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9 'LVWULEXWLRQ RI 0XGVNLSSHUV (Gobiidae: Oxudercinae) on the Selangor Coast .KDLURQL]DP 0= DQG 1RUPD5DVKLG < Institute of Biological Sciences, University of Malaya, 50603 Kuala Lumpur 9.1 Introduction 0XGVNLSSHUV RQH RI WKH ÀVKHV WKDW DUH UHVWULFWHG WR WKH FRDVWDO DQG HVWXDULQH KDELWDWV LQ WKH 2OG :RUOG DUH PHPEHUV RI WKH *RELLG VXEIDPLO\ 2[XGHUFLQDH 7KH VXEIDPLO\ FRPSULVHV PRUH WKDQ VSHFLHV 0XUG\ 1RW RQO\ FDQ WKH\ VXUYLYH RXW RI ZDWHU EXW DUH DOVR DEOH WR FOLPE WUHHV ZKLOH PDLQWDLQLQJ WKHLU DELOLW\ WR VZLP OLNH RWKHU ÀVKHV 7KH\ DUH UHVWULFWHG WR WKH VRIW PXG ERWWRP VKRUHV LQ LQWHUWLGDO DUHDV HVWXDULQH KDELWDWV DQG PDQJURYH VZDPSV RI WKH ,QGR3DFLÀF UHJLRQ *RUGRQ et al &OD\WRQ 0XUG\ UHSRUWHG WKH SUHVHQFH RI JHQHUD LQFOXGLQJ VSHFLHV ZRUOGZLGH 7KRVH FRQÀQHG WR WKH FRDVWDO DUHDV RI WKH ZHVW FRDVW RI 3HQLQVXODU 0DOD\VLD ZHUH VSHFLHV IURP JHQHUD 7KH LQWHUWLGDO ]RQHV RQ WKH FRDVW RI 3HQLQVXODU 0DOD\VLD FRQVLVW RI PDQJURYHV HVWXDULQH DQG PXGÁDW KDELWDWV ZKLFK KRVW D GLYHUVH PDULQH IDXQD .XDOD 6HODQJRU RQ WKH 6HODQJRU FRDVW KDV H[WHQVLYH PDQJURYHV 6HYHUDO DVSHFWV RI WKH PDQJURYH IDXQDO HFRORJ\ KDYH EHHQ UHYLHZHG E\ 0DFQDH %HUU\ 6DVHNXPDU DQG 6DVHNXPDU et al 0DK VWXGLHG WKH EHKDYLRXU ZKLOH RWKHUV )DULGDK +DQLP 0XKDPPDG $OL +DP]DK VWXGLHG PDQ\ DVSHFWV RI PXGVNLSSHU HFRORJ\ 7KH REMHFWLYHV RI WKLV SDSHU DUH WR UHYLHZ WKH VSHFLHV ULFKQHVV GLVWULEXWLRQ DQG RFFXUUHQFH RI PXGVNLSSHUV LQ WKH YDULRXV KDELWDWV RI WKH 6HODQJRU FRDVW 9.2 The study area 7KH VWXG\ ZDV FDUULHG RXW LQ WKH FRDVWDO DUHDV RI 0RULE ·1 ·( 6HPHQWD ·1 ·( -HUDP ·1 ·( .DPSXQJ .XDQWDQ ·1 ·( DQG .XDOD 6HODQJRU ·1 ·( LQ 6HODQJRU RQ WKH ZHVW FRDVW RI 3HQLQVXODU 0DOD\VLD 0RULE DQG -HUDP KDYH H[WHQVLYH PXGÁDWV RQ WKH IRUHVKRUH RI WKH PDQJURYHV 6DPSOHV ZHUH FROOHFWHG IURP PXGÁDWV ULYHU EDQNV DQG IRUHVWHG DUHDV IURP VHDZDUG IRUHVKRUH WR WKH HURGHG EDQNV +HUH WKH PDQJURYH IRUHVWV ZHUH GRPLQDWHG E\ Avicennia alba, Sonneratia alba and Rhizophora mucronata 7KH PDQJURYH IRUHVW KHUH FDQ EH GLYLGHG LQWR IRXU ]RQHV 1 Avicennia alba ]RQH ` PDQJURYH 2 Avicennia alba and Sonneratia alba PL[HG IRUHVW ]RQH ` SLRQHHU VKRUH 3 Sonneratia alba ]RQH ` ORFDWHG DW WKH Rhizophora mucronata ]RQH ` HURGHG EDQN DUHDV

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Page 1: LVWULEXWLRQ RI 0XGVNLSSHUV (Gobiidae: Oxudercinae) … · the sonneratia alba and rhizophora mucronata ]rqhv ri 0rule zkhuh wkh vxevwudwhv zhuh vdqg\ pxg 7kh pxgvnlsshuv zhuh irxqg

9

'LVWULEXWLRQ�RI�0XGVNLSSHUV�(Gobiidae: Oxudercinae) on the Selangor Coast

.KDLURQL]DP��0�=��DQG�1RUPD�5DVKLG��<�Institute of Biological Sciences, University of Malaya, 50603 Kuala Lumpur

9.1 Introduction0XGVNLSSHUV��RQH�RI�WKH�ÀVKHV�WKDW�DUH�UHVWULFWHG�WR�WKH�FRDVWDO�DQG�HVWXDULQH�KDELWDWV�LQ�WKH�2OG�:RUOG��DUH�PHPEHUV�RI�WKH�*RELLG�VXEIDPLO\�2[XGHUFLQDH��7KH�VXEIDPLO\�FRPSULVHV�PRUH�WKDQ�������VSHFLHV��0XUG\���������1RW�RQO\�FDQ�WKH\�VXUYLYH�RXW�RI�ZDWHU��EXW�DUH�DOVR�DEOH�WR�FOLPE�WUHHV�ZKLOH�PDLQWDLQLQJ�WKHLU�DELOLW\�WR�VZLP�OLNH�RWKHU�ÀVKHV��7KH\�DUH�UHVWULFWHG�WR�WKH�VRIW�PXG�ERWWRP�VKRUHV�LQ�LQWHUWLGDO�DUHDV��HVWXDULQH�KDELWDWV�DQG�PDQJURYH�VZDPSV�RI�WKH�,QGR�3DFLÀF�UHJLRQ��*RUGRQ�et al���������&OD\WRQ���������0XUG\��������UHSRUWHG�WKH�SUHVHQFH�RI����JHQHUD�LQFOXGLQJ����VSHFLHV�ZRUOGZLGH��7KRVH�FRQÀQHG�WR�WKH�FRDVWDO�DUHDV�RI�WKH�ZHVW�FRDVW�RI�3HQLQVXODU�0DOD\VLD�ZHUH���VSHFLHV�IURP���JHQHUD�

7KH� LQWHUWLGDO�]RQHV�RQ� WKH� FRDVW�RI�3HQLQVXODU�0DOD\VLD� FRQVLVW�RI�PDQJURYHV��HVWXDULQH�DQG�PXGÁDW�KDELWDWV�ZKLFK�KRVW�D�GLYHUVH�PDULQH�IDXQD��.XDOD�6HODQJRU�RQ�WKH�6HODQJRU�FRDVW�KDV�H[WHQVLYH�PDQJURYHV��6HYHUDO�DVSHFWV�RI�WKH�PDQJURYH�IDXQDO�HFRORJ\�KDYH�EHHQ�UHYLHZHG�E\�0DFQDH���������%HUU\���������6DVHNXPDU��������DQG�6DVHNXPDU�et al�� ��������0DK� ������� VWXGLHG� WKH�EHKDYLRXU�ZKLOH�RWKHUV� �)DULGDK�+DQLP��������0XKDPPDG�$OL��������+DP]DK��������VWXGLHG�PDQ\�DVSHFWV�RI�PXGVNLSSHU�HFRORJ\�

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9.2 The study area7KH�VWXG\�ZDV�FDUULHG�RXW� LQ� WKH�FRDVWDO�DUHDV�RI�0RULE� �����·1��������·(���6HPHQWD�����·1��������·(���-HUDP������·1��������·(���.DPSXQJ�.XDQWDQ������·1��������·(���DQG�.XDOD�6HODQJRU������·1��������·(��LQ�6HODQJRU��RQ�WKH�ZHVW�FRDVW�RI�3HQLQVXODU�0DOD\VLD�

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9.3 Materials and methods7KLV�VWXG\�DQG�RWKHU�VWXGLHV�UHSRUWHG�KHUH�ZHUH�FDUULHG�RXW�IURP�6HSWHPEHU������XQWLO�$XJXVW�������4XDGUDWHV�PHDVXULQJ����P2�ZHUH�HUHFWHG�LQ�YDULRXV�ORFDWLRQV�WR�VDPSOH�GLYHUVH�KDELWDWV��WKH�VL]HV�ZHUH�DOWHUHG�DFFRUGLQJ�WR�VWXG\�ORFDOLW\����[���P2�LQ�PDQJURYH�IRUHVW�ÁRRU�DQG�PXGÁDW��DQG���[���RU���[����P2�RQ�ULYHU�EDQNV���2WKHU�PHWKRGV�IRU�FROOHFWLQJ�PXGVNLSSHUV�ZHUH�E\�XVLQJ�KDQG�RU�VFRRS�QHWV��FDVW�QHWV��GLJJLQJ�EXUURZV�DQG�SXUFKDVLQJ�IURP�ÀVKHUPHQ·V�WUDZO�FDWFK�

7KH�ÀVKHV�ZHUH�LGHQWLÀHG�XVLQJ�WKH�NH\V�RI�.RXPDQV���������.RWWHODW�et al.���������0XUG\��������DQG�0XUG\�DQG�7DNLWD���������:H�IROORZHG�0XUG\��������IRU�QRPHQFODWXUH��)LVKHV�ZHUH�NLOOHG�ZLWK�����IRUPDOLQ��7KH�IUHVKO\�NLOOHG�VDPSOHV�ZHUH�ZDVKHG�ZLWK�ZDWHU�DQG�GULHG�ZLWK�D�SLHFH�RI�FORWK��7KH�FRORXU�RI�HDFK�VSHFLPHQ�ZDV�UHFRUGHG�

9.4 Results6SHFLHV�ULFKQHVV�DQG�VSDWLDO�GLVWULEXWLRQ�RI�PXGVNLSSHUV7KLUWHHQ�VSHFLHV�IURP���JHQHUD�ZHUH�IRXQG�LQ�WKH�VWXG\�DUHDV��7DEOH�����7KH�GLVWULEXWLRQ�RI�PXGVNLSSHU�VSHFLHV�UHYHDOHG�GLVWLQFW�SUHIHUHQFHV�IRU�FHUWDLQ�KDELWDWV�

Periophthalmus chrysospilos Bleeker7\SLFDOO\��Ps. chrysospilos��)LJ�����KDV�PDUNHG�RUDQJH�VSRWV�RQ�RSHUFOHV�DQG�ÁDQNV��'��ÀQ�EULJKW�RUDQJH��GXVN\�VWULSHV�RQ�'��ÀQ�DQG�ZLWK�D�GLVWLQFW��URXQG�GLVN�SHOYLF�ÀQ��Ps.

chrysospilos�LV�WKH�RQO\�VSHFLHV�WKDW�FRXOG�EH�IRXQG�LQ�DOO�ORFDOLWLHV�RI�WKH�VWXG\�DUHDV��7KLV�VSHFLHV�ZDV�IRXQG�LQ�KLJK�GHQVLWLHV�MXVW�VKRUHZDUG�WR�WKH�ORZ�WLGH�OLQH��,W�LV�GRPLQDQW�RQ�WKH�PXGÁDWV�DQG�ULYHU�EDQNV�LQ�PRVW�VWXG\�VLWHV��7KH\�DUH�IRXQG�LQ�WKH�PDQJURYH�IRUHVW�ÁRRU�QHDUE\�WKH�HURGHG�EDQNV��EXW�QRW�IRXQG�ZLWKLQ�PDQJURYH�IRUHVWV�LQ�6HPHQWD�DQG�.XDOD�6HODQJRU��7KXV�WKH�VWXG\�VKRZHG�Ps. chrysospilos�ZDV�GRPLQDQW�LQ�SLRQHHU�PDQJURYH�]RQHV�RI�Avicennia alba�DQG�PL[HG�IRUHVW�]RQHV�RI�Avicennia alba and Sonneratia

alba��+RZHYHU��LW�RFFXUUHG�LQ�ORZ�DEXQGDQFH�RQ�WKH�HURGLQJ�EDQNV�

Periophthalmus spilotus 0XUG\�DQG�7DNLWD7KLV�LV�D�QHZ�VSHFLHV�GHVFULEHG�E\�0XUG\�DQG�7DNLWD���������7KLV�VSHFLHV�LV�VLPLODU�WR�DQG�SUREDEO\�PRVW�UHODWHG�WR�Ps. chrysospilos��)LJ������,W�KDV�PDUNHG�DQG�QXPHURXV�VPDOO�

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Table 1��7KH�GLVWULEXWLRQ�RI�PXGVNLSSHU�VSHFLHV�LQ�WKH�PDQJURYH�IRUHVW�ÁRRU��ULYHU�EDQNV�DQG�PXGÁDWV�RQ�WKH�6HODQJRU�FRDVW�

No. Species Mangrove Forest Floor � � 0DQJURYH� (URGHG�� 7UXH�� 5LYHU�%DQNV� 0XGÁDWV� � 3LRQHHU� %DQN� 0DQJURYH 1. Periophthalmus chrysospilos �� �� �� �� �2. Periophthalmus novemradiatus �� �� �� �� �3. Periophthalmus gracilis �� �� �� �� ���� Periophthalmus argentilineatus �� �� �� �� ���� Periophthalmus spilotus �� �� �� �� ���� Periophthalmus walailakae �� �� �� �� ���� Periophthalmodon schlosseri �� �� �� �� �8. Periophthalmodon septemradiatus �� �� �� �� ���� Boleophthalmus boddarti �� �� �� �� ����� Pseudapocryptes elongatus �� �� �� �� �11. Parapocryptes serperaster �� �� �� �� �12. Scartelaos histophorus �� �� �� �� �13. 2[XGHUFHV�GHQWDWXV� �� �� �� �� �

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ZKLWLVK�EOXH�VSRWV�RQ�WKH�ÁDQNV�DQG�RSHUFOHV��'��ÀQ�LV�UHGGLVK�EURZQ��'��ZLWK�VLQJOH�GXVN\�VWULSH��D�UHGGLVK�VSRW�RQ�SHFWRUDO�ÀQV��DQG�IUHQXP�RI�SHOYLF�ÀQV�DUH�SURPLQHQW�DQG�IXVHG�ZLWK�WKH�SHOYLF�ÀQV�LQWR�D�URXQG�GLVN��7KLV�VSHFLHV�ZDV�IRXQG�WR�EH�DEXQGDQW�LQ�WKH�KLJK�VKRUH�PDQJURYH�IRUHVWV�LQ�6HPHQWD�

Fig. 1. Periophthalmus chrysospilos Bleeker.

Fig. 2. Periophthalmus spilotus�0XUG\�DQG�7DNLWD�

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Periophthalmus walailakae 'DUXPDV�DQG�7DQWLFKRGRN7KLV�LV�DOVR�D�QHZ�VSHFLHV��)LJ�����GHVFULEHG�E\�'DUXPDV�DQG�7DQWLFKRGRN���������7KH�VSHFLHV�LV�FKDUDFWHULVHG�E\�D�VLQJOH�URZ�RI�WHHWK�LQ�XSSHU�MDZ�DQG�VWURQJ�IUHQXP�WKDW�IXVHV�ZLWK�WKH�SHOYLF�ÀQ��IRUPLQJ�D�URXQG�GLVN��7KH�FRORXUDWLRQ�RI�KHDG�DQG�ÁDQNV�LV�JUHHQLVK�EURZQ��ZKLWH�VSRWV�DUH�REYLRXVO\�VHHQ�VFDWWHUHG�RQ�ÁDQNV��SHFWRUDO�ÀQV�DQG�RSHUFOHV��'��UHGGLVK�EURZQ��'��ZLWK�VLQJOH�GXVN\�VWULSH��SHFWRUDO�DQG�FDXGDO�ÀQ�VSLQHV�DUH�GXVN\�DQG�SHOYLF�ÀQV�WRWDOO\�IXVHG��IRUPLQJ�D�URXQG�GLVN��7KLV�VSHFLHV�OLYHV�V\PSDWULFDOO\�ZLWK�Ps. novemradiatus and Ps. gracilis�LQ�WKH�PL[HG�IRUHVW�RI�Sonneratia alba and Rhizophora mucronata�LQ�0RULE��7ZR�ELJ�VSHFLPHQV��!�����PP�6/��ZHUH�FDXJKW�LQ�WKH�XSSHU�VKRUH�PDQJURYHV�LQ�6HPHQWD��7KH�FRPPRQ�PXGVNLSSHUV�LQ�WKH�KDELWDW�ZHUH�Ps. novemradiatus, Ps. gracilis, Ps. spilotus, Pn schlosseri and B. boddarti.

Fig. 3. Periophthalmus walailakae�'DUXPDV�DQG�7DQWLFKRGRN�

Fig. 4. Periophthalmus gracilis Eggert.

Periophthalmus gracilis Eggert0RUSKRORJLFDOO\��Ps. gracilis� �)LJ�� ��� LV�GHWHUPLQHG�E\� WKH� DEVHQFH� RI� IUHQXP�� WKH�SHOYLF�ÀQV�DUH�HQWLUHO\�VHSDUDWHG�DQG�WKH�ÀUVW�GRUVDO�ÀQ�LV�UHODWLYHO\�VKRUW��FRQYH[�DQG�URXQGHG�ZLWK�QR�HORQJDWHG�VSLQH��'RUVDO�ÀQV�DUH�QRW�FRQQHFWHG��'��VKRUW��JUH\�ZLWK�EODFN�VSRWV�DQG�EODFN�VWULSH�WKDW�HQG�LQWR�D�ODUJH�EODFN�VSRW��'��HOHPHQWV�JUH\�ZLWK�D�VLQJOH�EODFN�SDOH�VWULSH��3HFWRUDO�DQG�FDXGDO�ÀQV�KDYH�HOHPHQWV�RI�JUH\LVK�EURZQ��$QDO�ÀQ�HOHPHQWV�DUH�VKRUW�DQG�GXVN\�\HOORZ��'RUVDO�EORWFKHV�JUH\�EXW�QRW�YHU\�REYLRXV��7KH�EODFN�VSRW�DQG�WKH�YHUWLFDO�ZKLWLVK�EOXH�VWULSH��FRPSRVHG�RI�VPDOO�VSRWV�ZHUH�XVHG�WR�GLIIHUHQWLDWH�WKH�Ps. gracilis IURP�Ps. novemradiatus. Ps. gracilis�LV�UHVWULFWHG�WR�PRUH�LQODQG�PDQJURYH�IRUHVWV�

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Fig. 5. Periophthalmus novemradiatus��+DPLOWRQ��

Periophthalmus novemradiatus��+DPLOWRQ�Ps. novemradiatus��)LJ�����LV�FKDUDFWHULVHG�E\�WKH�GLVWLQFW�VHSDUDWLRQ�RI�SHOYLF�ÀQV�DQG�WKH�H[LVWHQFH�RI�D�VPDOO�SHOYLF�IUHQXP��ZKLFK�LV�YLVLEOH�ZLWKRXW�PDJQLÀFDWLRQ��'��KHLJKW�PRGHUDWH��JUH\�ZLWK�UHGGLVK�FRORXUDWLRQ�RQ�WLSV�RI�VSLQHV��SDOH�PDUJLQ�DQG�D�VLQJOH�EODFN�VWULSH�RQ�HOHPHQWV�DQG�VRPH�EODFN�VSRWV�LQ�WKH�SRVWHULRU��'��JUH\�ZLWK�EODFN�SDOH�VWULSH�DQG�VRPH�UHGGLVK�VSRWV��%RWK�'��DQG�'��DUH�QRW�FRQQHFWHG��&DXGDO�DQG�SHFWRUDO�ÀQ�HOHPHQWV�DUH� UHGGLVK�EURZQ�ZLWK�EURZQ�VSRWV��$QDO�ÀQ�HOHPHQWV�DUH� VKRUW�DQG�WUDQVSDUHQW�LQ�FRORXU��7KLV�VSHFLHV�LV�GRPLQDQW�DQG�ZLGHO\�GLVWULEXWHG�LQ�WKH�PDQJURYH�IRUHVW�ÁRRU��7KHLU�FDUQLYRURXV�QDWXUH��IHHGLQJ�PDLQO\�RQ�LQVHFWV�DQG�FUXVWDFHDQV��OHDG�WKHP�WR�IRUDJH�DZD\�IURP�WKHLU�UHVWULFWHG�KDELWDWV�DQG�LQWR�DTXDFXOWXUH�VKULPS�SRQGV�

Periophthalmus argentilineatus�9DOHQFLHQQHV7KLV�VSHFLHV�ZDV�IRXQG�E\�6ZHQQHQ�et al����������LQ�WKH�IRUHVW�ÁRRU�QHDU�WKH�VWUHDPV�DQG�HVWXDU\�RI�3DWWDQL��7KDLODQG��,Q�WKLV�VWXG\��Ps. argentilineatus��)LJ�����ZDV�IRXQG�LQ�the Sonneratia alba and Rhizophora mucronata�]RQHV�RI�0RULE��ZKHUH�WKH�VXEVWUDWHV�ZHUH�VDQG\�PXG��7KH�PXGVNLSSHUV�ZHUH�IRXQG�FOLQJLQJ�RQWR�WKH�PDQJURYH�VWHPV��7KH\�FDQ�EH�GLVWLQJXLVKHG�E\�WKH�UHG�PDUNLQJV�ZLWK�D�ZLGH�EODFN�VWULSH�DQG�VPDOO�ZKLWH�VSRWV�RQ�'��'��PDUJLQ�UHG�ZLWK�D�ZLGH�EODFN�VWULSH�DQG�ODFN�VSRW��VPDOO�ZKLWLVK�EOXH�VSRWV�RQ�RSHUFOH��SHOYLF�ÀQV�ZHUH�WRWDOO\�VHSDUDWHG�DQG�QR�SHOYLF�IUHQXP�ZDV�REVHUYHG��7KH�VSHFLHV�ZDV�GLVWLQJXLVKHG� IURP�Ps. gracilis� E\� WKH� FRORXUDWLRQ�RI�'��ÀQ� �Ps. gracilis KDYLQJ�'��VKRUW��JUH\�ZLWK�EODFN�VSRWV��DQG�IURP�Ps. novemradiatus�E\�WKH�DEVHQFH�RI�SHOYLF�IUHQXP��SHOYLF�IUHQXP�LQ�Ps. novemradiatus�LV�SURPLQHQW��

Fig. 6. Periophthalmus argentilineatus�9DOHQFLHQQHV��6/�����PP�

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spilotus, Ps. ranongensis and B. boddarti��$GXOWV�ZHUH�PRUH�FRPPRQ�RQ�PXGÁDWV�DQG�ULYHU�EDQNV�WKDQ�LQ�WKH�PDQJURYH�IRUHVW��7KLV�VSHFLHV�LV�D�FDUQLYRUH��IRUDJLQJ�RQ�VPDOO�FUDEV�DQG�VPDOO�ÀVK�

Fig. 7. Periophthalmodon schlosseri��3DOODV��

Periophthalmodon septemradiatus��+DPLOWRQ�7KHUH�DUH�QXPHURXV�UHG�DQG�SDOH�EOXH�VSRWV�VFDWWHUHG�RQ�VQRXW��RSHUFOH�DQG�WUXQN��1R�SHOYLF�IUHQXP�DQG�SHOYLF�ÀQV�FRPSOHWHO\�VHSDUDWHG��7HHWK�RI�XSSHU�MDZ�LQ�WZR�URZV��DQWHULRU�WHHWK�RI�RXWHU�MDZ�VPDOO�FDQLQRLG�DQG�VPDOOHU�LQ�WKH�LQQHU�URZ��'RUVDO�ÀQV��LQ�IHPDOH��'��SRRUO\�GHYHORSHG�SRVVHVVLQJ�������VKRUW�VSLQHV�DQG��LQ�VRPH�VSHFLPHQV��DUH�QHJOLJLEOH��,Q�PDOH��'��LV�EODFN�ZLWK�UHGGLVK�PDUJLQ��KHLJKW�PRGHUDWH�ZLWK�ÀUVW�VSLQH�HORQJDWH��FRQFDYH�DQG�VHFRQG�GRUVDO�ÀQ��'���GXVN\�ZLWK�D�UHG�PDUJLQ��'��DQG�'��DUH�FRQWLJXRXV��7KH�'��VKDSH�FKDUDFWHULVHG�VH[XDO�GLPRUSKLVP��&DXGDO�DQG�SHFWRUDO�ÀQV�JUH\LVK�LQ�FRORXU�DQG�QXPHURXV�UHG�DQG�EOXH�VSHFNOHV�RQ�SHFWRUDO�ÀQV��$QDO�DQG�SHOYLF�ÀQV�DUH�GXVN\��)LJ������7KLV�VSHFLHV�ZDV�IRXQG�LQ�WKH�VPDOO�VWUHDPV�RI�WKH�6HODQJRU�ULYHU�EDQN�LQ�.DPSXQJ�.XDQWDQ��ZKHUH�WKH�VDOLQLW\�UHDFKHG���SSW�GXULQJ�WKH�ORZ�WLGHV�

Fig. 8. Periophthalmodon septemradiatus��+DPLOWRQ���6/�����PP�

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Fig. 9. Boleophthalmus boddarti��3DOODV���6/������PP�

Fig. 10. Scartelaos histophorus��9DOHQFLHQQHV��

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Fig. 13. Parapocryptes serperaster��5LFKDUGVRQ���6/������PP�

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novemradiatus, Ps. gracilis, Ps. spilotus, Ps. argentilineatus, Ps. walailakae, B. boddarti, and Pn. schlosseri.

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B. pectinirostris, B. dussumieri and Pss. borneensis�DOWKRXJK�WKHVH�ZHUH�UHSRUWHG�E\�RWKHU�DXWKRUV�IURP�ORFDOLWLHV�RXWVLGH�WKH�SUHVHQW�VWXG\�DUHDV�LQ�3HQLQVXODU�0DOD\VLD��7DEOH�����Ps. malaccencis�ZDV�GLVFRYHUHG�LQ�.XDOD�/DQJDW�E\�.RXPDQV��������DQG�B. pectinirostris ZDV� UHSRUWHG� WR� H[LVW� LQ�3HQDQJ�DFFRUGLQJ� WR� WKH�ROG� UHFRUGV�RI�&DQWRU� ������� DQG�.RXPDQV� ��������$OWKRXJK�Pss. borneensis�ZDV� UHSRUWHG� IURP�3HQDQJ�DQG�6DUDZDN�E\�.RXPDQV���������0XUG\��������RQO\�UHGLVFRYHUHG�WKLV�VSHFLHV�LQ�6DUDZDN��$�PRUH�UHFHQW�ZRUN�E\�7DNLWD�et al���������FDUULHG�RXW�LQ�.DPSXQJ�6XQJDL�<X��.XDOD�6HODQJRU�added B. dussumieri to the list.

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1R�� 6SHFLHV� &DQWRU� .RXPDQV� %HUU\� 0XUG\� 7DNLWD�et al. Present Study (1849) (1953) (1972) (1989) (1999) (Khaironizam & Norma-Rashid)1. Ps. chrysospilos �� �� �� �� �� �2. Ps. novemradiatus �� �� �� �� �� �3. Ps. gracilis �� �� �� �� �� ���� Ps. argentilineatus �� �� �� � � �� ���� Ps. spilotus �� �� �� �� �� ���� Ps. malaccencis �� ����������������������������������� �� �� �� �� � � ��.��/DQJDW���� Ps. walailakae �� �� �� �� �� �8. Pn. schlosseri ������������������� �� �� �� �� �� � �3HQDQJ���� Pn. septemradiatus �� �� �� � � �� ����� B. boddarti ������������������ �� �� �� �� �� � �3HQDQJ�11. B. pectinirostris ���������������� ����������������������� �� � � �� �� � �3HQDQJ�� ��3HQDQJ�12. B. dussumieri �� �� �� �� �� �13. Pss. elongatus ��� ��� �� �� �� �� � �3HQDQJ�� �3HQDQJ�� ����� Pss. borneansis �� ������������������������ �� � � �� �� � � ��3HQDQJ�� ����� Pa. serperaster �� �� �� �� �� ����� S. histophorus �� �� �� �� �� �� � �3HQDQJ�� �3HQDQJ������ 2��GHQWDWXV� �� ��� �� �� �� �� � ��3HQDQJ�

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Ps. argentilineatus��IRPHUO\�NQRZQ�DV�Ps. vulgaris��ZDV�HDUOLHU�UHSRUWHG�WR�LQKDELW�WKH�VWUHDP�EDQNV�ZLWKLQ�WKH�PDQJURYH�IRUHVW��%HUU\���������,W�ZDV�DOVR�UHSRUWHG�WR�SRSXODWH�

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Ps. chrysospilos�LV�WKH�PRVW�DEXQGDQW�VSHFLHV�RQ�WKH�LQWHUWLGDO�PXGÁDW��ULYHUEDQNV�DQG�LQ�WKH�PDQJURYH�SLRQHHU�DUHDV�RI�-HUDP�DQG�0RULE��$W�KLJK�WLGHV�LW�PD\�EH�IRXQG�FOLQJLQJ�RQWR�WKH�PDQJURYH�VWHPV�RU�GHDG�WUXQNV�

Pa. serperaster�VHHPV�WR�SUHIHU�VXEWLGDO�KDELWDWV�LQ�HVWXDULHV��6SHFLPHQV�LQ�WKLV�VWXG\�DQG�WKRVH�RI�7DNLWD�et al.��������ZHUH�FDXJKW�ZLWK�WUDZO�QHW�RU�FDVW�QHW��6DPSOHV�ZHUH�FDXJKW�LQ�WKH�ÀVKHUPHQ·V�WUDZO�DV�IRXQG�LQ�WKLV�VWXG\�DQG�WKDW�RI�7DNLWD�et al.���������7KH�SUHVHQFH�RI�Pss. elongatus�LQ�WKH�FDQDOV�QHDU�VKULPS�SRQGV�DQG�LQ�WKH�GUDLQV�RI�SDOP�RLO�HVWDWHV�FRXOG�EH�GXH�WR�LWV�IRUDJLQJ�KDELWV�LQ�H[SORLWLQJ�HDVLO\�DFFHVVLEOH�IRRG��+RUD��������IRXQG�WKDW�Pss. elongatus�ZDV�DEOH�WR�ZLWKVWDQG�GURXJKW�DQG�WRUSLG�ERWWRP�RI�GULHG�XS�SRROV��7KH�QDWXUDO�KDELWDW�RI�WKLV�VSHFLHV�DV�UHSRUWHG�E\�6ZHQQHQ��et al��������ZDV�EDUH�PXGÁDW�DUHDV�ZLWK�QR�WLGDO�LQÁXHQFH�

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References%HUU\��$�-���������7KH�QDWXUDO�KLVWRU\�RI�:HVW�0DOD\VLDQ�PDQJURYH�IDXQDV��Malayan

Nature Journal 25����������&DQWRU��7�(���������&DWDORJXH�RI�0DOD\DQ�ÀVKHV� Journal of the Asiatic Society of Bengal

18(2)���������&OD\WRQ��'�$���������0XGVNLSSHUV��2FHDQRJUDSK\�DQG�0DULQH�%LRORJ\�$QQXDO�5HYLHZ 31��

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�*RELLGDH�2[XGHUFLQDH��IURP�VRXWKHUQ�7KDLODQG��Phuket Marine Biological Center

Research Bulletin. 64����������)DULGDK�+DQLP��$���������%HEHUDSD�DVSHN�HNRORJL�LNDQ�WHPEDNXO�GL�3DQWDL�0RULE��%6F��

7KHVLV��(FRORJ\���'HSDUWPHQW�RI�=RRORJ\��8QLYHUVLW\�RI�0DOD\D��.XDOD�/XPSXU����SS�

*RUGRQ��0�6���%RHWLXV��-���(YDQV��'�+��DQG�2JOHVE\��/�&���������$GGLWLRQDO�REVHUYDWLRQV�RQ�WKH�1DWXUDO�+LVWRU\�RI�WKH�PXGVNLSSHU��Periophthalmus sobrinus. &RSLHD���������������

+DP]DK��0���������.HODNXDQ�Periophthalmus chrysospilos�GL�KXWDQ�EDNDX��%6F��7KHVLV��(FRORJ\���'HSDUWPHQW�RI�=RRORJ\��8QLYHUVLW\�RI�0DOD\D��.XDOD�/XPSXU����SS�

+RUD��6�/���������(FRORJ\�DQG�ELRQRPLFV�RI�WKH�JRELRLG�ÀVKHV�RI�WKH�*DQJHWLF�'HOWD��International Congress of Zoology 12����������

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.KDLURQL]DP��0�=�� DQG�1RUPD�5DVKLG�� <��� ������$� QHZ� UHFRUG� RI�PXGVNLSSHU�Periophthalmus spilotus��*RELLGDH��2[XGHUFLQDH��IURP�6HODQJRU�FRDVWV��3HQLQVXODU�0DOD\VLD��Natural History Bulletin of Siam Society 50(2)���������

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