrosperma English: giddy giddy (Australia) Fabaceae tree

Agave sisalana English: sisal, sisal hemp; Other: ndali, natali (Fiji) Agavaceae herbAilanthus altissima English: tree of heaven Simaroubaceae treeAlbizia chinensis English: Chinese albizia Fabaceae tree

Albizia lebbeck

English: siris-tree, rain tree, East Indian walnut, kokko, woman's-tonguetree, soros-tree, raom tree ; French: bois noir; Other: trongkon-mames,tronkon mames, mamis (Guam); kalaskas, trongkon-kalaskas (CNMI–Chamorro); schepil kalaskas (CNMI–Carolinian); ukall ra ngebard(Palau); gumorningabchey; ngumorrningobchey (Yap); 'ohai (Hawai'i);vaivai, vaivai ni vavalangi, vaivai ni vavalagi (Fiji); tamaligi pa’epa’e(Samoa); kasia (Tonga)

Fabaceae tree

Albizia lophantha English: plume albizia, plume acacia, stink bean, brush wattle Fabaceae shrub/tree

Aleuritesmoluccana

English: candlenut, Indian walnut; French: bancoulier; Other: lumbang(Guam), raguar (Caroline Islands); sakan, shakan (Pohnpei); lama(American Samoa and Samoa); kukui (Hawaii); tuitui (Niue, Tonga),'ama (Marquesas); lauthe, lauthi, toto, sikethi, tuitui, waiwai, sikeli,nggerenggere (Fiji); tahii tiairi, ti’a’iri, tutui, tahiri (French Polynesia).

Euphorbiaceae tree

Alpinia purpurata English: red ginger; Other: thevunga (Fiji); ‘awapuhi ‘ula’ula (Hawai'i) Zingiberaceae herb

Alstoniamacrophylla English: devil-tree Apocynaceae tree

Alternantheraphiloxeroides English: alligator weed Amaranthaceae herb

Anacardiumoccidentale

English: cashew, cashew nut; French: anacardier, pomme de cojou,acajou a pommes; Other: kasoi (Guam), 'apu initia (Samoa); kesiu(Tonga)

Anacardiaceae tree

Andropogongayanus English: gamba grass Poaceae grass

Andropogonglomeratus English: bush beardgrass, bushy bluestem Poaceae grass

Andropogonvirginicus English: broomsedge, yellow bluestem, whisky grass Poaceae grass

Angiopteris evecta English: king’s fern; Other: bersarm, bersarm dermarm (Palau), payuit,umpai (Pohnpei); mong, mongmong (Yap) Marattiaceae fern

Annona glabra English: pond apple, alligator apple, bullock's heart, cherimoyer; Other:uto ni mbulumakau, uto ni bulumakau, kaitambo, kaitambu (Fiji). Annonaceae tree

Anredera cordifolia English: Madeira vine, mignonette vine, lamb's tails; Other: filikafa (Niue) Basellaceae vine

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Scientific Name Common Names Family Habit

Antigononleptopus

English: Mexican creeper, mountain rose, Confederate vine chain-of-love, hearts on chain, love-vine, coral vine, queen’s jewels, kadena deamor, corallita; Other: rohsapoak (Pohnpei)

Polygonaceae vine

Araujia sericifera English: moth plant, moth catcher, white bladder flower Asclepiadaceae shrub

Ardisia crenata English: hen's eyes, Hilo holly, coral berry, Australian holly,scratchthroat; French: arbre à noël Myrsinaceae shrub

Ardisia elliptica English: Shoebutton ardisia;Other: ati popa'a, atiu (French Polynesia) Myrsinaceae shrub

Argyreia nervosaEnglish: elephant creeper, Hawaiian baby woodrose , silver morningglory, woolly morning glory; French: coup d'air, liane a minguet, liane d'argent

Convolvulaceae vine

Aristolochialittoralis English: Dutchman's pipe, calico flower Aristolochiaceae vine

Arthrostemaciliatum English: everblooming eavender Melastomataceae herb

Arundo donax English: giant reed, Spanish reed, wild cane; Other: ngasau nivavalangi (Fiji) Poaceae grass

Asclepiasphysocarpa English: balloon plant Asclepiadaceae shrub

Asparagusdensiflorus English: asparagus fern, sprengeri fern Liliaceae herb

Asparagussetaceus English: ornamental asparagus, climbing asparagus fern Liliaceae vine

Austroeupatoriuminulaefolium Asteraceae herb

Axonopuscompressus English: carpetgrass Poaceae grass

Axonopusfissifolius English: narrow-leaved carpetgrass Poaceae grass

Azadirachta indica English: neem, nim, margosa Meliaceae treeBarleria lupulina English: hophead Philippine violet Acanthaceae shrubBarleria prionitis English: barleria, porcupine flower Acanthaceae shrubBauhinia purpurea English: purple orchid tree; purple butterfly tree, pink butterfly tree (Fiji) Fabaceae treeBauhinia variegata English: orchid tree; mountain ebony, butterfly tree (Fiji) Fabaceae tree

Bidens pilosa

English: beggar’s tick, Spanish needle, cobbler's pegs; French: piquantsnoirs; Other: fisi'uli (Tonga); kofe tonga, kofetoga (Niue); piripiri (CookIslands); tae puaka (Futuna); batimadramadra, mbatimandramandra,mbatikalawau, matakaro, matua kamate (Fiji); ki, ki nehe, ki pipili, nehe(Hawai'i)

Asteraceae herb

Bischofia javanicaEnglish: bischofia, bishopwood; Other: 'o'a (American Samoa); toog(Hawai'i); koka (Cook Islands, Hawai'i, Fiji, Tonga, Futuna); tongotongo,koka ndamu, tongo, tongatonga, tea (Fiji)

Euphorbiaceae tree

Blainvillea gayana Asteraceae herbBocconiafrutescens English: bocconia Papaveraceae shrub

Boerhaviacoccinea

English: boerhavia, hog weed, hog feed, red spiderling, wineflower, tarvine; Other: mata pavo, patagon Nyctaginaceae herb

Brachiaria mutica English: California grass, para grass, buffalo grass, Mauritius grass,signal grass; Other: puakatau (Tonga) (Swarbrick, 1997) Poaceae grass

Brachiariasubquadripara English: brachiaria, green summer grass, tropical signalgrass Poaceae grass

Brillantaisialamium Acanthaceae shrub

Buddleia asiatica English: dog tail Loganiaceae shrubBuddleia davidii English: orange eye butterfly bush, summer lilac Loganiaceae shrubBuddleiamadagascariensis English: butterfly bush, smoke bush Loganiaceae shrub

Caesalpiniadecapetala

English: cats claw, Mysore thorn, Mauritius thorn, wait-a-bit; Other:puakelekino (Hawai'i) Fabaceae shrub

Caesalpinia majorEnglish:; yellow nickers; Other: pakao (Guam); 'anaoso (AmericanSamoa); kakalaioa, hihikolo, kinikini (Hawai'i); talamoa, talamoa foto(Niue)

Fabaceae vine

16

Scientific Name Common Names Family Habit

Calamus spp. English: rattan; Other: rotan, bejuko-n-halumtano, rotan bejuro-n-halumtan (Guam); bangerenguis ra ngebard (Palau) Arecaceae palm

Calliandracalothyrsus

English: powderpuffOther: kaliana (Samoa) Fabaceae shrub

Calliandrahaematocephala English: red powder puff Fabaceae shrub

Calliandrasurinamensis English: Surinamese stickpea Fabaceae tree

Callisia fragrans English: fragrant inch plant, basketplant, spironema Commelinaceae herbCalopogoniummucunoides English: calopo; Other: akankan-guakag (Guam) Fabaceae vine

Calotropis procera English: calotropis, rubber bush, apple of Sodom, mudar, madar, king’scrown, roostertree Asclepiadaceae shrub/tree

Cardiospermumgrandiflorum English: balloon vine; heart seed Sapindaceae vine

Cardiospermumhalicacabum

English: balloon vine, heart pea, love-in-a-puff; Other: wa niu, vo niu(Fiji); vinivinio (French Polynesia) Sapindaceae vine

Carpobrotus edulis English: ice plant, pigface, hottentot fig Aizoaceae succulent

Castilla elastica English: Panama rubber tree, Mexican rubber tree; Other: pulu mamoe(American Samoa) Moraceae tree

Casuarinaequisetifolia

English: casuarina, ironwood, Australian pine, she-oak, horsetail tree;French: arbe de fer; Other: nach (Yap); lach (Ulithi); laash (Woleai);weeku (Chuuk); gagu, gago (Guam, CNMI); weighu (CNMI-Carolinian);pinetree, nidil, bluegum (Marshall Islands); toa (American Samoa,French Polynesia, Samoa and Tonga) ‘aito (French Polynesia),nokonoko, te katurina (from Ecoport, source not given)

Casuarinaceae tree

Casuarina glauca English: swamp oak, saltmarsh ironwood, longleaf ironwood, graysheoak Casuarinaceae tree

Cecropiaobtusifolia English: trumpet tree, guarumo Cecropiaceae tree

Cecropia peltata English: trumpet tree; French: parasolier, faux-ricin, pisse-roux Cecropiaceae tree

Cedrela odorata English: cigar box cedar, Mexican cedar, West Indian cedar, Spanishcedar, Barbados cedar; Other: sita hina (Tonga) Meliaceae tree

Cenchrus brownii English: burgrass, burr grass, sand-bur, slimbristle sandbur Poaceae grassCenchrus ciliaris English: buffelgrass, African foxtail grass, anjan grass Poaceae grassCentrosemapubescens English: centro; Other: pi ni ndola (Fiji) Fabaceae herb

Cestrum diurnumEnglish: inkberry, day jessamine, day cestrum, China berry; Other: tinta‘n-China, tentanchinu, tintan China (Guam, CNMI); thauthau (Fiji), vaitohi(Tonga).

Solanaceae shrub

Cestrumnocturnum

English: night-flowering cestrum, night-flowering jasmine, queen (or lady)of the night; Other: dama-de-noche (Guam); juñoul rua awa [?] (MarshallIslands); iki he po (Niue); teine o le po, ali'i o le po (American Samoaand Samoa); lakau po'uli (Tonga); thauthau, thauthau ni mbongi, kara(Fiji)

Solanaceae shrub

Cestrum parqui English: green cestrum, willow-leaved jessamine, Chilean cestrum,green poison-berry Solanaceae shrub

Chamaecristanictitans

English: partridge pea, Japanese tea senna; Other: kobo-cha, nemu-cha(Japan) Fabaceae small shrub

Chloris barbata English: swollen fingergrass, purpletop chloris (Australia), airport grass(Fiji); Other: mau’u lei (Hawaii) Poaceae grass

Chrysobalanusicaco English: coco plum, icaco; Other: apolo (Fiji); icaque Chrysobalanaceae shrub

Chrysophyllumoliviforme English: satin leaf, caimitillo Sapotaceae tree

Cinchonapubescens English: quinine tree Rubiaceae tree

Cinnamomumburmannii English: padang cassia Lauraceae tree

Cinnamomumcamphora English: camphor tree, camphor laurel Lauraceae tree

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Scientific Name Common Names Family HabitCinnamomumverum

English: cinnamon tree; Other: ochod ra ngebard (Palau). tinamoni(American Samoa and Samoa) Lauraceae tree

Cirsium vulgare English: spear thistle, Scotch thistle, bull thistle, black thistle, plumethistle; Other: pua kali (Hawai'i) Asteraceae herb

Cissus nodosa English: grape ivy Vitaceae lianaCitharexylumcaudatum English: juniper berry Verbenaceae tree

Citharexylumspinosum English: fiddlewood; Other: masese (Fiji) Verbenaceae tree

Clausena excavata English: clausena Rutaceae treeClerodendrumbuchananii var.fallax

English: red clerodendrum, pagoda-flower; Other: talufe (Niue);amo’ula, amo’ule (Tonga) Verbenaceae shrub

Clerodendrumjaponicum English: glorybower Verbenaceae shrub

Clerodendrumpaniculatum

English: pagoda plant, pagoda flower; Other: butcherechár,butecherechar (Palau); tukehn sousou (Pohnpei) Verbenaceae shrub

Clidemia hirtaEnglish: Koster’s curse, soap bush; Other: kui, kúi (Palau); kauresinga,kaurasinga, roinisinga, ndraunisinga, mara na bulumakau, mbona nambulamakau, vuti (Fiji); kaurasiga (from FAO, no source given)

Melastomataceae shrub

Clitoria ternateaEnglish: butterfly pea, Asian pigeonwings; Other: buikike, bukike,paokeke, bukike paokeke, capa de la reina, kapa de la raina (Guam);putitainubia (CNMI); pepe (Niue); latoela, nawa (Fiji), paipa (Tonga)

Fabaceae vine

Clusea rosea English: signature tree, autograph tree, copey, Scotch attorney,pitchapple Guttiferae tree

Coccinia grandis English: ivy gourd, scarlet-fruited gourd; Other: aipikohr (Pohnpei);kundru (Fiji) Cucurbitaceae vine

Coffea arabica English: coffee; Other: kove, kofe (Fiji), kofi (Tonga) Rubiaceae treeConocarpuserectus English: sea mulberry, buttonwood, button mangrove Combretaceae shrub/tree

Cordia alliodora English: laurel, Ecuador laurel, salmwood, Spanish elm; Other: kotia(Samoa, Tonga) Boraginaceae tree

Cordia curassavica English: black sage Boraginaceae shrubCordia glabra English: broad-leaved cordia Boraginaceae treeCortaderia jubata English: jubata grass, Andean pampas grass, purple pampas grass Poaceae grassCortaderia selloana English: pampas grass, silver pampas grass, Uruguayan pampas grass Poaceae grassCorynocarpuslaevigatus English: New Zealand laurel, karakara nut Corynocarpaceae tree

Crassocephalumcrepidoides

English: thickhead, fireweed, redflower ragleaf; Other: fua lele, pualele(American Samoa and Samoa); maraburubo (Solomon Islands); fisipuna (Tonga); se vuka (Fiji)

Asteraceae herb

Crotalariaanagyroides English: crotalaria; Other: pine kotalelia, pile Fabaceae shrub

Cryptostegiagrandiflora

English: rubber vine, India rubber vine, Palay rubbervine; Other: lianede gatope (New Caledonia) Asclepiadaceae vine

Cryptostegiamadagascariensis English: Madagascar rubbervine Asclepiadaceae vine

Cupaniopsisanacardioides English: carrotwood Sapindaceae tree

Cuscutacampestris

English: golden dodder, field dodder, five-angled dodder; Other:navereverelangi, wa vereverelangi, wa lawala, wa mbosuthu, wandanga, wa tikaivu, ndithangi (Fiji)

Convolvulaceae vine

Cynodon dactylon

English: Bermuda grass, giant Bermuda grass, bahama grass, devil'sgrass, couch grass, Indian doab, grama, devilgrass, couchgrass, balamagrass; French: chiendent, petit chiendent, chiendent pied-de-poule;Other: manini, manienie (Hawaii); motie molulu (Niue); kambuta, kabuta(Fiji); mosie molulu (Niue); herbe de couverture (New Caledonia); pastobermuda, zacate bermuda, grama dulce, gramón, hierba fina, grama-seda, (from Ecoport, no source given)

Poaceae grass

Cyperusalternifolius English: umbrella sedge, umbrella plant, dwarf papyrus grass Cyperaceae sedge

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Scientific Name Common Names Family Habit

Cyperus rotundus

English: nut grass, nutsedge, purple nutsedge, cocograssFrench: souchet rond, souchet à tubercules, herbe à oignon; Other:chaguan humatag (Guam); tuteoneon (Marshall Islands); kili'o'opu(Hawaii); oni ani (Cook Islands); soro na kambani, sora na kambani,soro ni kabani, ivako, malanga, vucesa, motha, vuthesa mot ha (Fiji);mumuta (Samoa, Tokelau); pakopako (Tonga), te mumute (Kiribati)

Cyperaceae sedge

Dalbergia sissoo English: Indian dalbergia, Indian rosewood, shisham Fabaceae treeDerris indica English: pongam, pongame oiltree Fabaceae tree

Derris malaccensis English: New Guinea creeper; Other: 'ava niukini (American Samoa);nduva, tuva, nduva ni niukini (Fiji); akau niukini (Niue); kava fisi (Tonga). Fabaceae vine

Desmodiumnicaraguense

Spanish: engorda-caballo, vara de arco, pie de paloma, vara larga, varablanca, Juana de Arco, gutao, Juana Larga Fabaceae shrub

Dichrostachyscinerea

English: aroma, Kalahari Christmas tree, Chinese lantern tree, sicklebush, marabou-thorn, marabú, Sen Domeng, Saint Domingue, acaciaSaint Domingue

Fabaceae shrub

Digitaria insularis English: cottongrass, sourgrass, feather-top grass Poaceae grassDigitariaviolascens English: smooth crabgrass, violet crabgrass; Other: sau (Palau) Poaceae grass

Dilleniasuffruticosa English: shrubby dillenia Dilleniaceae shrub/tree

Duranta erecta English: duranta, golden dewdrop, pigeon-berry, sky-flower; Other:‘olive (Tonga) Verbenaceae shrub

Echinochloapolystachya English: aleman grass, carib grass, creeping rivergrass Poaceae grass

Egeria densa English: egeria, dense waterweed Hydrocharitaceae aquaticEhrharta stipoides English: meadow ricegrass Poaceae grass

Eichhorniacrassipes

English: water hyacinth; French: jacinthe d'eau; Other: bung el ralm(Palau); mbekambekairanga, ndambendambe ni nga, jal khumbe,bekabekairaga, dabedabe ne ga, jal khumbe (Fiji); riri vai (Cook Islands);jacinto de agua, lirio acuatico

Pontederiaceae aquaticherb

Elaeagnusumbellata English: autumn olive, oleaster, silverberry Elaeagnaceae shrub

Elaeis guineensis English: African oil palm; Other: apwiraiasi (Pohnpei) Arecaceae tree

Elephantopusmollis

English: elephantopus, elephant’s foot, tobacco weed; French: fauxtabac; Other: papago' vaca, papago' halom tano, papago' baka, papago'halomtano' (Guam); lata hina, lau veveli (Tonga); tavako ni veikau, janglitambaku (Fiji); tapua erepani (Cook Islands)

Asteraceae herb

Erigeronkarvinskianus English: Mexican daisy, seaside daisy, daisy fleabane Asteraceae herb

Eriobotrya japonica English: loquot, Japanese plum; Other: loketi (Tonga) Rosaceae tree

Eugenia uniflora English: Surinam cherry, red Brazil cherry; French: cerisier de Cayenne;Other: kafika, kafika palangi (Niue), pitanga (Brazil) Myrtaceae shrub, tree

Eupatoriumcatarium English: praxelis Asteraceae herb

Ficus altissima English: lofty fig, false banyan, council tree Moraceae treeFicus benghalensis English: banyan, Indian banyan, vada tree Moraceae treeFicus benjamina English: weeping fig, baka; Other: ovava' fisi (Tonga) Moraceae tree

Ficus elastica English: India rubber tree, rubber plant; Other: komunoki, komunokí(Palau); rapah (Pohnpei); gak'iynigoma (Yap) Moraceae tree

Ficus macrophylla English: Moreton Bay fig Moraceae treeFicus nota English: tibig Moraceae treeFicus rubiginosa English: rusty-leaved fig, Port Jackson fig Moraceae tree

Flacourtia rukam English: Indian plum, Indian prune, governor's plum; Other: filimoto (Fiji,Futuna, Samoa and American Samoa, Tonga) Flacourtiacae tree

Flemingiamacrophylla Fabaceae tree

Flemingiastrobilifera

English: luck plant; French: sainfoin du bengale; Other: besungelaiei(Palau) Fabaceae shrub

Flindersiabrayleyana English: Queensland maple, silkwood Rutaceae tree

Fuchsia boliviana English: fuchsia Onagraceae tree

19

Scientific Name Common Names Family HabitFuchsiamagellanica English: hardy fuchsia, earring flower; Other: kulapepeiao (Hawai'i) Onagraceae shrub

Funtumia elastica English: African rubber tree, silkrubber; Other: pulu vao (Samoa) Apocynaceae tree

Furcraea cubensis English: Cuban hemp, hempOther: cubuya (Galapagos Islands) Agavaceae succulent

Furcraea foetida English: Mauritius hemp, sisal; Other: toua (Niue); faumalila, fau malila(Tonga) Agavaceae succulent

Grevillea banksii English: kahili flower, Banks grevillea; Other: haiku (Hawai'i) Proteaceae treeGrevillea robusta English: silk oak, silky oak, she-oak, silver oak; Other: oke' (Tonga) Proteaceae treeGrewia asiatica English: phalsa Tiliaceae treeGymnocoronisspilanthoides English: Senegal tea, temple plant Asteraceae herb

Haematoxylumcampechianum English: logwood, bloodwood tree, campeachy wood, campeche Fabaceae tree

Harunganamadagascariensis English: harungana Clusiaceae tree

Hedychiumflavescens

English: yellow ginger; Other: awaphuhi melemele (Hawai'i), opuhi rearea (French Polynesia) Zingiberaceae herb

Hedychiumgardnerianum English: kahili ginger; Other: cevuga dromodromo (Fiji); sinter weitahta Zingiberaceae herb

Heliocarpuspopayanensis English: moho, white moho Tiliaceae tree

Hemigraphisalternata English: metal leaf; red ivy, cemetery plant (Fiji) Acanthaceae herb

Heterocentronsubtriplinervium English: pearl flower Melastomataceae herb

Heterospathe elataOther: palma brava, palma braba (Guam); asbo, demailei, demailéi,domaile, domailei, ebouch (Palau); buag bbuag (Yap); Sagisi palm(Philippines?)

Arecaceae tree

Hiptagebenghalensis English: hiptage; French: liane de cerf Malpighiaceae shrub

Hydrilla verticillata English: hydrilla, Florida elodea, water thyme Hydrocharitaceae herbHylocereusundatus

English: night-blooming cereus, strawberry pear, red pitaya, dragon fruit,pitahaya, pitaya; Other: paniniokapunahou, papipi pua (Hawai'i) Cactaceae climber

Hymenachneamplexicaulis English: hymenachne, trompetilla, West Indian marsh grass Poaceae grass

Hyparrhenia rufa English: jaragua grass, thatching grass Poaceae grassHypochoerisradicata English: hairy cat’s ear, gosmore Asteraceae herb

Hyptis capitata English: knobweed; Other: botones, batunes (Guam); t'aiegarabao,pathpath (Yap), maa uupo (French Polynesia) Lamiaceae herb

Hyptis pectinataEnglish: comb hyptis, comb bushmint, mint weed, purple top; French:fausse menthe; Other: mumutun lahe, mumutun palaoan, mumutanademelon (Guam); tamole ni veikau, tamoli ni vavalangi, timothi nivavalangi, wavuwavu, ndamoli, ben tulsia (Fiji)

Lamiaceae herb

Hyptis suaveolens English: wild spikenard; Other: mumutun (Guam) Lamiaceae herbImpatienswalleriana

English: garden impatiens, balsam, busy lizzy, bizzy-lizzie, patient lucy;Japanese balsam (Fiji) Balsaminaceae succulent

Imperata cylindricaEnglish: blady grass, cogon grass, satintail; French: paille de dys,paillotte, impérata cylindrique; Other: gi, ngi (Fiji); alang-alang, lalang,carrizo, kunai

Poaceae grass

Indigoferasuffruticosa

English: indigo; Other: aniles (Guam); ‘iniko, inikoa, kolu (Hawai'i);'akauveli (Tonga). Fabaceae shrub

Ischaemumrugosum

English: muraina grass, wrinkle duck-beak, saramattagrass; Other: thomuraina, co muraina (Fiji) Poaceae grass

Ischaemumtimorense English: centipede grass, stalkleaf murainagrass; Other: waidoi grass Poaceae grass

Jacarandamimosifolia English: jacaranda Bignoniaceae tree

Jasminumfluminense English: Brazilian jasmine Oleaceae vine

20

Scientific Name Common Names Family Habit

Jatropha curcasEnglish: physic nut, purging nut; Other: wiriwiri, wiriwiri ni vavalangi, utoni vavalangi, banidakai, mbanindakai, manggele, maqele, ndrala, fiki(Fiji); tuba-tuba (Guam)

Euphorbiaceae shrub

Jatrophagossypifolia English: bellyache bush, cotton-leaved physic nut Euphorbiaceae shrub

Justicia carnea English: pink plume-flower, pink jacobinia Acanthaceae shrub

Kalanchoe pinnataEnglish: life plant, air plant, resurrection plant, Canterbury bells,cathedral bells, Mexican love plant; Other: kibilia (Marshall Islands);bulatawamudu (Fiji); tupu he lau, tupu noa (Niue); teang (Kiribati)

Crassulaceae herb

Khayasenegalensis English: African mahogany, Senegal mahogany Meliaceae tree

Kyllinga polyphylla English: Navua sedge; Other: tuise tele, tuise fiti (Samoa), pakopako(Tonga) Cyperaceae sedge

Lemna spp. English: duckweed Lemnaceae herbLeonotisnepetifolia English: lion’s tail, lion’s ear, Christmas candlestick Lamiaceae herb

Leptospermumericoides English: tree manuka, tree manuba; kanuka (New Zealand) Myrtaceae tree

Leptospermumscoparium English: New Zealand tea; manuka (New Zealand) Myrtaceae tree

Licuala grandis English: ruffled fan palm; French: palmier-cuillère Arecaceae tree

Ligustrum spp. English: privet Oleaceae shrubs,small trees

Lonicera japonica English: Japanese honeysuckle, Hall’s honeysuckle; Other: honekakala(Hawai'i) Caprifoliaceae vine

Lophostemonconfertus English: brush box, Brisbane box, vinegar tree Myrtaceae tree

Ludwigia peruviana English: ludwigia, Peruvian primrose, water primrose Onagraceae shrubMacfadyenaunguis-cati English: cat's-claw climber Bignoniaceae vine

Macroptiliumatropurpureum English: siratro, purple bushbean Fabaceae herb

Maesopsis eminii English: umbrella-tree, musizi Rhamnaceae tree

Medinilla magnifica English: chandelier tree, showy melastome, showy medinilla, Malaysianorchid Melastomataceae shrub

Medinilla venosa English: holdtight Melastomataceae shrubMelaleucaquinquenervia

English: melaleuca, cajeput, paper bark tree, punk tree; Other: niaouli(New Caledonia) Myrtaceae tree

Melastomacandidum English: Asian melastome, Indian rhododendron, Malabar melastome Melastomataceae tree

Melastomasanguineum English: fox-tongued melastoma, red melastome Melastomataceae shrub/tree

Melia azedarachEnglish: Chinaberry, pride-of-India, indian lilac, Persian lilac, whitecedar, tira; French: lilas des Indes; Other: paraiso, para'isu (Guam);lelah (Pohnpei); prais (Yap);sili, tili (Niue); dake, bakain (Fiji), sita(Tonga)

Meliaceae tree

Melicoccusbijugatus English: Spanish lime Sapindaceae tree

Melinis minutiflora English: melinis, molasses grass; French: herbe molasses; Other:puakatau (Tonga) Poaceae grass

Melochia umbellata English: melochia Sterculiaceae treeMemecylonfloribundum Melastomataceae shrub

Merremia tuberosa English: wood rose, Spanish arborvine, yellow morning-glory; Other:pilikai (Hawaii) Convolvulaceae vine

Miconia calvescens English: miconia, velvet leaf, purple plague, bush currant Melastomataceae treeMikania scandens English: climbing hempweed, climbing hempvine Asteraceae vine

Mimosa invisa

English: giant sensitive plant, nila grass; French: grande sensitive,sensitive gèante; Other: singbiguin sasa (Saipan); mechiuaiu (Palau);limemeihr laud (Pohnpei); vao fefe palagi (American Samoa andSamoa), la'au fefe tele, la'au fefe palagi (Samoa); wa ngandrongandrolevu, wa ngandrongandro ni wa ngalelevu, co gadrogadro (Fiji)

Fabaceae shrub

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Scientific Name Common Names Family Habit

Mimosa pigraEnglish: catclaw mimosa, thorny sensitiveplant, giant mimosa, giantsensitive plant, black mimosa, zaraz, dormilona, bashful plant, amouretteviolet; French: amourette riviére

Fabaceae shrub

Mimosa pudica

English: sensitive plant, sleeping grass; French: sensitive; Other:betguen sosa (Guam); memege (Niue); mechiuaiu (Palau); limemeihr(Pohnpei); ra kau pikikaa (Cook Islands); vao fefe (American Samoa andSamoa); mateloi (Tonga); tho ngandrongandro, tho kandrodandro,cogadrogadro (Fiji); pohe ha'avare, pope haavare (Tahiti); tuitui, lajwania(from Ecoport, source not given)

Fabaceae herb

Momordicacharantia

English: balsam-apple, bitter-melon, bitter gourd, balsam pear, squirtingcucumber, cerasee, peria; Other: almagosa, atmagosu (Guam);atmagoso (Guam, CNMI); markoso (Palau); kerala (Fiji); meleni 'aekuma, vaine 'initia (Tonga)

Cucurbitaceae vine

Montanoahibiscifolia English: montanoa, tree daisy, Anzac flower Asteraceae shrub

Muntingia calaburaEnglish: jam tree, strawberry tree, Jamaican cherry, Singapore cherry,Panama cherry, Panama berry, ornamental cherry, calabura, sirsen;Other: budo (Yap, Palau); manzanilla, mansanita, manzanita (Guam,CNMI), capulin (Fiji, French Polynesia)

Tiliaceae tree

Murraya paniculata English: orange jessamine, satin-wood, Chinese box Rutaceae shrub/treeMyriophyllumaquaticum English: parrot's feather, water feather, Brazilian water milfoil Haloragidaceae aquatic

herbNeonotonia wightii English: glycine Fabaceae vineNymphaea spp. English: water lily Nymphaeaceae herbOchna serrulata English: ochna, Mickey-Mouse plant, bird’s-eye bush Ochnaceae shrubOchna thomasiana English: Micky Mouse plant, bird’s-eye bush Ochnaceae shrub/treeOchromapyramidale English: balsa, corkwood, down tree Bombacaceae tree

Odontonematubaeforme English: fire spike, cardinal flower Acanthaceae shrub

Olea europaea English: oliveOther: ‘oliwa, ‘oliwa haole (Hawai'i) Oleaceae tree

Operculinaventricosa

English: paper rose, St. Thomas lidpod; Other: alalag (Guam); palulu(Samoa); fue hina [?] (Tonga) Convolvulaceae vine

Opuntia spp. English: prickly pear; Other: lengua de vaca (Guam) Cactaceae succulentshrub

Ossaea marginata MelastomataceaeOxysporapaniculata English: oxyspora, bristletips Melastomataceae shrub

Paederia foetida English: stink vine; Other: maile pilau, maile ka kahiki (Hawai'i). Rubiaceae vine

Panicum maximum English: Guinea grass, green panic, buffalograss; French: herbe deGuinéa, panic élevé, capime guiné, fataque; Other: saafa (Tonga) Poaceae grass

Panicum repens English: torpedo grass, couch panicum, creeping panic; French: panicrampant; Other: wainaku grass (Hawaii) Poaceae grass

Parkinsoniaaculeata English: parkinsonia, Jerusalem thorn, horse-bean, retama Fabaceae tree

Paspalumdilatatum

English: paspalum, dallis grass, water grass; French: paspalum dilaté,herbe sirop, herbe de miel; Other: hiku nua (Niue) Poaceae grass

Paspalumfimbriatum

English: fimbriate paspalum, winged paspalum, Panama paspalum,Panama crowngrass, Columbia grass Poaceae grass

Paspalumorbiculare

English: rice grass; ditch millet (Fiji)Other: co duru levu, tho nduru levu, tho ndina, tho ni ndina (Fiji); karasi(Solomon Islands)

Poaceae grass

Paspalumpaniculatum English: Russell river grass, galmarra grass Poaceae grass

Paspalum urvillei English: Vasey grass Poaceae grassPassiflora alata English: wild passion fruit Passifloraceae shrubPassiflora caerulea English: blue passion flower Passifloraceae shrubPassiflora coccinea English: scarlet passion flower, Red Passion-flower, Red Granadilla Passifloraceae shrub

Passiflora edulisEnglish: passion fruit, yellow passion fruit, purple passion fruit, qarandila,purple granadilla; Other: liliko’i (Hawaii), pompom (Pohnpei); vaine tonga(Tonga); pasio (Samoa)

Passifloraceae vine

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Scientific Name Common Names Family Habit

Passiflora laurifoliaEnglish: yellow granadilla, belle apple, yellow water-lemon; French:pomme-liane; Other: pasio (Samoa); vaine tinetina, vaine ‘ae kuma(Tonga)

Passifloraceae vine

Passiflora ligularis English: sweet granadilla, yellow passionfruit; French: barbadine Passifloraceae vinePassifloramaliformis

English: hard-shelled passionfruit, sweet calabash, sweet cup; French:pomme d'or; Other: pasio (Samoa), vaine kai (Tonga) Passifloraceae vine

Passifloramollissima English: banana poka, banana passionfruit, bananadilla Passifloraceae vine

Passiflorapulchella English: two-lobed passion flower, wingleaf passionfruit Passifloraceae shrub

Passifloraquadrangularis

English: granadilla, giant granadilla; Other: kudamono (Palau);parapotina maata (Cook Islands); palasini, palatini, vine fua lalahi, vinepalasini, tinitini (Niue); pasione (Tonga); pasio (Samoa); para pautini(French Polynesia)

Passifloraceae vine

Passiflora rubra English: red passionfruit, Dutchman's laudanum Passifloraceae vine

Passiflorasuberosa

English: wild passionfruit, devil's pumpkin, indigo berry, corkypassionflower, corkstem passionflower; French: passiflore, grenadille;Other: huehue haole (Hawai'i)

Passifloraceae vine

Paulowniatomentosa English: paulownia, princess-tree, Chinese empress tree, karri-tree Scrophulariaceae tree

Pennisetumclandestinum English: kikuyu grass, kikuyo Poaceae grass

Pennisetumsetaceum English: fountain grass Poaceae grass

Persea americanaEnglish: avocado, alligator pear; Other: apoka (Cook Islands); pea (Fiji);avoka (Niue), avoka (Tonga); aviota (Samoa); bata (Palau); alageta(Chamorro, Guam)

Lauraceae tree

Philadelphuskarwinskyanus English: mock orange Saxifragaceae shrub

Phormium tenax English: New Zealand flax, New Zealand hemp, bush flax Agavaceae herbPhylostachys nigra English: black bamboo Poaceae shrubPimenta dioica English: pimento, allspice Myrtaceae treePimenta racemosa English: bay tree, bay rum tree, bay oil tree Myrtaceae tree

Pinus caribaea English: Caribbean pine, Bahamas pineOther: paina (Samoa) Pinaceae tree

Piper aduncum English: spiked pepper; Other: yaqona ni Onolulu, yanggona ni Onolulu(Fiji) Piperaceae tree

Piper auritumEnglish: eared pepper, anise piper, Veracruz pepper; Spanish: hojasanta, anisillo, hinojo, sabalero, hoja de la estrella; Other: Hawaiiansakau, false sakau, false kava (Pohnpei)

Piperaceae shrub

Pistia stratiotesEnglish: water lettuce, tropical duckweed; French: laitue d'eau, pistie;Other: lechuguita de agua, repollo de agua , apon-apon, apoe-apoe,beo-cai, chawk

Araceae aquaticherb

Pithecellobiumdulce

English: Madras thorn, Manila tamarind, monkeypod; Other: kamachuri(Chuuk); camachili, kamachile, kamachiles, kamachili (Guam);kamachili, kamatire (CNMI Chamorro), ghamasiligh (CNMI-Carolinian);kamatsiri, kamatsíri, opiuma (Palau): 'opiuma (Hawaii); kataiya (Fiji)

Fabaceae tree

Pittosporumpentandrum English: pittosporum, Taiwanese cheesewood Pittosporaceae shrub/tree

Pittosporum tobira English: tobira, Japanese pittosporum, Japanese cheesewood, mockorange, Australian laurel Pittosporaceae shrub/tree

Pittosporumundulatum

English: Victorian box, Victorian laurel, Australian cheesewood, mockorange, sweet pittosporum Pittosporaceae tree

Platyceriumbifurcatum English: elkhorn fern, staghorn fern Selaginellaceae fern

Plectranthusamboinicus

English: Mexican mint, Spanish thyme, Cuban oregano; Other: rhaivoki,sage (Fiji); pasiole (Niue, Tonga) Lamiaceae herb

Plectranthusscutellarioides

English: coleus; Other: lata, lau lata (Fiji); weleweka (Hawai'i); selevese(Niue); koramahd, koaramahd (Pohnpei); pate, patiale (Samoa) Lamiaceae herb

Pluchea indica English: Indian fleabane, Indian pluchea, Indian camphorweed Asteraceae shrubPlucheasymphytifolia English: sour bush Asteraceae shrub

Prosopis spp. English: mesquite, algaroba Fabaceae tree

23

Scientific Name Common Names Family Habit

Psidiumcattleianum

English: strawberry guava, cherry guava, Cattley guava, Chinese guava;French: goyavier de Chine; Other: kuahpa (Pohnpei); waiawi (Hawai'i);ngguava (Fiji), tuava tinito (French Polynesia)

Myrtaceae tree

Psidium guineense English: Brazilian guava, Guinea guava Myrtaceae shrub/tree

Pueraria lobataEnglish: kudzu, Japanese arrowroot; Other: deday (Yap); aka(American Samoa, Tonga, Niue, Wallis and Futuna); a'a (AmericanSamoa and Samoa); yaka , wa yaka, nggariaka (Fiji); akataha,fue’aepuaka (Tonga); acha, nepalem

Fabaceae vine

Puerariaphaseoloides English: tropical kudzu, puero Fabaceae vine

Pyrostegia venusta English: flame vine, flame flower, golden shower, orange trumpet vine Bignoniaceae vineQuisqualis indica English: Rangoon creeper Combretaceae vineRavenalamadagascariensis

English: traveler’s palm, traveler’s tree, ravenale; French: arbre àvoyageur Strelitziaceae tree

Rhodomyrtustomentosa English: rose myrtle, downy myrtle, Isenberg bush Myrtaceae shrub

Rhynchelytrumrepens

English: Natal redtop, Natal grass, red Natal grass, Holme's grass,blanketgrass; French: herbe du Natal, herbe rose, herbe pappangue;tricholène (New Caledonia); Other: salapona (Tonga)

Poaceae grass

Ricinus communisEnglish: castor bean, castor-oil plant; French: ricin; Other: agaliya(Guam); gelug, maskerekur, uluchula skoki (Palau); tuitui, tuitui fua ikiiki(Niue); koli (Hawaii); lama palagi (American Samoa and Samoa); lepo,lepohina (Tonga); mbele ni vavalagi, toto ni vavalagi, utouto (Fiji).

Euphorbiaceae shrub

Rivina humilis English: baby pepper, bloodberry, coral berry, rouge plant; Other: polo(Tonga) Phytolaccaceae herb

Rubus alceifolius English: giant bramble Rosaceae shrub

Rubus rosifolius English: roseleaf raspberry, thimbleberry, Mauritius raspberry; French:framboisier; Other: ola'a, ‘akala, akalakala (Hawai'i) Rosaceae shrub

Rubus spp. English: raspberries, blackberries, brambles Rosaceae shrubRuellia prostrata English: black weed, bell weed; Other: vao uli, vao uliuli (Samoa) Acanthaceae herbSalvinia molesta English: salvinia, water fern, kariba weed, African payal Salviniaceae fernSambucusmexicana English: elderberry, Mexican elder Caprifoliaceae small tree

Sanchezia nobilis English: sanchezia. Acanthaceae shrubSancheziaparvibracteata English: Sanchezia Acanthaceae shrub

Sansevieratrifasciata

English: bowstring hemp, mother-in-law’s tongue; Other: tigre (Guam),kitelel (Palau) Agavaceae herb

Sapium sebiferum English: Chinese tallow tree, popcorn tree, vegetable tallow, white waxberry, chicken tree, Florida aspen Euphorbiaceae tree

Scheffleraactinophylla English: octopus tree, umbrella tree, ivy palm Araliaceae tree

Scheffleraarboricola English: dwarf brassia, dwarf schefflera, Hawaiian elf schefflera Araliaceae shrub

Schinus molle English: pepper tree, California pepper tree, Peruvian peppertree,pepperina Anacardiaceae tree

Schinusterebinthifolius

English: Christmas-berry, Brazilian pepper, Brazilian holly, Florida holly;French: faux poivrier, poivre rose; Other: warui (Fiji) Anacardiaceae tree

Schizachyriumcondensatum English: little bluestem, bush beardgrass Poaceae grass

Seneciomadagascarensis English: fireweed Asteraceae herb

Seneciomikanioides English: German ivy, Italian ivy, cape ivy Asteraceae vine

Senna obtusifolia English: habucha, coffeeweed, sicklepod Fabaceae herbSennaseptemtrionalis

English: buttercup bush, yellow shower; Other: mosimosi, nasenikarakarawa, winivinikau (Fiji); kolomona, kalamona (Hawai'i) Fabaceae shrub/tree

Senna siamea English: cassod tree, kassod tree, Siamese cassia Fabaceae tree

Sesbania puniceaEnglish: red sesbania, coffee weed, rattle-pod, sesban, red seine bean,Brazil rattlebox, coffee of the coast, tame acacia, false poinciana, blackacacia, purple sesbane

Fabaceae shrub

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Scientific Name Common Names Family HabitSetaria pallide-fusca

English: foxtail, garden bristle grass, yellow bristlegrass, Queenslandpigeon grass (Australia), cat's tail grass (Fiji) Poaceae grass

Setaria palmifolia English: palmgrass, short pitpit, hailans pitpit, broadleaved bristlegrass Poaceae grassSolandra maxima English: golden cup, cup of gold Solanaceae lianaSolanummauritianum

English: bugweed, wild tobacco, tree tobacco; Other: pua nana honua(Hawai'i), pula (Tonga) Solanaceae shrub

Solanumseaforthianum English: potato creeper, Brazilian nightshade Solanaceae vine

Solanumtampicense English: wetland nightshade, aquatic soda apple Solanaceae aquatic

herbSolanum viarum English: tropical soda apple Solanaceae herb?

Sorghumhalepense

English: Johnson grass, Aleppo grass, Aleppo milletgrassFrench: sorgho d' Alep, sorgo de Alepo, herbe de CubaOther: kola (Tonga); gumai (Russia); zacate Johnson, grama China,cañuela, Don Carlos

Poaceae grass

Sorghumsudanense English: Sudan grass; Other: kola (Tonga) Poaceae grass

Spathodeacampanulata

English: African tulip tree, fireball, fountain tree; French: tulipier duGabon, pisse-pisse; Other: apär (CNMI); rarningobchey (Yap); tuhkedulip (Pohnpei); tiulipe (Tonga), taga mimi (Fiji)

Bignoniaceae tree

Sphaeropteriscooperi English: Australian tree fern Cyatheaceae fern

Sporoboluselongatus English: wiregrass, rat-tail dropseed Poaceae grass

Sporobolus indicus English: smutgrass, wiregrass, Indian dropseed Poaceae grass

Stachytarphetaurticifolia

English: blue rat's tail, dark-blue snakeweed, false verbena, nettleleafvelvetberry; French: herbe bleue; Other: ouchung, sakura (Chuuk),louch beluu (Palau); mautofu tala, mautofu vao, matofu fualanumanoa(American Samoa and Samoa); te uti (Kiribati); mautofu Samoa, motofuSamoa (Niue); hiku `i kuma, hiku'kuma, 'iku 'i kuma (Tonga); turulakaka,tumbutumbu, serakawa, lavenia, se karakarawa (Fiji)

Verbenaceae herb

Stipa cernua English: nodding tussockgrass Poaceae grassStizolobiumpruriens

English: cow itch, velvet bean, Bengal bean, Mauritius beanOther: akangkang dangkulo (Guam) Fabaceae vine

Stylosanthesguianensis English: stylo, Brazilian lucerne, tropical lucerne, Nigerian stylo Fabaceae herb

Syncarpiaglomulifera English: turpentine tree, lustre wood Myrtaceae tree

Syzygium cuminiEnglish: Java plum; jambolan plum; French: faux-pistachier, jamelon-guier; Other: duhat (Guam); mesegerak, mesekerrak, mesekerrák,mesigerak (Palau); jamelonguier (New Caledonia); kavika ni India,jammun (Fiji)

Myrtaceae tree

Syzygium jambosEnglish: malabar plum, rose apple, Malay apple; French: jambosier,pommier rose; Other: iouen wai, youenwai, apel en wai (Pohnpei);kavika, kavika ni vavalangi, kavika ni India (Fiji), ahi'a popa'a (FrenchPolynesia), fekika papalangi (Tonga)

Myrtaceae tree

Tabebuiaheterophylla

English: pink tecoma, pink trumpet tree, white cedar; Other: calice dupape Bignoniaceae tree

Tetrazygia bicolor English: Florida clover ash Melastomataceae shrub/treeThunbergia alata English: black-eyed susan vine; Other: tagamimi (Samoa) Acanthaceae vineThunbergiafragrans English: white lady Acanthaceae vine

Thunbergiagrandiflora English: Bengal trumpet, blue trumpet vine, clock vine, skyvine Acanthaceae vine

Thunbergialaurifolia English: purple allamanda, laurel-leaved thunbergia, laurel clock vine Acanthaceae vine

Tibouchinaherbacea English: glorybush, cane ti, tibouchina Melastomataceae herb

Tibouchinaurvileana English: glorybush, lasiandra, princess flower Melastomataceae herb

Tillandsiausneoides

English: Spanish moss, old man's beard, grandfather’s whiskers, airplant; French: cheveaux du roi Bromeliaceae bromeliad

Timonius timon English: liberal (Palau); Other: sakosia (from Ecoport, no source given) Rubiaceae tree

25

Scientific Name Common Names Family HabitTithoniadiversifolia

English: tree marigold, shrub sunflower, Mexican sunflower, Japanesesunflower; Other: matala (Niue) Asteraceae shrub

Toona ciliata English: Australian red cedar; Other: tuna (Samoa) Meliaceae treeTradescanthiazebrina English: wandering zebrina, wandering jew, inchplant Commelinaceae herb

Trema orientalisEnglish: charcoal tree, gunpowder tree; Other: agaunai, banahl, talamama (CNMI); elodechoel (Palau); uanin (Yap); ndrou, ndroundrou,ndrikanaithembe (Fiji); mangele (Niue); bulasisi (from Ecoport, no sourcegiven)

Ulmaceae tree

Tribulus cistoides English: puncture vine, caltrop, burrnut, Jamaican feverplant; Other: temaukinikini (Kiribati) Zygophyllaceae herb

Tribulus terrestris English: puncture vine, goat head Zygophyllaceae herb

Triphasia trifolia English: limeberry, lemon ChinaOther: limon-China, lemon de china, limoncito (Guam) Rutaceae shrub

Triplarisweigeltiana English: long john, long jack Polygonaceae tree

Tripsacumlatifolium English: wideleaf gamagrass Poaceae grass

Triumfettarhomboidea

English: Chinese burr, paroquet burr, burr bush; Other: dadangsi,masiksik lahe (Guam); mo'osipo (Tonga); mosipo (Niue); mautofu(American Samoa and Samoa); qatima (Fiji), urio (French Polynesia)

Tiliaceae shrub

Triumfettasemitriloba English: Sacramento bur; Other: dadangsi, masiksik lahe (Guam) Tiliaceae shrub

Turbina corymbosa English: turbina, Christmas vine Convolvulaceae vineVitex parviflora English: smallflower chastetree; Other: molauin, molave (Philippines) Verbenaceae shrub/treeWaterhousiafloribunda English: weeping lilly pilly, weeping myrtle Myrtaceae tree

Zantedeschiaaethiopica English: calla lily, arum lily Araceae herb

Ziziphusmauritiana

English: Indian jujube, Chinese date, Chinee apple; Other: manzanas,manzanita, jujube (Guam); baer, bahir, baher (Fiji) Rhamnaceae tree

NOTE: The common names in this and the following tables are from the PIER database and web site and includenames in use at various locations throughout the Pacific. Additional information, including the locations where thevarious names are used, can be found in the individual species summaries at the PIER web site,http://www.hear.org/pier, or on the PIER CD.

26

Table 2. Introduced species that are invasive elsewhere and are also invasive or potentiallyinvasive in Kosrae.

Scientific Name Common Names Family Habit

Adenanthera pavonina

English: Coral bean tree, red sandalwood tree, red bead tree, lopa, bead tree,false wiliwili, peacock flower-fence; Other: kaikes (Pohnpei); mwetkwem(Kosrae); colales, culalis, kolales, kulales, kulalis (Guam, CNMI); metekam,metkam, metkem, (Kosrae); telengtúngd, telentundalel (Palau); pomea (Fiji,Niue), lera, lere ndamu, vaivai, vaivai ni vavalangi (Fiji), lopa (American Samoa,Samoa and Tonga); paina, pitipitio (French Polynesia).

Fabaceae tree

Chromolaena odorataEnglish: Siam weed, triffid weed , bitter bush, Jack in the bush; French: herbedu Laos; Other: kesengesil, masigsig (Guam); otuot (Chuuk),wisolmatenrehwei (Pohnpei); mahsrihsrihk (Kosrae); hagonoy, agonoi,huluhagonoi (Philippines).

Asteraceae herb

Clerodendrum chinense English: Honolulu rose, stickbush; Other: losa Honolulu (American Samoa andSamoa); pikake hohono, pikake wauke (Hawai'i), pitate mama (Cook Islands) Verbenaceae shrub

Clerodendrumquadriloculare

English: bronze-leaved clerodendrum; Other: tuhkehn palau (Pohnpei),bagauac (Philippines) Verbenaceae shrub

Dieffenbachia maculata English: dieffenbachia, dumb cane; Other: yalu ni vavalagi (Fiji) Araceae herb

Lantana camara

English: lantana; Other: landana, rantana, rahndana, tukasuweth (Pohnpei); rosfonacni (Kosrae); kauboica, kaumboitha, mbonambulumakau, mbona rambulumakau, tokalau, waiwai, lanitana (Fiji); latana (American Samoa andSamoa); latora moa, tatara moa (Tahiti); lakana (Hawai'i); talatala, talatalatalmoa (Tonga); te kaibuaka, te kaibuaja (Kiribati); taramoa (Cook Islands);migiroa (Nauru), taratara hamoa (French Polynesia)

Verbenaceae shrub

Leucaena leucocephala

English: leucaena, wild tamarind, lead tree; French: faux-acacia; faux mimosa(New Caledonia); Other: koa haole (Hawaii); tangantangan, tangan-tangan,talantayan (Guam, CNMI, Marshall Islands); talntangan (CNMI); ganitnityuwantangantan (Yap); tuhngantuhngan, rohbohtin (Kosrae); telentund (Palau); lopasamoa (American Samoa); fua pepe (American Samoa and Samoa); lusina(Samoa); pepe (Niue and Samoa); nito (Cook Islands); siale mohemohe(Tonga); vaivai, vaivai ni vavalangi, vaivai dina, balori (Fiji); cassis (Vanuatu); tekaitetua (Kiribati); namas (from Ecoport, source not given)

Fabaceae tree

Mikania micranthaEnglish: mile-a-minute weed, Chinese creeper, American rope, bittervine;French: liane américaine; Other: fue saina (American Samoa, Samoa andNiue); wa mbosuthu, wa mbosuvu, wa mbutako, wa ndamele, ovaova, wabosucu, usuvanua (Fiji); kwalo koburu, (from Ecoport, no source given)

Asteraceae vine

Paraserianthesfalcataria

English: Molucca albizia; Other: tamaligi palagi (American Samoa); tuhkekerosene, tuhkehn karisihn (Pohnpei); ukall ra ngebard (Palau); tamaligi uliuli(Samoa)

Fabaceae tree

Scindapsus aureus English: pothos, money plant; Other: yalu, wa yalu, wa lu, alu, halu, waloa,matha, nanggalanggala, nanggalinggali, tonga, tanga (Fiji) Araceae climber

Syngoniumpodophyllum English: arrowhead plant, goosefoot plant Araceae climber

Tradescanthiaspathacea

English: oyster plant, boat plant, boat lily, moses in a boat; Other: talotalo,laupapaki (Niue)

Commelinaceae herb

27

Table 3. Species that are reported as invasive or weedy elsewhere and are cultivated, common orweedy in Kosrae.

Scientific Name Common Names Family Habit

Abelmoschusmoschatus

English: fautia, musk mallow, musk okra; Other: kamang, ka'mang (Guam);karereon, karereon nikapwerik nik, kareron, likonokon (Chuuk); gongul(Palau); metei, mety, methey (Pohnpei); hathongethong, kamwayang,nikapwerik, setmwechin, sotumo (Yap); wakiwaki, wakewake, wakeke,vakeke, aukiki, okeoke, o'e'e (Fiji); fou ingo (Niue), fau ingo (Wallis andFutuna); loa, fau’ingo (Tonga)

Malvaceae herb

Acacia auriculiformis English: Papuan wattle, auri, earleaf acacia, northern black wattle, ear-podwattle; Other: tuhkehn pwelmwahu (Pohnpei) Fabaceae tree

Allamanda catharticaEnglish: yellow trumpet vine, golden trumpet, allamanda, brownbudallamanda, golden allamanda, golden cup; French: monette jaune; Other:puataunofo (American Samoa and Samoa), lani-ali'i (Hawai'i)

Apocynaceae shrub

Annona muricata

English: soursop, prickly custard apple; French: corossolier, cacheimantierepineux; Other: laguana, laguaná, laguanaha, laguanaba, labuanaha(Guam); sei, sae, truka shai (Pohnpei); jojaab (Marshall Islands); saasaf,sasaf (Chuuk); sausab (Palau); sausau (Yap); sosap (Kosrae); syasyap(Northern Mariana Islands), talapo fotofoto (Niue), ‘apele ‘initia (Tonga);tapotapo papaa (French Polynesia)

Annonaceae tree

Annona squamosaEnglish: sugar apple, sweetsop, custard apple, sugar apple; Other: atis, ates(Guam); ngel ra ngebard (Palau), ‘apele papalangi (Tonga), tapotapo(French Polynesia)

Annonaceae tree

Asystasia gangetica English: Chinese violet, Philippine violet, coromandel Acanthaceae herb

Bambusa spp.

English: bamboo; French: bambou; Other: moor (Yap); iich (Chuuk); pehrien sapahn (Pohnpei); bambuu (Palau); bambu (Kosrae); pi’ao, pi’ao palaoan(Guam and Northern Marianas-Chamorro); bwai (Guam and NorthernMarianas-Carolinian); kaho palangi (Niue); ofe, ‘ohe (French Polynesia),kofe, pitu (Tonga).

Poaceae tree

Bauhinia monandraEnglish: orchid-tree, St. Thomas-tree, Napoleon's plume; flamboyant(Pohnpei), flores mariposa (CNMI); mariposa (Guam); pink butterfly tree(Fiji); Other: pine fua loloa (Niue)

Fabaceae tree

Blechum brownei English: blackweed (Samoa), Browne’s blechum Acanthaceae herb

Bothriochloa bladhii English: blue grass, Australian beardgrass, Caucasian bluestem; Other:desum (Palau); latoka grass, thamboni grass (Fiji) Poaceae grass

Calotropis gigantea English: calotrope, crown flower, mudar; Other: pua-kalaunu (Hawai'i);tahinu (French Polynesia) Ascelepiadaceae shrub

Cananga odorata

Other: ilang-ilang, alang-ilang (Guam, Philippines); ylang-ylang, lengileng(CNMI); chiráng, irang (Palau); ilanlang, ilahnglahng (Kosrae); pwanang,pwuur, pwalang (Chuuk); pur-n-wai, pwurenwai, sair-n-wai, seirin wai, seiren wai (Pohnpei); ilanilan, ilañlañ (Marshall Islands); motoi (Niue); makasoi(Fiji); moso'oi (Am. Samoa and Samoa); mohokimohokoi, mohokoi (Tonga);moto'oi, mata'oi (Cook Islands and Tahiti); makosoi, mokosoi, mokohoi (fromEcoport, source not known)

Annonaceae tree

Cascabela thevetia English: yellow oleander, be-still tree, lucky nutOther: koneta (Chuuk); nohomalie (Hawai'i); irelepsech (Yap) Apocynaceae small tree

Ceiba pentandra

English: kapok, kapok tree, silk-cotton tree, pacae; French: kapokier; Other:kkoton (Chuuk); algodon de Manila, atgodon di Manila, algidon, atgidon deManila (Guam); koatoa, atagodon, arughuschel (Saipan), batte ni gan' ken(Yap); bulik, kotin (Marshall Islands); cottin, koatun, koatoa (Pohnpei);kuhtin, cutin (Kosrae); kalngebard, kalngebárd, kerrekar ngebard (Palau);vauvau ni vavalangi, semar (Fiji); vavae (American Samoa, Samoa, Niue,Tonga); vavau ni lokoloko (from Ecoport, source not given)

Bombacaceae tree

Cenchrus echinatus

English: burgrass, sand-bur, southern sandbur, Mossman River grass(Australia); French: herbe a cateaux (Mauritius); Other: loklok, lellik, lekelik,karmwijmwij (Marshall Islands); se mbulabula (Fiji); motie vihilago, mosievihilango (Niue); piri-piri, pipiri (French Polynesia); cauit-cauitan(Philippines); konpeito-gusa, 'ume'alu (Hawai'i); tuitui, vao tuitui (AmericanSamoa and Samoa); hefa (Tonga); te anti, te kateketeke (Kiribati), cram-cram (New Caledonia)

Poaceae grass

Chloris radiata English: plush-grass, radiate fingergrass Poaceae grass

28

Scientific Name Common Names Family Habit

Commelina diffusa

English: commelina, dayflower, wandering Jew, spiderwort; Other:semprebiban-damalong (Guam); honohono (Hawai'i); mau'utoga, mau’uTonga (Samoa); mohuku vai, musie matala pulu (Tonga); ai rorongi, airongorongo, airogorogo, cobulabula, rongomatailevu, thombulambula,matembulambula, drano,duludauwere, ndrano, ndulandauwere, luna, thonggalonggalo (Fiji), ma’apape (French Polynesia).

Commelinaceae herb

Delonix regiaEnglish: flame tree, flamboyant, poinciana; Other: arbol del fuego, atbot,atbot det fuegu, atbut (Guam and CNMI–Chamorro); fayarbaw (CNMI–Carolinian); nangiosákura, nangyo (Palau); pilampwoia weitahta (Pohnpei);sakuranirow (Yap); sekoula (Fiji); pine (Niue); ‘ohai (Tonga)

Fabaceae tree

Derris ellipticaEnglish: derris; Other: tuba, bagin (Guam); oop, op (Kosrae); dub (Palau);up (Chuuk, Pohnpei); peinuhp, uhp, upanai, upanyap, upkesetik (Pohnpei);yuub (Yap); nduva, duva ni vavalagi, nduva ni vavalangi (Fiji)

Fabaceae vine

Desmanthus virgatus English: slender mimosa, virgate mimosa, wild tantan Fabaceae herbDesmodiumtortuosum English: Florida beggarweed, Spanish clover, dixie ticktrefoil Fabaceae herb

Digitaria ciliarisEnglish: Henry's crabgrass, smooth crabgrass, tropical crab grass, largecrab grass, southern crabgrass, fingergrass, summer grass; Other:kukaepua'a (Hawai'i); Saulangi (Niue)

Poaceae grass

Dissotis rotundifolia English: dissotis, Spanish shawl, pink lady Melastomataceae herb

Eleusine indica

English: goosegrass, wiregrass, goose foot, crow's foot, bullgrass; French:chiendent patte de poule; Other: umog (Guam); deskim, keteketarmalk(Palau); reh takai (Pohnpei); mahkwekwe (Kosrae); katejukjuk (MarshallIslands); manienie ali’i (Hawaii); fahitalo, mosie fahitalo (Niue); kavoronaisivi,vorovoroisivi, ghoraya (Fiji); lau ta'ata'a, ta'ata'a (American Samoa andSamoa); takataka, takataka 'a leala, mohuku siamane (Tonga); tamamau(French Polynesia), te uteute (from Ecoport, source not given)

Poaceae grass

Ficus microcarpa English: Chinese banyan, Malayan banyan, Indian laurel; Other: lulk(Palau); nunu (Chamorro, CNMI) Moraceae tree

Gliricidia sepium English: mother of cacao, quickstick; Spanish: madre de cacao Fabaceae tree

Hedychiumcoronarium

English: white ginger, butterfly lily; Other: tunun, sinser (Chuuk); sinterpwetepwet (Pohnpei); tolon (Puluwat); thevunga, ndrove, cevuga vula,dalasika (Fiji)

Zingiberaceae herb

Hippobromalongiflora English: star of Bethlehem, fetia, madamfate; Other: pua hōkū (Hawai'i) Campanulaceae herb

Impatiens balsamina English: impatiens, balsam, garden balsam, rose balsam, spottedsnapweed; Other: kamantigi (Guam) Balsaminaceae succulent

Ipomoea aquaticaEnglish: aquatic morning glory, swamp cabbage, water spinach, ung-choi,kang kong; Other: kangkong (Yap); cancon, kangkun (Guam); ota karisa,ota karisi, wa kumala, ndrinikava, luve ne tombithi (Fiji), lorenzo (Nauru), tekang kong (Kiribati)

Convolvulaceae aquaticherb

Ipomoea spp. English: morning glory (non-native) Convolvulaceae herb

Moringa oleiferaEnglish: horseradish tree, drumstick tree, ben nut, morango; Other:malungkai, marronggai, marungai, marunggai, malungay, katdes (Guam);sajina (Fiji)

Moringaceae tree

Ocimum gratissimum English: wild basil, clove basil, tree basil; French: basilic; Other: la’ausauga (Samoa) Lamiaceae herb

Orthosiphon aristatus English: cat’s whiskers; Other: emadecharebub (Palau), kumi ni pusi (Fiji),kava ‘i pusi (Tonga) Lamiaceae herb

Paspalumconjugatum

English: T grass, ti grass, sour grass; sour palpalum, Hilo grass (Hawaii);French: herbe créole; herbe de tauère (New Caledonia); Other: fetinwumwune (Chuuk); muhsrasre (Kosrae); udel ra ngebei (Palau); rehn wai(Pohnpei); moise vailima, motie vailima (Niue and Tonga); vao lima(American Samoa and Samoa), vailima matafao (Samoa); vailima, (Samoa,Tonga, Niue)

Poaceae grass

Passiflora foetida

English: love-in-a-mist, wild passion fruit, passionflower, stinkingpassionflower; French: passiflore; Other: kinahulo' atdao, dulce (Saipan);kudamono (Palau); bombom (Chuuk), pompom, pwomwpwomw (Pohnpei);tomates (Yap); pasio vao (American Samoa and Samoa); vine vao (Niue);vaine ‘initia (Tonga); tea biku (Kiribati); pohapoha (Hawai'i); sou, loliloli nikalavo, qaranidila (Fiji).

Passifloraceae vine

Pennisetumpolystachyon

English: mission grass, feathery pennisetum; French: queue de chat;Other: mechen katu (Chuuk), pwokso, dipw rais (Pohnpei), o tamata (Fiji) Poaceae grass

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Scientific Name Common Names Family Habit

Pennisetumpurpureum

English: elephant grass, napier grass, merker grass; French: herbeéléphant, fausse canne à sucre; Other: bokso (Palau); puk-soh (Pohnpei);acfucsracsracsr (Kosrae).

Poaceae grass

Psidium guajava

English: guava; French: goyavier; Other: kuafa (Chuuk); abas (Guam,Saipan-Chamorro, Yap); apas (Saipan); abwas (Saipan-Carolinian);guabang, kuabang (Palau); guahva, kuahpa (Pohnpei); kuhfahfah (Kosrae);kuava (Cook Islands, Tonga); kuava, quwawa, nguava, ngguava ni India,amrut (Fiji); kautoga, kautonga, kautonga tane (Niue); ku'ava, ku'avu, tu'avu(American Samoa and Samoa); te kuawa (Kiribati); kuwawa (Nauru); tuava,tumu tuava, tuvava (French Polynesia); kuawa (Hawai'i); guayaba(Galapagos)

Myrtaceae tree

Saccharumspontaneum

English: wild cane, false sugarcane; Other: banga ruchel (Palau); ac(Kosrae); tebu salah; ahlek, ahlec, (Pohnpei); to aheo (French Polynesia) Poaceae grass

Samanea samanEnglish: monkeypod, rain tree, saman; Other: tronkon mames, trongkon-mames (CNMI-Chamorro); filinganga (CNMI-Carolinian); gumor ni spanis(Yap); 'ohai (Hawaii); vaivai ni vavalangi, vaivai ni vavalagi (Fiji), kasia(Tonga)

Fabaceae tree

Senna alata

English: candle bush; candelabra bush, Roman candle tree, emperor'scandlesticks, ringworm bush (Australia); French: epis d’or; Other: arakak(Chuuk); Acapulco, akapuku, andadose, candalaria, take-biha (Guam);kerula besokel, yult (Palau); rakau honuki, truke-n-kili-n-wai, tuhkehn kilinwai, tirakahonuki (Pohnpei); flay-n-sabouw (Yap); mulamula (Niue); bakauplant (Solomon Islands); mbai ni thangi (Fiji); la'au fai lafa (American Samoa,Samoa and Tonga); te'elango (Tonga)

Fabaceae shrub

Sesbania grandifloraEnglish: hummingbird tree, sesban, scarlet wisteria tree; French: agati agrandes fleurs; Other: agati, agathi (Fiji); oufai, ofai, ouai (FrenchPolynesia); katurai (Guam); sepania (Samoa)

Fabaceae small tree

Solanum torvumEnglish: prickly solanum, devil's fig, turkeyberry, terongan; French: fausseaubergine, aubergine sauvage épineuse; Other: piko (Vanuatu); tisaipale(Tonga); kausoni, soni, kauvoto-votua, kaisurisuri, katai, bhankatiya, soni nivavalagi (Fiji)

Solanaceae shrub

Tecoma stans English: yellow bells, yellow-elder, yellow trumpetbush; Other: peeal(Puluwat), piti (French Polynesia) Bignoniaceae small tree

Turnera ulmifolia English: yellow alder, ramgoat dashalong Turneraceae herb

Urena lobata

English: hibiscus burr, aramina, caesarweed, pink Chinese burr, urena burr;French: jute africain; Other: dadangsi, dadangsi apaka, dadangsi machingat,dádangse (Guam); chosuched e kui, osuched a rechui (Palau); karap, korop(Pohnpei); nognuk, ocher (Chuuk); motipo, mosipo (Niue); mautofu,manutofu (American Samoa and Samoa); qatima, gataya, nggatima (Fiji);mo'osipo (Tonga), piripiri (French Polynesia)

Malvaceae shrub

Wedelia trilobataEnglish: wedelia, trailing daisy, Singapore daisy, creeping ox-eye; Other:ngesil ra ngebard (Palau); dihpw ongohng, tuhke ongohng (Pohnpei);rosrangrang (Kosrae); atiat (Puluwat); ate (Tonga)

Asteraceae herb

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Table 4. Native species (or early introductions) that exhibit aggressive behavior

Scientific Name Common Names Family Habit

Caesalpinia bonduc

English: gray nickers, wait-a-bit; Other: nickaeoo (Chuuk), pacap, pakao,(Guam); tochedulik (Palau); sers mekemad; togodulik (Palau); jeimota (MarshallIslands); alamoa, talatalamoa, talmoa foto (Niue); kakalaioa , hihikolo (Hawai'i);papati, papali (French Polynesia); tataramoa (French Polynesia, Cook Islands);talatala'amoa (Tonga); 'anoso, 'anaoso (Samoa); soni (Fiji); kakalaioa (Hawai'i)

Fabaceae vine

Canna indica

English: canna, canna lily, Indian shot; Other: mongos halum-tano (Guam);luiuenwai (Pohnpei); apeellap, oruuru (Puluwat); fanamanu (American Samoa);fagamanu (American Samoa and Samoa); gasau ni ga (Fiji); te riti (Kiribati),misimisi (Tonga) (Swarbrick, 1997); fagafaga (Futuna); ali'ipoe, li'ipoe, poloka(Hawai'i); pia-raroto’a (French Polynesia)

Cannaceae herb

Chrysopogonaciculatus

English: Mackie's pest, lovegrass, seed grass, golden beardgrass, seedy grass(Solomon Islands); French: herbe plate, herbe à piquant; Other: inifuk, palaii(Guam); iul (Palau); manienie 'ula (Hawai'i); mutia tai (Samoa); mosie fisi(Niue); matapekepeke, matapekapeka (Tonga); herbe plate (Vanuatu), papapa(French Polynesia)

Poaceae grass

Costus speciosus English: crepe ginger, crape ginger, wild ginger, Malay ginger, canereed;Other: isebsab (Palau) Zingiberaceae herb

Ischaemumpolystachyum English: paddle grass; Other: mah (Kosrae); reh padil (Pohnpei) Poaceae grass

Kyllinga nemoralisEnglish: white kyllinga, whitehead spikesedge; Other: kili'o'opu (Hawai'i), matieupo’o, matie tahiti, mo’u upo’onui, mo'u upo'o (Tahiti); tuise (Samoa); pakopako,pakopako 'ae kuma (Tonga)

Cyperaceae sedge

Merremia peltata

English: merrimia; Other: wachathal (yap); lohl, iol, yol (Pohnpei); kebeas(Palau); lagon, lagun (Guam); fitau, fitaw (Chuuk); pala, pul, puhlah (Kosrae);fue, fue vao, fue kula (Niue); fue lautetele (American Samoa and Samoa); fuemea (Tonga); abui, grobihi, arosumou (Solomon Islands); wa mbula, wa bula,wa damu, wa ndamu, viliyawa, wiliviwa, veliyana, wiliao (Fiji); pohue (FrenchPolynesia)

Convolvulaceae vine

Miscanthus floridulus

English: miscanthus, swordgrass, sawgrass, reed grass, Pacific Islandsilvergrass, Japanese silvergrass, Chinese silvergrass, Chinese fairygrass,eulalia; Other: nete, neti, tupon nette, tupun-neti (Guam); nette (Guam,Saipan); mah (Kosrae); sapala, sapeleng, sapalang (Pohnpei); aset, ene(Chuuk); banga ruchel, medecherecher bokso, pagaluel (Palau); 'u (AmericanSamoa, Samoa and Futuna); ngasau (Fiji); kaho (Niue and Tonga); kaho tonga,kaho foi’iki (Tonga), aeho (French Polynesia).

Poaceae grass

Rubus moluccanusEnglish: Molucca raspberry, broad leafed bramble, Molucca bramble, eelkek;French: piquant lou-lou (Mauritius); Other: kohkihl (Kosrae); soni, wa sori, wangandrongandro, wa votovotoa, (Fiji)

Rosaceae shrub

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Appendix 3

Invasive species present in Guam, Palau or Hawai'i but not present in Kosrae

Species present: .

Scientific Name Guam Pohnpei Hawai'iAcacia confusa . . .Acacia farnesiana . . .Acacia mangium . . .Acacia mearnsii . . .Acacia melanoxylon . . .Acacia spp. . . .Agave sisalana . . .Ailanthus altissima . . .Albizia chinensis . . .Albizia lebbeck . . .Aleurites moluccana . . .Alpinia purpurata . . .Alstonia macrophylla . . .Anacardium occidentale . . .Andropogon glomeratus . . .Andropogon virginicus . . .Angiopteris evecta . . .Annona glabra . . .Anredera cordifolia . . .Antigonon leptopus . . .Ardisia crenata . . .Ardisia elliptica . . .Argyreia nervosa . . .Aristolochia littoralis . . .Arthrostema ciliatum . . .Arundo donax . . .Asclepias physocarpa . . .Asparagus densiflorus . . .Asparagus setaceus . . .Axonopus fissifolius . . .Barleria lupulina . . .Bauhinia purpurea . . .Bidens pilosa . . .Bischofia javanica . . .Bocconia frutescens . . .Boerhavia coccinea . . .Brachiaria mutica . . .Brachiaria subquadripara . . .Buddleia asiatica . . .Buddleia davidii . . .

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Scientific Name Guam Pohnpei Hawai'iBuddleia madagascariensis . . .Caesalpinia decapetala . . .Calamus spp. . . .Calliandra haematocephala . . .Calliandra surinamensis . . .Callisia fragrans . . .Calopogonium mucunoides . . .Calotropis procera . . .Cardiospermum grandiflorum . . .Cardiospermum halicacabum . . .Castilla elastica . . .Casuarina equisetifolia . . .Casuarina glauca . . .Cecropia obtusifolia . . .Cecropia peltata . . .Cedrela odorata . . .Cenchrus brownii . . .Cenchrus ciliaris . . .Centrosema pubescens . . .Cestrum diurnum . . .Cestrum nocturnum . . .Chamaecrista nictitans . . .Chloris barbata . . .Chrysobalanus icaco . . .Chrysophyllum oliviforme . . .Cinchona pubescens . . .Cinnamomum burmannii . . .Cinnamomum camphora . . .Cinnamomum verum . . .Cirsium vulgare . . .Cissus nodosa . . .Citharexylum caudatum . . .Citharexylum spinosum . . .Clausena excavata . . .Clerodendrum buchananii var.fallax

. . .

Clerodendrum japonicum . . .Clerodendrum paniculatum . . .Clidemia hirta . . .Clitoria ternatea . . .Clusea rosea . . .Coccinia grandis . . .Coffea arabica . . .Conocarpus erectus . . .Cordia glabra . . .Cortaderia jubata . . .

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Scientific Name Guam Pohnpei Hawai'iCorynocarpus laevigatus . . .Crassocephalum crepidoides . . .Crotalaria anagyroides . . .Cryptostegia grandiflora . . .Cuscuta campestris . . .Cynodon dactylon . . .Cyperus alternifolius . . .Cyperus rotundus . . .Derris indica . . .Digitaria insularis . . .Digitaria violascens . . .Dillenia suffruticosa . . .Duranta erecta . . .Egeria densa . . .Ehrharta stipoides . . .Eichhornia crassipes . . .Elaeagnus umbellata . . .Elaeis guineensis . . .Elephantopus mollis . . .Erigeron karvinskianus . . .Eriobotrya japonica . . .Eugenia uniflora . . .Ficus elastica . . .Ficus nota . . .Ficus rubiginosa . . .Flemingia strobilifera . . .Flindersia brayleyana . . .Fuchsia boliviana . . .Fuchsia magellanica . . .Furcraea foetida . . .Grevillea banksii . . .Grevillea robusta . . .Haematoxylum campechianum . . .Hedychium flavescens . . .Hedychium gardnerianum . . .Heliocarpus popayanensis . . .Hemigraphis alternata . . .Heterocentron subtriplinervium . . .Hiptage benghalensis . . .Hylocereus undatus . . .Hyparrhenia rufa . . .Hypochoeris radicata . . .Hyptis capitata . . .Hyptis pectinata . . .Hyptis suaveolens . . .Impatiens walleriana . . .

34

Scientific Name Guam Pohnpei Hawai'iImperata cylindrica . . .Indigofera suffruticosa . . .Ischaemum rugosum . . .Ischaemum timorense . . .Jacaranda mimosifolia . . .Jasminum fluminense . . .Jatropha curcas . . .Jatropha gossypifolia . . .Justicia carnea . . .Kalanchoe pinnata . . .Lemna spp. . . .Leonotis nepetifolia . . .Leptospermum ericoides . . .Leptospermum scoparium . . .Licuala grandis . . .Ligustrum spp. . . .Lonicera japonica . . .Lophostemon confertus . . .Macfadyena unguis-cati . . .Macroptilium atropurpureum . . .Medinilla venosa . . .Melaleuca quinquenervia . . .Melastoma candidum . . .Melastoma sanguineum . . .Melia azedarach . . .Melinis minutiflora . . .Melochia umbellata . . .Merremia tuberosa . . .Miconia calvescens . . .Mikania scandens . . .Mimosa invisa . . .Mimosa pudica . . .Momordica charantia . . .Montanoa hibiscifolia . . .Muntingia calabura . . .Murraya paniculata . . .Myriophyllum aquaticum . . .Neonotonia wightii . . .Nymphaea spp. . . .Ochna serrulata . . .Ochna thomasiana . . .Ochroma pyramidale . . .Odontonema tubaeforme . . .Olea europaea . . .Operculina ventricosa . . .

35

Scientific Name Guam Pohnpei Hawai'iOpuntia spp. . . .Oxyspora paniculata . . .Paederia foetida . . .Panicum maximum . . .Panicum repens . . .Parkinsonia aculeata . . .Paspalum dilatatum . . .Paspalum fimbriatum . . .Paspalum orbiculare . . .Paspalum paniculatum . . .Paspalum urvillei . . .Passiflora caerulea . . .Passiflora edulis . . .Passiflora laurifolia . . .Passiflora ligularis . . .Passiflora maliformis . . .Passiflora mollissima . . .Passiflora pulchella . . .Passiflora quadrangularis . . .Passiflora suberosa . . .Pennisetum clandestinum . . .Pennisetum setaceum . . .Persea americana . . .Philadelphus karwinskyanus . . .Phormium tenax . . .Phylostachys nigra . . .Pinus caribaea . . .Piper auritum . . .Pistia stratiotes . . .Pithecellobium dulce . . .Pittosporum tobira . . .Pittosporum undulatum . . .Plectranthus scutellarioides . . .Pluchea indica . . .Pluchea symphytifolia . . .Prosopis spp. . . .Psidium cattleianum . . .Psidium guineense . . .Pueraria lobata . . .Pueraria phaseoloides . . .Quisqualis indica . . .Rhodomyrtus tomentosa . . .Rhynchelytrum repens . . .Ricinus communis . . .Rivina humilis . . .

36

Scientific Name Guam Pohnpei Hawai'iRubus rosifolius . . .Rubus spp. . . .Ruellia prostrata . . .Sambucus mexicana . . .Sanchezia nobilis . . .Sanseviera trifasciata . . .Schefflera actinophylla . . .Schinus molle . . .Schinus terebinthifolius . . .Schizachyrium condensatum . . .Senecio mikanioides . . .Senna obtusifolia . . .Senna septemtrionalis . . .Setaria pallide-fusca . . .Setaria palmifolia . . .Solandra maxima . . .Solanum mauritianum . . .Solanum seaforthianum . . .Sorghum halepense . . .Sorghum sudanense . . .Spathodea campanulata . . .Sphaeropteris cooperi . . .Sporobolus elongatus . . .Sporobolus indicus . . .Stachytarpheta urticifolia . . .Stipa cernua . . .Stizolobium pruriens . . .Syncarpia glomulifera . . .Syzygium cumini . . .Syzygium jambos . . .Tabebuia heterophylla . . .Tetrazygia bicolor . . .Thunbergia alata . . .Thunbergia fragrans . . .Thunbergia grandiflora . . .Thunbergia laurifolia . . .Tibouchina herbacea . . .Tibouchina urvileana . . .Tillandsia usneoides . . .Tithonia diversifolia . . .Toona ciliata . . .Tradescanthia zebrina . . .Trema orientalis . . .Tribulus cistoides . . .Tribulus terrestris . . .Triphasia trifolia . . .

37

Scientific Name Guam Pohnpei Hawai'iTripsacum latifolium . . .Triumfetta rhomboidea . . .Triumfetta semitriloba . . .Vitex parviflora . . .Zantedeschia aethiopica . . .Ziziphus mauritiana . . .

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Appendix 3

Scientific name synonyms

This table lists commonly used scientific name synonyms of the listed invasive species. Names sometimes changebecause better information is available on the taxonomy of a species or precedence is given to an earlier author.

Synonym Listed as FamilyAcacia adansonii Acacia nilotica FabaceaeAcacia adstringens Acacia nilotica FabaceaeAcacia arabica Acacia nilotica FabaceaeAdenanthera falcataria Paraserianthes falcataria FabaceaeAdenoropium gossypifolium Jatropha gossypifolia EuphorbiaceaeAgave cubensis Furcraea cubensis AgavaceaeAgave gigantea Furcraea foetida AgavaceaeAgave rigida Agave sisilana AgavaceaeAgrostis indica Sporobolus indicus PoaceaeAlbizia distachya Albizia lophantha FabaceaeAlbizia falcataria Paraserianthes falcataria FabaceaeAlbizia saman Samanea saman FabaceaeAlbizia stipulata Albizia chinensis FabaceaeAleurites triloba Aleurites moluccana EuphorbiaceaeAllamanda hendersonii Allamanda cathartica ApocynaceaeAndropogon bladhii Bothriochloa bladhii PoaceaeAndropogon condensatus Schizachyrium condensatum PoaceaeAndropogon halepensis Sorghum halepense PoaceaeAndropogon intermedius Bothriochloa bladhii PoaceaeAndropogon marcrourus Andropogon glomeratus PoaceaeAndropogon rufus Hyparrhenia rufa PoaceaeAntelaea azadirachta Azadirachta indica MeliaceaeAraujia hortorum Araujia sericifera AsclepiadaceaeArdisia humilis Ardisia elliptica MyrsinaceaeArdisia solanacea Ardisia elliptica MyrsinaceaeArdisia squamulosa Ardisia elliptica MyrsinaceaeArgyreia speciosa Argyreia nervosa ConvolvulaceaeAristolochia elegans Aristolochia littoralis AristolochiaceaeArum seguine Dieffenbachia maculata AraceaeAsclepias procera Calotropis procera AsclepiadaceaeAsparagopsis densiflora Asparagus densiflorus LiliaceaeAsparagus plumosus Asparagus setaceus LiliaceaeAsparagus sprengeri Asparagus densiflorus LiliaceaeAxonopus affinis Axonopus fissifolius PoaceaeAxonopus poiretii Sporobolus indicus PoaceaeBambusa nigra Phyllostachys nigra PoaceaeBidens alba Bidens pilosa AsteraceaeBignonia venusta Pyrostegia venusta BignoniaceaeBlechum pyramidatum Blechum brownei AcanthaceaeBlochmannia weigeltiana Triplaris weigeltiana PolygonaceaeBoerhavia diffusa Boerhavia coccinea NyctaginaceaeBothriochloa caucasica Bothriochloa bladhii PoaceaeBoussingaultia baselloides Anrederia cordifolia Basellaceae

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Synonym Listed as FamilyBoussingaultia cordifolia Anrederia cordifolia BasellaceaeBrachiaria purpurascens Brachiaria mutica PoaceaeBrassaia actinophylla Schefflera actinophylla AraliaceaeBryonia grandis Coccinea grandis CucurbitaceaeBryophyllum pinnatum Kalanchoë pinnata CrassulaceaeBuddleja asiatica Buddleia asiatica LoganiaceaeBuddleja davidii Buddleia davidii LoganiaceaeBuddleja madagascarienses Buddleia madagascarienses LoganiaceaeCaesalpinia crista Caesalpinia bonduc FabaceaeCaesalpinia sepiaria Caesalpinia decapetala FabaceaeCalla aethiopica Zantedeschia aethiopica AraceaeCaryophyllus racemosus Pimenta racemosa MyrtaceaeCassia aeschynomene Chamaecrista nictitans FabaceaeCassia alata Senna alata FabaceaeCassia laevigata Senna septemtrionalis FabaceaeCassia lechenaultiana Chamaecrista nictitans FabaceaeCassia leschenaultiana Chamaecrista nictitans FabaceaeCassia obtusifolia Senna obtusifolia FabaceaeCassia septemtrionalis Senna septemtrionalis FabaceaeCassia siamea Senna siamea FabaceaeCastilloa elastica Castilla elastica MoraceaeCasuarina litorea Casuarina equisetifolia CasuarinaceaeCasuarina littorea Casuarina equisetifolia CasuarinaceaeCedrela australis Toona ciliata MeliaceaeCedrela toona Toona ciliata MeliaceaeCenchrus brevisetus Cenchrus echinatus PoaceaeCenchrus pungens Cenchrus echinatus PoaceaeCenchrus quinquevalvis Cenchrus echinatus PoaceaeCenchrus setosus Pennisetum polystachyon PoaceaeCenchrus viridis Cenchrus echinatus PoaceaeChalcas paniculata Murraya paniculata RutaceaeChloris inflata Chloris barbata PoaceaeChloris paraguayensis Chloris barbata PoaceaeChlorocyperus rotundus Cyperus rotundus CyperaceaeCinchona succirubra Cinchona pubescens RubiaceaeCinnamomum zeylanicum Cinnamomum verum LauraceaeCitharexylum quadrangulare Citharexylum spinosum VerbenaceaeClerodendrum fragrans Clerodendrum chinense VerbenaceaeClerodendrum philippinum Clerodendrum chinense VerbenaceaeClerodendrum speciosissimum Clerodendrum buchananii VerbenaceaeCoccinea cordifolia Coccinea grandis CucurbitaceaeColeus amboinicus Plectranthus amboinicus LamiaceaeColeus blumei Plectranthus scutellarioides LamiaceaeColeus scutellarioides Plectranthus scutellarioides LamiaceaeCordia collococca Cordia glabra BoraginaceaeCordia macrostachya Cordia curassavica BoraginaceaeCrotalaria micans Crotalaria anagyroides FabaceaeCroton moluccanus Aleurites moluccana EuphorbiaceaeCuscuta pentagona Cuscuta campestris ConvolvulaceaeCyathea cooperi Sphaeropteris cooperi CyatheaceaeCyperus aromaticus Kyllinga polyphylla CyperaceaeCyperus flabelliformis Cyperus alternifolius Cyperaceae

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Synonym Listed as FamilyCyperus involucratus Cyperus alternifolius CyperaceaeCyperus kyllingia Kyllinga nemoralis CyperaceaeCyperus purpuro-variegatus Cyperus rotundus PoaceaeCyperus stoloniferum pallidus Cyperus rotundus PoaceaeCyperus tetrastachyos Cyperus rotundus PoaceaeCyperus tuberosus Cyperus rotundus PoaceaeDaubentonia punicea Sesbania punicea FabaceaeDelairea odorata Senecio mikanioides AsteraceaeDichanthium bladhii Bothriochloa bladhii PoaceaeDichrostachys glomerata Dichrostachys cinerea FabaceaeDichrostachys natans Dichrostachys cinerea FabaceaeDieffenbachia picta Dieffenbachia maculata AraceaeDieffenbachia seguine Dieffenbachia maculata AraceaeDipteracanthus prostrata Ruellia prostrata AcanthaceaeDolichos pruriens Stizolobium pruriens FabaceaeDoxantha unguis-cati Macfadyena unguis-cati BignoniaceaeDuranta repens Duranta erecta VerbenaceaeElephantopus scaber Elephantopus mollis AsteraceaeEpipremnum aureum Scindapsus aureus AraceaeEpipremnum pinnatum Scindapsus aureus AraceaeEugenia cumini Syzygium cumini MyrtaceaeEugenia jambos Syzygium jambos MyrtaceaeEupatorium inulaefolium Austroeupatorium inulaefolium AsteraceaeEupatorium odoratum Chromolaena odorata AsteraceaeFicus thonningii Ficus microcarpa MoraceaeFurcraea gigantea Furcraea foetida AgavaceaeGliricidia maculata Gliricidia sepium FabaceaeGlycine javanica Neonotonia wightii FabaceaeGlycine wightii Neonotonia wightii FabaceaeGrewia subinaequalis Grewia asiatica TiliaceaeGuilandina bonduc Caesalpinia bonduc FabaceaeGynura crepidioides Crassocephalum crepidioides AsteraceaeHemigraphis colorata Hemigraphis alternata AcanthaceaeHeptapleurum arboricola Schefflera arboricola AraliaceaeHibiscus abelmoschus Abelmoschus moschatus MalvaceaeHiptage madablota Hiptage benghalensis MalpighiaceaeImperata arundinacea Imperata cylindrica PoaceaeImperata conferta Imperata cylindrica PoaceaeIpomoea peltata Merremia peltata ConvolvulaceaeIpomoea tuberosa Merremia tuberosa ConvolvulaceaeIschaemum digitatum var. polystachyum Ischaemum polystachyum PoaceaeIsotoma longiflora Hippobroma longiflora CampanulaceaeJacobinia carnea Justicia carnea AcantaceaeJasminum azoricum Jasminum fluminense OleaceaeJussiaea grandiflora Ludwigia peruviana OnagraceaeJussiaea grandiflora Ludwigia peruviana OnagraceaeJussiaea peruviana Ludwigia peruviana OnagraceaeJussiaea peruviana Ludwigia peruviana OnagraceaeKunzea ericoides Leptospermum ericoides MyrtaceaeKunzea scoparium Leptospermum scoparium MyrtaceaeKyllinga cephalotes Kyllinga nemoralis CyperaceaeKyllinga monocephala Kyllinga nemoralis Cyperaceae

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Synonym Listed as FamilyLaurentia longiflora Hippobroma longiflora CampanulaceaeLeucaena glauca Leucaena leucocephala FabaceaeMeibomia nicaraguense Desmodium nicaraguense FabaceaeMeibomia rensonii Desmodium nicaraguense FabaceaeMelastoma decemfidum Melastoma sanguineum MelastomataceaeMelia azadirachta Azadirachta indica MeliaceaeMelicoccus bijuga Melicoccus bijugatus SapindaceaeMelinis repens Rhynchelytrum repens PoaceaeMelochia compacta Melochia umbellata SterculiaceaeMelochia indica Melochia umbellata SterculiaceaeMemecylon caeruleum Memecylon floribundum MelastomataceaeMerremia nymphaeifolia Merremia peltata ConvolvulaceaeMetrosideros glomulifera Syncarpia glomulifera MyrtaceaeMiconia magnifica Miconia calvescens MelostomataceaeMicrolaena stipoides Ehrharta stipoides PoaceaeMimosa cinerea Dichrostachys cinerea FabaceaeMimosa diplotricha Mimosa invisa FabaceaeMimosa dulcis Pithecellobium dulce FabaceaeMimosa lebbeck Albizia lebbeck FabaceaeMiscanthus japonicus Miscanthus floridulus PoaceaeMoghania strobilifera Flemingia strobilifera FabaceaeMoringa pterygosperma Moringa oleifera MoringaceaeMucuna pruriens Stizolobium pruriens FabaceaeMurraya exotica Murraya paniculata RutaceaeMyrtus dioica Pimenta dioica MyrtaceaeMyrtus pimenta Pimenta dioica MyrtaceaeNassella cernua Stipa cernua PoaceaeNotonia wightii Neonotonia wightii FabaceaeOchna kirkii Ochna thomasiana OchnaceaeOchroma lagopus Ochroma pyramidale BombacaceaeOcimum scutellarioides Plectranthus scutellarioides LamiaceaeOdontonema callistachyum Odontonema tubaeforme AcanthaceaeOdontonema strictum Odontonema tubaeforme AcanthaceaeOperculina peltata Merremia peltata ConvolvulaceaeOperculina turpethum Operculina ventricosa ConvolvulaceaeOplismenus polystachyus Echinochloa polystachya PoaceaeOrthosiphon stamineus Orthosiphon aristatus LamiaceaePaederia scandens Paederia foetida RubiaceaePanicum barbinode Brachiaria mutica PoaceaePanicum guadeloupense Brachiaria mutica PoaceaePanicum melinis Melinis minutiflora PoaceaePanicum minutiflora Melinis minutiflora PoaceaePanicum muticum Brachiaria mutica PoaceaePanicum palmaefolium Setaria palmifolia PoaceaePanicum purpurascens Brachiaria mutica PoaceaePanicum subquadriparum Brachiaria subquadripara PoaceaeParaserianthes lophantha Albizia lophantha FabaceaePaspalum cartilagineum Paspalum orbiculare PoaceaePaspalum scrobiculatum Paspalum orbiculare PoaceaePassiflora bicornis Passiflora pulchella PassafloraceaePassiflora mixta Passiflora mollissima PassafloriaceaePassiflora tripartita Passiflora mollissima Passifloriaceae

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Synonym Listed as FamilyPennisetum cenchroides Cenchrus ciliaris PoaceaePennisetum ciliare Cenchrus ciliaris PoaceaePennisetum polystachion Pennisetum polystachyon PoaceaePennisetum ruppelii Pennisetum setaceum PoaceaePennisetum setosum Pennisetum polystachyon PoaceaePhaseolus atropurpureus Macroptilium atropurpureum FabaceaePhlomis nepetaefolia Leonotis nepetaefolia LamiaceaePhyllostachys henionis Phyllostachys nigra PoaceaePhysianthus albens Araujia sericifera AsclepiadaceaePimenta officinalis. Pimenta dioica MyrtaceaePiscidia punicea Sesbania punicea FabaceaePithecellobium saman Samanea saman FabaceaePluchea odorata Pluchea symphytifolia AsteraceaePoinciana regia Delonix regia FabaceaePothos aureus Scindapsus aureus AraceaePothos pinnata Scindapsus aureus AraceaePraxelis clematidea Eupatorium catarium AsteraceaeProasparagus plumosus Asparagus setaceus LiliaceaePseudechinolaena polystachya Echinochloa polystachya PoaceaePsidium aracá Psidium guineense MyrtaceaePsidium littorale Psidium cattleianum MyrtaceaePsidium molle Psidium guineense MyrtaceaePsidium schiedeanum Psidium guineense MyrtaceaePueraria harmsii Pueraria lobata FabaceaePueraria montana Pueraria lobata FabaceaePueraria novo-guiniensis Pueraria lobata FabaceaePueraria thunbergiana Pueraria lobata FabaceaePueraria triloba Pueraria lobata FabaceaeRacosperma longifolia Acacia longifolia FabaceaeRacosperma melanoxylon Acacia melanoxylon FabaceaeRaphidophora aurea Scindapsus aureus AraceaeRhaphidophora aurea Scindapsus aureus AraceaeRhaphidophora pinnata Scindapsus aureus AraceaeRhoeo discolo Tradescanthia spathacea CommelinaceaeRhoeo spathacea Tradescanthia spathacea CommelinaceaeRivina laevis Rivina humilis PhytolaccaceaeRuellia alternata Hemigraphis alternata AcanthaceaeSaccharum floridulum Miscanthus floridulus PoaceaeSalvinia auriculata Salvinia molesta SalviniaceaeSanchezia speciosa Sanchezia nobilis AcanthaceaeSetaria pumila ssp. pallidifusca Setaria pallide-fusca PoaceaeSolandra hartwigii Solandra maxima SolanaceaeSolandra nitida Solandra maxima SolanaceaeSolanum auriculatum Solanum mauritianum SolanaceaeSolanum houstonii Solanum tampicense SolanaceaeSolanum verbascifolium Solanum mauritianum SolanaceaeSolenostemon scutellarioides Plectranthus scutellarioides LamiaceaeSorghum bicolor ssp. drummondii Sorghum sudanense PoaceaeSporobolus diander Sporobolus indicus PoaceaeSporobolus poiretii Sporobolus indicus PoaceaeStachytarpheta cayennensis Stachytarpheta urticifolia VerbenaceaeStachytarpheta urticaefolia Stachytarpheta urticifolia Verbenaceae

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Synonym Listed as FamilyStizolobium aterrimum Stizolobium pruriens FabaceaeStizolobium deeringianum Stizolobium pruriens FabaceaeStizolobium niveum Stizolobium pruriens FabaceaeStylosanthes erecta Stylosanthes guianensis FabaceaeStylosanthes gracilis Stylosanthes guianensis FabaceaeStylosanthes guineensis Stylosanthes guianensis FabaceaeSwietenia senegalensis Khaya senegalensis MeliaceaeSyncarpia laurifolia Syncarpia glomulifera MyrtaceaeSyngonium angustatum Syngonium podophyllum AraceaeSyzygium floribundum Waterhousia floribunda MyrtraceaeTabebuia pallida Tabebuia heterophylla BignoniaceaeTabebuia pentaphylla Tabebuia heterophylla BignoniaceaeTerminalia erecta Conocarpus erectus CombretaceaeThelechitonia trilobata Wedelia trilobata AsteraceaeThevetia nerifolia Cascabela thevetia ApocynaceaeThevetia peruviana Cascabela thevetia ApocynaceaeThunbergia harrisii Thunbergia laurifolia AcanthaceaeToona australis Toona ciliata MeliaceaeTrachypogon rufus Hyparrhenia rufa PoaceaeTradescanthia discolor Tradescanthia spathacea CommelinaceaeTrema cannabina Trema orientalis UlmaceaeTrichachne insularis Digitaria insularis PoaceaeTricholaena repens Rhynchelytrum repens PoaceaeTricholaena rosea Rhynchelytrum repens PoaceaeTriplaris surinamensis Triplaris weigeltiana PolygonaceaeTriumfetta bartramia Triumfetta rhomboidea TiliaceaeUrena sinuata Urena lobata MalvaceaeUrochloa mutica Brachiaria mutica PoaceaeUrochloa subquadripara Brachiaria subquadripara PoaceaeWedelia gossweileri Blainvillea gayana AsteraceaeZebrina pendula Tradescantia zebrina Commelinaceae