occurrence of fishes at 38 sites in the mississippi ... · information for 38 fish sampling sites...

Occurrence Of Fishes At 38 Sites in the Mississippi Embayment Study Unit, 1996-98

By B.G. Justus and B.J. Caskey

U.S. Geological Survey

Open-File Report 99-605

National Water-Quality Assessment Program

Pearl, Mississippi

2000

U.S DEPARTMENT OF THE INTERIOR

BRUCE BABBITT, Secretary

U.S. GEOLOGICAL SURVEY

Charles G. Groat, Director

Any use of trade, product, or firm names is for identification purposes only and does not constitute endorsement by the U.S. Geological Survey.

For additional information write to:

District Chief U.S. Geological Survey 308 South Airport Road Pearl, MS 39208-6649

Copies of this report can be purchased from:

U.S. Geological Survey Branch of Information Services Box 25286 Denver, CO 80225-0286

Information regarding the National Water-Quality Assessment (NAWQA) Program is available on the Internet via the World Wide Web. You may connect to the NAWQA Home Page using the Universal Resource Locator (URL) at:

http://wwwrvares.er.usgs. gov/nawqa/nawqa_home.html

FOREWORD

The mission of the U.S. Geological Survey (USGS) is to assess the quantity and quality of the earth resources of the Nation and to provide informa tion that will assist resource managers and policymak- ers at Federal, State, and local levels in making sound decisions. Assessment of water-quality conditions and trends is an important part of this overall mission.

One of the greatest challenges faced by water- resources scientists is acquiring reliable information that will guide the use and protection of the Nation's water resources. That challenge is being addressed by Federal, State, interstate and local water-resource agencies and by many academic institutions. These organizations are collecting water-quality data for a host of purposes that include: compliance with permits and water-supply standards; development of remedia tion plans for a specific contamination problem; opera tional decisions on industrial, wastewater, or water- supply facilities; and research on factors that affect water quality. An additional need for water-quality infor mation is to provide a basis on which regional and national-level policy decisions can be based. Wise decisions must be based on sound information. As a society we need to know whether certain types of water-quality problems are isolated or ubiquitous, whether there are significant differences in conditions among regions, whether the conditions are changing over time, and why these conditions change from place to place and over time. The information can be used to help determine the efficacy of existing water-quality policies and to help analysts determine the need for and likely consequences of new policies.

To address these needs, the Congress appropri ated funds in 1986 for the USGS to begin a pilot pro gram in seven project areas to develop and refine the National Water-Quality Assessment (NAWQA) Pro gram. The NAWQA Program builds upon an existing base of water-quality studies of the USGS, as well as those of other Federal, State, and local agencies. The objectives of the NAWQA Program are to:

Describe current water-quality conditions for a large part of the Nation's freshwater streams, rivers, and aquifers.

Describe how water quality is changing over time.

Improve understanding of the primary natural and human factors that affect water-quality conditions.

This information will help support the development and evaluation of management, regulatory, and monitoring decisions by other Federal, State, and local agencies to protect, use and enhance water resources.

The goals of the NAWQA Program are being achieved through ongoing and proposed investigations of more than 50 of the Nation's most important river basins and aquifer systems, which are referred to as study units. These study units are distributed through out the Nation and cover a diversity of hydro-geologic settings. More than two-thirds of the Nation's freshwa ter use occurs within the more than 50 study units, and more than two-thirds of the people served by public water-supply systems live within their boundaries.

National synthesis of data analysis, based on aggregations of comparable information obtained from the study units, is a major component of the program. This effort focuses on selected water-quality topics using nationally consistent information. Comparative studies will explain difference and similarities in observed water-quality conditions among study areas and will identify changes and trends and their causes. The first topics addressed by the national synthesis are pesticides, nutrients, volatile organic compounds, and aquatic biology. Discussions on these and other water- quality topics will be published in periodic summaries of the quality of the Nation's ground and surface water as the information becomes available.

This report is an element of the comprehensive body of information developed as part of the NAWQA Program. The program depends heavily on the advice, cooperation, and information from many Federal, State, interstate, Tribal, and local agencies and the public. The assistance and suggestions of all are greatly appreciated.

Robert M. Hirsch Chief Hydrologist

iii

CONTENTSPage

Abstract.................................................................................................................................................... 1

Introduction............................................................................................................................................. 1

Purpose and Scope................................................................................................................................... 1

Acknowledgments................................................................................................................................... 2

Site selection criteria and site information.............................................................................................. 2

Methods of sampling and processing ...................................................................................................... 2

Results..................................................................................................................................................... 3

References............................................................................................................................................... 4

ILLUSTRATION

Figure 1. Map showing locations of 38 sites sampled in the Mississippi Embayment during 1996-98 5

TABLES

Table 1. Information for 38 fish sampling sites in the Mississippi Embayment Study Unit............. 6

2. Scientific and common names of fishes collected from 38 sites in the Mississippi

Embayment Study Unit, 1996-98 ..................................................................................... 7

3. Number of individuals for fishes collected at 38 sites in the Mississippi Embayment

Study Unit, 1996-98.......................................................................................................... 9

4. Percent relative abundance of fishes collected at 38 sites in the Mississippi Embayment

Study Unit, 1996-98.......................................................................................................... 17

CONVERSION FACTORS AND ABBREVIATIONS

Divide By To obtain

cubic meter per second (m3/s)

centimeter (cm)

meter (m)

meter per second (m/s)

f\

square kilometer (km)

0.02832

0.4

3.28

3.28

2.59

cubic foot per second (ft3/s)

inch (in.)

foot (ft)

foot per second (ft/s)

square mile (mi2)

The following acronyms are used in this report:

MAP Mississippi Alluvial PlainMISE Mississippi Embayment Study UnitNAWQA National Water-Quality AssessmentUSGS U.S. Geological Survey

VI

Occurrence of Fishes at 38 Sites in the Mississippi Embayment Study Unit, 1996-98

By B.C. Justus and B.J. Caskey

Abstract

Fishes were sampled at 38 sites (58 sampling reaches) in the Mississippi Embayment Study Unit in parts of Arkansas, Kentucky, Louisiana, Mississippi, Missouri, and Tennessee from 1996 through 1998 as part of the National Water-Quality Assessment Program. Fishes were collected by seining and electrofishing during low- flow periods from late spring to late summer. All fishes collected were identified to the lowest possible taxon (usually species), weighed, measured, and examined for anomalies. A total of 95 taxa, representing 94 species and 18 families, were collected during the study. All 94 species were known to occur in the study area. The number of species collected at each of the sampling reaches ranged from 13 to 37.

INTRODUCTION

In 1994, the U.S. Geological Survey (USGS) began a study of the Mississippi Embayment Study Unit (MISE) as part of the National Water-Quality Assessment (NAWQA) Program (fig. 1). The NAWQA Program is designed to assess status and trends in the quality of the Nation's water resources and to determine the natural and human factors affecting these resources (Hirsch and others, 1988). The Program will eventually integrate physical, chemical, and biological data from more than 50 study units across the Nation.

Fish community structure is one aspect of the biological data collected as part of the NAWQA Program. Most of the MISE overlies the Mississippi Alluvial Plain (MAP) Ecoregion, an area where fish communities have not been intensively sampled.

The MAP Ecoregion has been extensively altered by human activity; about 75 percent (or about 16 million acres) of the original forested wetlands has been cleared and drained (Nature Conservancy, 1992) and most streams in the MAP Ecoregion have some degree of channel and hydrological modifications. Land use is dominated by agriculture (85.1 percent) and is used for growing row crops, such as cotton, corn, and soybeans, and small grains, such as rice and wheat.

The climate of the MISE is characterized as humid, with southern parts being subtropical and northern parts being temperate. Mean annual temperatures range from about 64 °F in the south to about 57 °F in the north. Annual precipitation ranges from about 56 inches in the south to about 48 inches in the north (U.S. Department of Commerce, 1995).

Purpose and Scope

This report documents the occurrence of fishes sampled at 38 sites in the MISE (36 of which are in the MAP Ecoregion) from 1996 to 1998. These and other ecological data collected by NAWQA can be used to help describe water-quality conditions and improve understanding of the factors that affect water quality.

INTRODUCTION 1

Acknowledgments

The authors would like to acknowledge three museum curators for their willingness to verify identification of specimens: Dr. Neil Douglas at the University of Louisiana at Monroe; Dr. Todd Slack at the Mississippi Museum of Natural Sciences, and Dr. Steve Ross at the University of Southern Mississippi. Additional thanks are extended to Dr. Neil Douglas and Dr. Todd Slack for archiving museum specimens.

SITE SELECTION CRITERIA AND SITE INFORMATION

In 1995, eight sites were selected for which fish community sampling was planned from 1996 to 1998. The eight sites (which were located on seven streams in Arkansas, Kentucky, Louisiana, Mississippi, Missouri, and Tennessee, and within the MISE boundary) were selected because they either integrated most types of land use in the MISE or were representative of a specific land use in the MISE, collectively provided spatial coverage of the MISE, and had long-term gage records. Six of the eight sites and five of the seven streams are located in the MAP Ecoregion; of the two remaining sites, one site is located in the Mississippi Valley Loess Plains Ecoregion, and the other is located in the Southeastern Plains Ecoregion. As planned, fish communities were sampled annually from 1996 to 1998 at the eight sites.

In 1998, fish communities also were sampled at 30 sites that had not been previously sampled by NAWQA (table 1). These 30 sites were added to increase spatial coverage of the MAP Ecoregion. Two criteria were used for selection of these 30 additional sites: first, ten sites were chosen to represent a gradient of crop intensity for each of three major crops grown in the MAP Ecoregion~corn, rice, and cotton. Secondly, as with the first eight sites sampled, the remaining sites were selected that provided the best spatial coverage of the MAP Ecoregion. County-level land use information for 1995 and 1996 was used to

determine crop intensities. Photographs and maps showing the sampling locations at each of the 38 sampling sites can be viewed at

<http://ms.water.usgs.gov/misenawqa/>.

METHODS OF SAMPLING AND PROCESSING

Stream reaches, or lengths of the stream where sampling was to be done, were designated at each of the 38 sites before sampling. To designate reaches, a visual assessment of the stream was made, and lengths physically representative of the stream were measured and marked for sampling. A reach of stream 500-m long was selected for sampling at 36 sites; samples were collected from a 250-m reach at each of two small streams (LaGrue Bayou near Dewitt, Arkansas, and Second Creek near Palestine, Arkansas) because fallen trees and beaver dams restricted access.

Three separate 500-m reaches were sampled at two of the eight sites sampled in 1996 (the Bogue Phalia at Leland, Mississippi, and the Cache River at Cotton Plant, Arkansas) to assess sampling variability; consequently, fish were sampled from 12 stream reaches that year. Data from these samples are included in table 3 and 4.

In 1997 and 1998, only one reach was sampled at each of the two sites where three reaches were sampled in 1996. Because the three reaches at each of the two sites were physically similar, accessibility was the main consideration for selecting which reach would be sampled for both years. At the Bogue Phalia at Leland, Mississippi, the most upstream reach (reach A) was sampled. At the Cache River at Cotton Plant, Arkansas, the most downstream reach (reach C) was sampled. Only one reach was sampled at each of the 36 remaining sites in 1998.

Fishes were collected by seining and electrofishing during low-flow periods from late spring to late summer. In general, methods were consistent with NAWQA fish

2 OCCURRENCE OF FISHES AT 38 SITES IN THE MISSISSIPPI EMBAYMENT STUDY UNIT, 1996-98

sampling protocols (Meador and others, 1993).

Seining consisted of sampling available habitats with a 4-m x 2-m seine having a mesh size of 0.5 cm. In streams having some areas that were wadeable, seining was done by two people wading and pulling the seine through the water. Areas that were unwadeable but near-shore were seined by two people standing facing each other about 2 m apart at the edge of the water. Each person held the seine by the top of one of the seine-poles with the bottom of the poles resting on the bank about 1.5 m from the edge of water. On a signal, the seine was swung overhead, with the lead-line on the outer edge of the arc, and into the water as far away from the bank as possible. After the lead-line sank to the stream bottom, the seine was retrieved. The time spent seining at each site was approximately 45 minutes, and the number of seining efforts (hauls) ranged from 6 to 9. After each seine haul, specimens were placed in 10 percent formalin. All fishes collected by seining were taken to a lab at the USGS office in Pearl, Mississippi, where they were identified to the lowest possible taxon (usually species), weighed, measured, and examined for anomalies.

At 57 of the 58 reaches sampled, both banks of the sampling reach were sampled by electrofishing (one bank at the Skuna River was not sampled in 1996 because of equipment problems). Electrofishing was done using a 4.7-m X 1-m aluminum boat with a commercially manufactured electrofishing unit consisting of a 2,500-watt generator and a pulsator.

An electrofishing team consisted of a boat driver and a person positioned on the bow of the boat to net fish. For sites where the water was clear enough that samplers could see fish habitat (submerged woody debris was the dominant habitat type), the boat was maneuvered downstream and along the bank until the electrodes were positioned near habitat, at which time electrofishing began. At sites that lacked fish habitat or where fish habitat was not visible due to turbid water, the boat was maneuvered in the

same manner as above but with the electrofishing gear in continual use.

Samples from each side of the stream were collected and processed independently of each other. Electrofishing time averaged about 21 minutes per bank sample. Fish large enough were identified to the lowest taxon possible (usually species), and were weighed, measured, examined for anomalies, and then released. At sites where there were large numbers of fish, the time that the fish were out of water (and fish mortality) was reduced by subdividing the sampling reach and processing fish from each subreach immediately after they were collected. Fish that were too small for positive identification, or too small to be weighed and measured in the field, were preserved in 10 percent formalin and taken to the USGS office in Pearl, Mississippi for processing.

Standard quality-assurance and quality- control procedures were taken to ensure that the fish data were of high quality (Walsh and Meador, 1998). Common and scientific names reported were those established by the American Fisheries Society's Committee on Names of Fishes (Robins and others, 1991). Although most identifications were made by the study unit biologist; some individuals were of a size or species that made them challenging to identify. To ensure data quality, those specimens were also identified by curators of fish museums in Louisiana and Mississippi. Voucher specimens are stored in fish collections at two museums: the Museum of Natural Sciences in Jackson, Mississippi; and the Museum of Natural History (Zoology) at the University of Louisiana at Monroe in Monroe, Louisiana.

RESULTS

A list of the fishes collected from 1996 to 1998 is provided in table 2. A total of 95 fish taxa representing 94 species and 18 families were collected at the 38 sites (table 3). The literature indicates that all 94 species were known to occur in the study area (Douglas, 1974; Pfleiger, 1975;

RESULTS 3

Robison and Buchanan, 1988; Etnier and Starnes, 1993). The number of taxa collected at the sites for all sampling dates ranged from 13 at Bogue Phalia near Leland, Mississippi in 1996 to 37 at Little River Ditch near Morehouse, Missouri in 1996. The number of individuals collected on all sampling dates is provided; however, it should be noted that immature or damaged individuals were not identified beyond the family or genus level (table 3). Data in table 3 were used to calculate percent relative abundance of fishes at all taxonomic levels on all sampling dates (table 4).

REFERENCES

Douglas, N.H., 1974, Freshwater fishes of Louisiana: Claitor's Publishing Division, Baton Rouge, La., 443 p.

Etnier, D.A., and Starnes, W.C., 1993, The fishes of Tennessee: The University of Tennessee Press, Knoxville, Tenn., 681 p.

Hirsch, R.M., Alley, W.M., and Wilbur, W.G., 1988, Concepts for a National Water-Quality Assessment Program: U.S. Geological Survey Circular 1021, 42 p.

Meador, M.R., Cuffney, T.F., and Gurtz, M.E., 1993, Methods for sampling fish communities as part of the National Water-Quality Assessment Program: U.S. Geological Survey Open-File Report 93-104,40 p.

Nature Conservancy, 1992, The forested wetlands of the Mississippi River: An ecosystem in crisis, The Nature Conservancy, Baton Rouge, LA, 25 p.

Pflieger, W.L., 1975, The fishes of Missouri: Missouri Department of Conservation, 343 p.

Robins, C.R., Bailey, R.M., Bond, C.E., Brooker, J.R. Lachner, E.A., Lea, R.N., and Scott W.B., 1991, Common and scientific names of fishes from the United States and Canada: Bethesda, Md., American Fisheries Society, Special Publication 20,183 p.

Robison, H.W., and Buchanan, T.M., 1988, Fishes of Arkansas (5th ed.): The University of Arkansas Press, Fayetteville, Ark., 536 p.

U.S. Department of Commerce, 1995,Monthly station normals of temperature, precipitation, and heating and cooling degree days 1961-1990: Climatography of the United States No. 81, National Oceanic and Atmospheric Administration, National Climatic Data Center, Asheville, North Carolina.

U.S. Geological Survey, 1999,http://ms.watcr.usgs.gov/misenawqa. Homepage to the Mississippi Embayment National Water-Quality Assessment Program.

Walsh, S.J., and Meador, Michael R., 1998, Guidelines for quality assurance and quality control of fish taxonomic data collected as part of the National Water- Quality Assessment Program: U.S. Geological Survey Open-File Report 98-4239, 33 p.


ILLINOIS

KENTUCKY

TENNESSEE

ARKANSAS MISSISSIPPI

LOUISIANA

EXPLANATIONMississippi Embayment Study Unit

Mississippi Alluvial Plain Ecoregion

Streams

Base fromU.S. Geological Survey digital data, 1:2,000,000 Geographic projection

(3J>) Sampling site and number

0 40 MILES

0 40 KILOMETERS

Figure 1. Locations of 38 sites sampled in the Mississippi Embayment Study Unit during 1996-98.

Tabl

e 1.

Inf

orm

atio

n (li

sted

by

site

num

ber)

for

38 fi

sh s

ampl

ing

site

s in

the

Mis

siss

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baym

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dy U

nit [

km2,

squa

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3/s, c

ubic

met

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m

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C/3 rn C/3

U O)

C/3 1 Z ^ rn c/5 C/3 w

w TJ 2 m > m H C/3

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No.

of

Site

nam

e S

ite

time

s S

tatio

n nu

mbe

r sa

mpl

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num

ber

St.

John

s D

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near

Sik

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n, M

OLi

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Riv

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no.

1 ne

ar M

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, M

OS

pillw

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at H

wy

102

near

Eas

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MO

Littl

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Ditc

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1 ne

ar L

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m,

MO

Obi

on C

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nea

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an,

KY

Mai

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at H

wy

153

near

Whi

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MO

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t H

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103,

TN

Elk

Chu

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onet

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Fran

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R

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flow

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rk,

MS

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bel

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1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38

1 3 1 1 1 1 1 1 1_

_ 1 3 1 1 1 1 1 3 1 5 1 1 1 1 1 3 1 1 5 1 1 1 1 1 1 3 3 1 1

0704

3300

0704

3500

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4160

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2500

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3800

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1120

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7050

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6515

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0496

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7380

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4660

0704

7700

0704

7520

0707

7700

0704

7947

0703

0392

0704

7950

0707

7555

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7950

0707

8040

0727

9950

0726

5099

0728

0900

0728

3000

0728

8570

0728

8500

0728

8650

0736

7659

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2887

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7700

0728

8700

0728

8870

0728

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8955

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0737

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3656

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3644

5436

3320

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3610

1835

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3522

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3323

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5818

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8921

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3036

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9056

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9028

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3436

9106

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9054

4089

1448

9053

1091

1919

9050

44

9116

5790

1557

9131

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2028

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9024

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3235

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4791

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2830

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(k

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1,14

418

662

778

4

356

751

218

146

6,15

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1,81

641

01,

367

13,7

741,

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111

543

1,98

32,

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1,16

0

594

4,93

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536

668

651

2,01

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301

376

231

1,82

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1,12

247

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721

2,14

11,

311

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cent

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sin

in

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lture

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char

ge

at g

age

(m3/

s)2.

222.

960.

933.

601.

22

2.20

0.68

0.70

3.31

11.0

8.84

4.71

4.59

36.1

4.61

2.52

5.04

8.69

14.6

6.33

0.00

90.7

0.00

2.41

0.49

5.77

15.0

5.35

9.80

0.00

6.17

46.0

2.88

0.00

405

2.35

6.80

1.56

Mea

n ch

anne

l w

idth

(m

)10

.933

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.5

19.9

13.9

11.7

53.1

24.6

21.7

115.

819

.058

.928

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15.1

15.6

24.6

35.0

17.8

8.1

37.5

25.5

62.2

37.4

16.3

23.2

37.7

14.6

19.2

45.0

89.8

49.6

20.4

91.4

19.3

55.4

48.4

Mea

n ch

anne

l de

pth

(m)

0.52

0.35

0.40

0.73

0.84

0.33

0.16

0.20

1.06

2.90

2.34

0.74

0.81

1.27

2.62

0.73

1.29

2.09

2.00

2.21

0.61

4.60

3.54

0.85

1.18

1.65

2.19

1.34

1.20

1.71

2.32

3.98

2.02

0.13

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2

1.26

2.02

1.75

Mea

n in

stan

tane

ous

velo

city

(m

/s)

0.25

0.13

0.15

0.27

0.17

0.36

0.26

0.14

0.08

0.45

0.18

0.11

0.21

0.46

0.06

0.28

0.30

0.18

0.22

0.23

0.00

0.50

0.00

0.04

_0.2

6__

0.24

0.56

0.09

0.45

0.00

0.07

0.15

0.08

0.00

0.49

0.26

0.08

0.02

* In

clud

es a

ll ar

eas

used

for

the

prod

uctio

n of

cro

ps s

uch

as c

orn,

soy

bean

s, v

eget

able

s, t

obac

co,

and

cotto

n, a

s w

ell a

s sm

all g

rain

s su

ch a

s w

heat

and

ric

e.

Exc

lude

s ar

eas

used

for

the

prod

uctio

n of

hay

and

pas

ture

.

Table 2. Scientific and common names of fishes collected from 38 sites in the Mississippi EmbaymentStudy Unit, 1996-98, listed in phylogenetic order [Rare, occurs at 2 or less (approximately 5 percent) of the sites sampled; common, occurs at 19 or more (50 percent) of the sites sampled; 1, indicates species was rare or common; 0, indicates the species was not rare or common; (34), indicates the number of sites where the species was collected]

Scientific namePetromyzontidae - lampreys

Petromyzontidae speciesLepisosteidae - gars

Lepisosteus oculatus (Winchell, 1864)Lepisosteus osseus (Linnaeus, 1758)Lepisosteus platostomus Rafinesque, 1820

Amiidae - bowflnsAmiacalva Linnaeus, 1766

Hiodontidae - mooneyesHiodon alosoides (Rafinesque, 1819)

Anguillidae - freshwater eelsAnguilla rostrata (Lesueur, 1817)

Clupeidae - herringsDorosoma cepedianum (Lesueur, 1818)Dorosoma petenense (Gunther, 1867)

Cyprinidae - carps and minnowsCtenopharyngodon idella (Valenciennes, 1844)Cyprinella camura (Jordan and Meek, 1884)Cyprinella lutrensis (Baird and Girard, 1853)Cyprinella spiloptera (Cope, 1868)Cyprinella venusta Girard, 1856Cyprinus carpio Linnaeus, 1758Hybognathus hayi Jordan, 1885Hybognathus nuchalis Agassiz, 1855Lythrurus fumeus (Evermann, 1892)Lythrurus umbratilis (Girard, 1856)Macrhybopsis aestivalis (Girard, 1856)Macrhybopsis storer/ana (Kirtland, 1847)Notemigonus crysoleucas (Mitchill, 1814)Notropis ammophilus Suttkus and Boschung, 1990Notropis amnis Hubbs and Greene, 1951Notropis atherinoides Rafinesque, 1818Notropis buchanani Meek, 1896Notropis chalybaeus (Cope, 1869)Notropis maculatus (Hay, 1881)Notropis texanus (Girard, 1856)Notropis volucellus (Cope, 1865)Opsopoeodus emiliae Hay, 1881Phenacobius mirabilis (Girard, 1856)Pimephales notatus (Rafinesque, 1820)Pimephales promelas Rafinesque, 1820Pimephales vigilax (Baird and Girard, 1856)Semotilus atromaculatus (Mitchill, 1818)

Catostomidae - suckersCarp/odes carp/o (Rafinesque, 1820)Carp/odes cyprinus Lesueur, 1817Erimyzon obtongus (Mitchill, 1814)Hypentelium nigricans (Lesueur, 1817)Ictiobus bubalus (Rafinesque, 1818)Ictiobus cyprinellus (Valenciennes, 1 844)Ictiobus niger (Rafinesque, 1819)Minytrema melanops (Rafinesque, 1820)Moxostoma macrolepidotum (Leuseur, 1817)Moxostoma poecilurum (Jordan, 1877)

Ictaluridae - bullhead cattrshesAmeiurus me/as (Rafinesque, 1820)Ameiurus natalis (Lesueur, 1819)

Common name

lamprey

spotted garlongnose garshortnose gar

bowfin

goldeye

american eel

gizzard shadthreadfin shad

grass carpbluntface shinerred shinerspotfin shinerblacktail shinercommon carpcypress minnowmississippi silvery minnowribbon shinerredfin shinerspeckled chubsilver chubgolden shinerorangefin shinerpallid shineremerald shinerghost shinerironcolor shinertaillight shinerweed shinermimic shinerpugnose minnowsuckermouth minnowbluntnose minnowfathead minnowbullhead minnowcreek chub

river carpsuckerquillbackcreek chubsuckernorthern hogsuckersmallmouth buffalobigmouth buffaloblack buffalospotted suckershorthead redhorseblacktail redhorse

black bullheadyellow bullhead

Rare

1(1)

00

0

0

1(1)1(1)

00

000100000

1(1)0

1(2)0

1(1)1(2)

00

1(1)

1(2)1(2)

00

1(1)0

1(1)0

1(2)

00

1(2)1(1)

0000

1(2)1(1)

00

Common

0

1(35)0

1(30)

0

0

0

1(35)0

0000

1(24)

1(35)0000

00000000000

0000

1(25)0

0000

1(32)1(19)1(24)

000

00

Introduced

0

000

0

0

0

00

100001000000000000000000100

0000

000000

00

Table 2. Scientific and common names of fishes collected from 38 sites in the Mississippi Embayment Study Unit, 1996-98-Continued.

Scientific nameIctaluridae - bullhead catfishes - - continued

Ictalurus furcatus (Lesueur, 1840)Ictalurus punctatus (Rafinesque, 1818)Noturus gyrinus (Mitchill, 1817)Noturus miurus Jordan, 1877Noturus nocturnus Jordan and Gilbert, 1886Noturus phaeus Taylor, 1969Pylodictis olivaris (Rafinesque, 1818)

Esocidae - pikesEsox americanus vermiculatus Lesueur, 1846

Aphredoderidae - pirate perchesAphredoderus sayanus (Gilliams, 1824)

Cyprinodontidae - kllllfishesFundulus chrysotus (Gunther, 1866)Fundulus notatus (Rafinesque, 1820)Fundulus olivaceus (Storer, 1845)

Poeciliidae - livebearersGambusia affinis (Baird and Girard, 1853)

Atherinidae - silversidesLabidesthes sicculus (Cope, 1865)Menidia beryllina (Cope, 1866)

Percichthyidae - temperate bassesMorone chrysops (Rafinesque, 1820)Morone mississippiensis Jordan and Eigenmann, 1887Morone saxatilis (Walbaum, 1792)

Centrarchidae - sunfishesAmbloplites ariommus Viosca, 1936Elassoma zonatum Jordan, 1877Lepomis cyanellus Rafinesque, 1819Lepomis gulosus (Cuvier, 1829)Lepomis humilis (Girard, 1858)Lepomis macrochirus Rafinesque, 1819Lepomis marginatus (Holbrook, 1 855)Lepomis megalotis (Rafinesque, 1820)Lepomis microlophus (Gunther, 1859)Lepomis miniatus Evermann, 1899Lepomis hybridMicropterus punctulatus (Rafinesque, 1819)Micropterus salmoides (Lacepede, 1802)Pomoxis annularis Rafinesque, 1818Pomoxis nigromaculatus (Lesueur, 1829)

Percidae - perchesAmmocrypta beani Jordan, 1877Ammocrypta Clara Jordan and Meek, 1885Ammocrypta vivax Hay, 1882Etheostoma asprigene (Forbes, 1878)Etheostoma chlorosomum (Hay, 1 881 )Etheostoma fusiforme (Girard, 1854)Etheostoma gracile (Girard, 1859)Etheostoma histrio Jordan and Gilbert, 1887Etheostoma proeliare (Hay, 1881)Etheostoma stigmaeum (Jordan, 1877)Percina caprodes (Rafinesque, 1818)Percina maculate (Girard, 1859)Percina sciera (Swain, 1883)

Sciaenidae - drumsAplodinotus grunniens Rafinesque, 1819

Common name

blue catfishchannel catfishtadpole madtombrindled madtomfreckled madtombrown madtomflathead catfish

grass pickerel

pirate perch

golden topminnowblackstripe topminnowblackspotted topminnow

western mosquitofish

brook silversideinland silverside

white bassyellow bassstriped bass

shadow bassbanded pygmy sunfishgreen sunfishwarmouthorangespotted sunfishbluegilldollar sunfishlongear sunfishredear sunfishredspotted sunfishsunfish hybridspotted basslargemouth basswhite crappieblack crappie

naked sand darterwestern sand darterscaly sand dartermud darterbluntnose darterswamp darterslough darterharlequin dartercypress darterspecked darterlogperchblackside darterdusky darter

freshwater drum

Rare

000

1(1)1(2)1(1)

0

0

0

1(2)00

0

01(2)

000

01(2)

00000000

1(2)0000

1(1)1(1)

000

1(1)0

1(2)0

1(1)1(1)

00

0

Common

01(30)

0000

1(19)

0

0

000

1(37)

00

000

00

1(27)1(30)1(30)1(34)

01(32)

0000

1(35)1(32)1(24)

0000000000000

1(37)

Introduced

0000000

0

0

000

0

00

000

000000000000000

0000000000000

0


Table 3. Number of individuals for fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98 [96,1996; A, one of three reaches sampled; , no individuals collected]

II

SCIENTIFIC NAMEPetromyzontidae - lampreys

Petromyzontidae species Lepisosteidae - gars _~^_^,

Lepisosteus oculatus {Winchell, 1864) Lepisosteus osseus {Linnaeus, 1758) ^^^~ Lepisosteus platostomus Rafinesque, 1820 Lepisosteus species

Amiidae - bowfins Amia calva Linnaeus, 1766 flHH^

Hiodontidae - mooneyes Hiodon alosoides {Rafinesque, 1819)

Anguiiiidae - freshwater eels Anguilla rostrate (Lesueur, 1817) ^Hflffc

Clupeidae - herrings Dorosoma cepedianum {Lesueur, 1818) ~ ~ Dorosoma petenense {Gunther, 1867) Dorosoma species

Cyprinidae - carps and minnows Cteriopharyngodon idella (Valenciennes, 1844) Cyprinella camura {Jordan and Meek, 1884) Cyprinella lutrensis {Baird and Girard, 1853) Cyprinella spitoptera {Cope, 1868) Cyprinella venusta Girard, 1856 _^|HB Cyprinus carpio Linnaeus, 1758

Hybognathus hayi Jordan, 1885 Hybognathus nuchalis Agassiz, 1855 *^^BIB Lythrurus fumeus {Evermann, 1892) Lythrurus umbratilis {Girard, 1856) Macrhybopsis aesSvalis {Girard, 1856) Macrhybopsis storeriana {Kirtland, 1847) Notemigonus ciysoleucas {Mitchill, 1814) Notropis ammophilus Suttkus and Boschung, 1990 Notropis amnis Hubbs and Greene, 1951 Notropis atherinoides Rafinesque, 1818 Notropis buchanani Meek, 1896

Notropis maculatus (Hay, 1881) Notropis texanus (Girard, 1856) ! Notropis volucellus :Co^e, 1865 ^^^^^^

Opsopoeodus emiliae hay, 1881 Phenacobius mlrabllis (Girard, 1856) Pimephales notatus {Rafinesque, 1820) Pimephales promelas Rafinesque, 1820 Pimephales vigilax {Baird and Girard, 1856! St.', "jr.;, , acu/afus (Mitchill, 1818)

Catostomidae - suckers CarpKXtes carpio (Rafinesque, 1820) __ Carptodes cyprinus Lesueur, 1817

Erimyzon species Hypentelium nigncans (Lesueur, 181?) ^^^HK Ictiobus bubalus {Rafinesque, 1818) ^^^^^ Ictiobus cypnnellus (Valenciennes, 184tt^m^p. Ictiobus niger (Rafinesque, 1819) Ictiobus species Minytrema melanops (Rafinesque, 1820) Moxostoma macrolepidotum (Leuseur, 1817) Moxostoma poecilurum (Jordan, 1877)

Catostomidae species

96 A

8

-

30 32

-

-

._

-

-

-

-

-

-

6

-

-

96 B

7

8

33 17

-

-

I

-

-

-

-

-

-

5

1

96 C

15

4

5 14

-

-

-

-

-

~

-

10

4

-

4

97

14

3

1 1

1

43 4

-

17

-

-

-

-

-

10

-

2

96

6

6

38

164

5 25 5

-

7

-

-

~-

24 1 4

-

-

96 A

4 2 3

1

8 14

9

-

_

-

-

6

-

-

5

-

-

96 B

6

2

-

26 11

-

-

-

-

8

-

-

21

-

-

96 C

7

5

3

1

2 12 2 2

-

1

-

-

7

-

-

2

-

97

4

4

3

41 15

10

1

178

1

1

16

_

-

7 1

-

-

98

3 1 1

3

17

21 17 1

72

5

I

-

-

5

1

-

14 3 6

-

-

96

42 7

77 36

30

-

_

-

-

-

-

-

62 5 1

-

97

3 22

35

21 14

-

3

-

-

-

_

-

48 1 4

-

-

98

3

5

4

2 13

-

-

-

-

6

-

-

36 10 22

-

-

96

2

10

193 9

3 1

38

21

1

751

g

-

6

2

I

-

97

4

1

70 5

1

26

17

53

2 3 1

5

-

98

3

3

122 6

5

15

1

18

29

13 _

1 5

2

-

Table 3. Number of individuals for fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98 Continued

SCIENTIFIC NAMEIctaluridae - bullhead catflshes

Ameiurus me/as (Rafinesque, 1820)

Ictalurus furcatus (Lesueur, 1840) Ictalurus punctatus (Rafinesque, IB 18) ^^HBV Noturus gyrinus (Mitchill, 1817)

Noturus phaeus Taylor, 1969 Pylodictis olivaris (Rafinesque, 1818)

Esocldae - pikes Esox americanus vermiculatus Lesueur, 1846

Aphredoderidae - pirate perches Aphredoderus sayanus (Gilliams, 1824)

Cyprinodontidae - kiliif ishes jgffE*.

Fundulus notatus (Rafinesque, 1820) jjffgf Fundulus olivaceus (Storer, 1845)

Poeciiiidae - iivebearersGambusia affinis (Baird and Girard, 1853) ^ff^^

Atherinldae - silversides Labidesthes sicculus (Cope, 1865) ^^^Bi

Menidia beryllina (Cope, 1B66) Percichthyidae - temperate basses

Morons chrysops (Rafinesque, 1820)

Morons saxatilis (Walbaum, 1792) Centrarchldae - sunfishes

Amb/oplites ariommus Viosca, 1936

Lepomis cyanellus Rafinesque, 1819 Lepomisgulosus (Cuvier, 1829) Lepomis humilis (Girard, 1858) Lepomis macrochirus Rafinesque, 1819 Lepomis marginatus (Holbrook, 1855) Lepomis megalotls (Rafinesque, 1820) Lepomis microlophus (Gunther, 1859) Lepomis mlniatus Evermann, 1899

Lepomis species Micropterus punctulatus (Rafinesque, 1819) Mlcroptarus salmoides iLaca^ede, 1802) Micropterus species Pomoxls annularis Rafinesque, 1818 ^g^^^p Pomoxis nigromaculatus (Lesueur, 1829)

Percidae - perches

Ammocrypta clara Jordan and Meek, 1885

Etheostoma asprigene (Forties, 1878) Etheostoma chlorosomum (Hay, 1881)

Etheostoma gracile (Girard, 1859)

Etheostoma proeliare (Hay, 1881) ^^^^ Etheostoma stigmaeum (Jordan, 1877)

Percina maculata (Girard, 1859)

Sciaenidae - drumsAplodinotus gwnniens Rafinesque, 1819

Unclassified fishes Unknown fry

96 A

1

8

2

-

1

~

:1 1

34

3

3

»

-

-

-

7

96 B

-

4

1

-

-

::4 12 3

1

-

-

-

-

-

-

4

<D96 C

--

1

3

-

3

_

1 4 25 9

1

7

-

-

-

-

-

19

97

-

4

1

-

31

~

1 102 19

1

1

6

5

-

-

-

-

-

17

98

-

1

-

-

53

~

:5 2 19 13

1

526

1 1

37 4

-

-

-

1

-

7

196 A

-

2 2

1

3

1

5

:7 7

2

1

3 2

8 4

-

2

-

-

-

10

96 B

-

2

4

1

1

:2 3 9

1

1

9

-

-

-

-

-

8

o96 C

-

67

2

-

-

__

4 3 1 4 1 4

8 2

161

-

2

-

-

8

97

-

3 2

2

1

5

34

~~

::3 2

4

2

2

11

-

41

-

~

1

tt

2

98

-

9 2

4

11

7

56

::10 10 15

26

3

5

27

-

1

3

-

-

26

1

I96

-

3

8

-

~

9 1

7

1 2

9 1

-

-

-

-

-

11

0"

97

-

2

1

-

35

~~

'_

5 17 9

2

2

15 2

-

6

-

-

5

12

98

-

7

5

-

136

_

:2

14

2

108

6

-

8

-

-

4

4

96

-

8

2

1

4 2

2

_

13

14 5 3 16

35

12

1 9 67

1

1 1

-

4

1

10

w97

-

6

1

3

3

~

1

10

11 3 1 5

35

6

1 8 5

1

-

-

-

-

-

5

98

-

6

3

1

m? --T

12

6 -^m.

-

>

23 2

19

64

2

5 16 10

1

-

-

-

-

1

11


Table 3. Number of individuals for fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98-Continued

SCIENTIFIC NAME'" Petromyzontidae - lampreys

Petromyzontidae species Leplsosteidae - gars

Lepisosteus oculatus (Winchell, 1864) Lepisosteus osseus (Linnaeus, 1758) Lepisosteus platostomus Rafinesque, 1820

Amildae - bowfins Amutcalva Linnaeus, 1766


Anguillidae - freshwater eelsAnguilla rostrate (Lesueur, 1817)

Clupeidae - herrings Dorosoma cepedianum (Lesueur, 1818) Dorosoma petenense (Gunther, 1867) ^ ""soma species

Cyprinidae - carps and minnows

Cyprinella camura (Jordan and Meek, 1884) Cyprinella lutransis (Baird and Girard, 1853)

Cyprinella veriusta Girard, 1856 Cyprinus carpio Linnaeus, 1 758 Cyprinidae speciesHybognathus hayi Jordan, 1885

Lythrurus fumeus (Evermann, 1892) Lythrurus umbratilis (Girard, 1856)

Macrhybopsis storariana (Kirtland, 1847) Notemigonus crysoleucas (Mitchiil, 1814)

Notropis amnis Hubbs and Greene, 1951 Notropis atherinoides Rafinesque, 1818 Notropis buchanani Meek, 1896Notropis chalybaeus (Cope, 1869)Notropis maculatus (Hay, 1881)Notropis texanus (Girard, 1856) Notropis volucellus (Cope, 1865)

Opsopoeodus emiliae Hay, 1881Phenacobius mirabilis (Girard, 1856) Pimephales notatus (Rafinesque, 1820) Pimephales promelas Rafinesque, 1820 Pimephales vigilax (Baird and Girard, 1856)Semotilus atromaculatus (Mitchiil, 1818)

Catostomidae - suckersCarp/odes carpio (Rafinesque, 1820)

Erimyzon oblongus (Mitchiil, 1814)

Hypentelium nigricans (Lesueur, 1817) ictiobus bubalus (Rafinesque. 1818) Ictiobus cyprinellus (Valenciennes, 1844) Ictiobus niger (Rafinesque, 1819) Ictiobus species Minytrema melanops (Rafinesque, 1820)

Moxostoma poecilurum (Jordan, 1877)

96

2 1

-

8

-

251 1

-

-

10

-

-

28

-

1 1

-

c?97

9 1 1

-

2

6

1007

-

1

23 1

-

-

18

1

1 1

-

98

7 1 1

-

11

9

541 5 1

3

1

9

-

1

1

26 1

1

22

-

96

7

8

-

14

1

5 2

-

-

1

-

-

-

-

15 1

5

-

1

197

8

3

-

2

-

8 21

-

-

18

-

-

-

-

30

1

-

98

4

2

-

5

2

2

-

-

18

-

-

7

-

29

3

--

/96

3

1

15

57

1

223 3

-

1 11

-

1

2

r

2

4

197

7

-

3

60

1

3

-

3

-

-

1

-

1

1

5

98

1

-

6

23

91

-

1

-

-

:

-

4 1

1

6

3

96

2 1 14

31

5584

-

_

-

1

12

-

-

-

-

2

-

I97

1 2 2

-

19 60

-

4

-

-

14 3

1

1

1

-

2

-

98

2 2

-

171 836

2

3

-

22

161 1

5

1

-

2

-

/I98

15 1 1

-

12

12

-

27 2

16

-

1

13

-

3

-

29 9 3

-

/I / 03

98

1

-

8

25

2 3

-

-

-

--

-

30

-

-

-

/I / °>

98

3

4

-

23 15 15

1

302

18 11

-

-

1 1

--

-

29

-

41 1 8

-

/I / °>

98

11 4 3

-

28

195

2

18

9

-

22

-

-

2

2

-

29 6

2

-

/I98

7

2

1

21

-

16 11

2

-

2

-

-

15

1

-

27

4

-

11


SCIENTIFIC NAMEIctalurldae - bullhead catfishes

Ameiurus me/as (Rafinesque, 1820) Amelurus natalls (Lesueur. 1819) Ictalurus furcatus (Lesueur. 1840) Ictalurus punctatus (Rafinesque. 1818) Noturus gyrinus (Mitchill, 1817) Noturus miurus Jordan, 1877 Noturus nocturnus Jordan and Gilbert. 1886 Noturus phaeus Taylor, 1969 Pylodictis olivaris (Rafinesque, 1818)

Esocidae - pikes Esox americanus vermiculatus Lesueur. 1 846 j|} Aphredoderidae - pirate perches Aphredoderus sayanus (Gilliams, 1824)

Cyprinodontldae - kllllflshes Fundulus chrysotus (Gunther, 1866) Fundulus notatus (Rafinesque, 1820) Fundulus olivaceus (Storer, 1845) Fundulus species

Poeclllldae - livebearers Gambusia affinis (Baird and Girard, 1853 1

Atherinidae - sllversldes Labidesthes slcculus (Cope, 18651 Menidia beryllina (Cope, 1866)

Percichthyidae - temperate basses Morone chrysops (Ra*inesque, 1820) Morone mississippiensis Jordan and Morone saxatilis (Walbaum, 1792)

Centrarchldae - sunfishes Ambloplites ariommus Viosca, 1936 Elassoma zonatum Jordan, 1877 Lepomis cyanellus Rafinesque, 1819 Lepomis gulosus (Cuvier, 1829) Lepomis humilis (Girard, 1858) Lepomis macrochirus Rafinesque, 1819 Lepomis marginatus (Holbrook, 1855) Lepomis megalobs (Rafinesque, 1820) Lepomis microlophus (Gunther, 1859) Lepomis mmiatus Evermann, 1899 Lepomis hybrid Lepomis species Micropterus punctulatus (Rafinesque, 1819) Micropterus salmoides (Lacepede 1802) Micropterus species Pomoxis annularis Rafinesque, 1818 Pomoxis nigromaculatus (Lesueur, 1829) Pomoxis species

Percldae - perches Ammocrypta beam Jordan, 1877 Ammocrypta tiara Jordan and Meek, 1885 Ammocrypta vivax Hay. 1882 Etheostoma asprigene (Fortes, 1878) Etheostoma chlorosomum (Hay, 1881) Etheostoma fusiforme (Girard, 1854) Etheostoma gracile (Girard, 1859) Etheostoma histrio Jordan and Gilbert, 1887 Etheostoma proeliare (Hay, 1881) Etheostoma stigmaeum (Jordan, 1877) Percina caprodes (Rafinesque, 1818) Percina maculata (Girard, 1859) Percina sctera (Swain, 1883)

Sciaenidae - drums Apkxtnolusgrunniens Rafinesque 1H1Q

Unclassified fishes

/ £96

-

13

5

1

3

1 1

-

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-

7

97

5

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46

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3 1 2 3

4 1

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-

8

98

4 5

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8

97

6

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10

2 18

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22

-

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12

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98

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78

2 1 6

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59

8 1

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14

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15

1

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8 4 8 3 4

1 11 1

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2

2

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97

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21

2

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22 2 4

7

10 2

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7

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8

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1 1

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30 2

20 4 3

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87

-

7

82

8

5

9 9

13

59 4

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15

97

10 9

1

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42

14

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14

1

z14

-

-

12

98

8 41

2

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123

3 1

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6 21 9

277

5

S

8

1175

-

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-

81

/of98

8

1

6

41

34 18

128

147

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68 4

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2

-

25

$98

2

-

81

9

5 3 10 3

29

81

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9

-

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s>

I98

1

5

-

73

8 21

17

16

1

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-

-

18

£98

1

1

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36

2

1 3 16 25

10

3

9

377

6

2

1

10

»&98

-

1

-1 2

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-1

1

- 1

1 9 I 1

16

14 1 -1

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81

13f

t

-1

7*

31

- 1

1-t

8*


Table 3. Number of individuals for fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98~Continued


Lepisosteidae - garsLepisosteus oculatus (Winchell, 1864) Lepisosteus osseus (Linnaeus, 1758) Lepisosteus platostomus Rafinesque, 1820

Amiidae - bowfins /Am/a calva Linnaeus, 1766

Hiodontidae - mooneyes

Anguiiiidae - freshwater eeis

Ciupeidae - herringsDorosoma cepedianum (Lesueur, 1818) Dorosoma petenense (Gunther, 1867)


Cyprinella lutrensis (Baird and Girard, 1853)

Cyprinella venusta Girard, 1856 Cyprinus carpio Linnaeus, 1758 Cyprinidae species Hybognathus hayi Jordan, 1885

Lythrurusfumeus (Evermann, 1892)

Macrhybopsis aestivalis (Girard, 1856)

Notemigonus crysoleucas (Mitchill, 1814)

Notropis atherinoides Rafinesque, 1818 Notropis buchanani Meek, 1896 Notropis chalybaeus (Cope, 1869)

Notropis volucellus (Cope, 1865)

Opsopoeodus emiliae Hay, 1881

Pimephales notatus (Rafinesque, 1820)

Pimephales vigilax (Baird and Girard, 1856)Semotilus atromaculatus (Mitchill, 1818)

Catostomidae - suckers Carp/odes carpio (Rafinesque, 1820) Carpiodes cyprinus Lesueur, 1817

Ictiobus bubalus (Rafinesque, 1818) Ictiobus cyprinellus (Valenciennes, 1844) Ictiobus niger (Rafinesque, 1819) Ictiobus species Minytrema melanops (Rafinesque, 1820)

1 S1

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92

18 5

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/C?98

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196

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118

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177

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2061

10

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13

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11

12

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598

1

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1

10

-

6

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-

-

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49

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/ ;J98

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9

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4 1

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79 4

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43

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27

-

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41

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73

14 10

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127

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13

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7

18

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°98

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1 12

s98

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92 10 6

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13


SCIENTIFIC NAMEIctalurldae - bullhead catfishes

Ameiurus me/as (Rafinesque, 1820) Ameiurus natatis (Lesueur, 1819) Ictalurus furcatus (Lesueur, 1840) ICtalurus punctatus (Rafinesque, 1818) Noturus gyrinus (Mitchill, 1817)

Noturus nocturnus Jordan and Gilbert, 1886Noturus phaeus Taylor, 1969 Pylodictis olivaris (Rafinesque, 1818)

Esocldae - pikes Esox americanus vermiculatus Lesueur, 1846


Cyprinodontidae - killifishM Funuulus chrysotus (Gunther, 1866) Fundulus notatus (Rafinesque, 1820) Fundulus olivaceus (Storer, 1845)

Poeciliidae - llvebearersGambus/a affinis (Baird and Girard, 1853)

Atherinldae - silversides Labidesthes sicculus (Cope, 1865)

Percichthyidae - temperate bassesMorone chrysops (Rafinesque, 1820)

Morone saxatilis (Walbaum, 1792) Centrarchidae - sunfishes

Ambloplites ariommus Viosca, 1936

Lepomis cyanellus Rafinesque, 1819 Lepomis gulosus (Cuvier, 1829) Lepomis humilis (Girard, 1858) Lepomis macrochirus Rafinesque, 1819

Lepomis megalotis (Rafinesque, 1820) Lepomis microlophus (Gunther, 1859) Lepomis miniatus Evermann, 1899 Lepomis hybrid Lepomis species Micropterus punctulatus (Rafinesque, 1819) Micropterus salmoides (Lacepede, 1802)

Pomoxis annularis Rafinesque, 1818 Pomoxis nigromaculatus (Lesueur, 1829)

Percldae - perches

Ammocrypta Clara Jordan and Meek, 1 885 Ammocrypta vivax Hay, 1882 Etheostoma asprigene (Forbes, 1878) Etheostoma chlorosomum (Hay, 1881)


Etheostoma proeliare (Hay, 1881)

Percina maculata (Girard, 1859)

Sclaenidae - drums /(psxftwlu* grunniens Rafinesque, 1819

Unclassified fishes

98

2

-

1

217

1

-

~~

2 3 6

12

4 9 15

161

-

-

-

2

1

98

2 1

-

2

124

-

1

1 4 1

59

5

51

-

-

-

15

98

-

-

2

11

-

__

6 6 16 16

2

66

1

111

-

-

-

13

98

1

-

-

215

1

-

__

6 3 5 10

13

1

8

-

-

-

14

98

1 1 4 1

-

-

S71

-

16 t

6

1

7 5

-

-

-

3

98

1 6

-

5

3

T

-

4

10 2

21

72 3

134 32 6 7 13

1

-

1

-

9

98

1 2

-

1

22

_

2 1

15

1

13 1

-

-

-

9

98

1

1 1

-

1

86

-

~

4 14 23

12

28

5

14

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-

1

98

1

102

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1

137

-

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24 1

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53

9 3

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6

98

5

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6 22 3

85

37 2 3

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98

8

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11

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1 7

22

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12 1

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98

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6

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237

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56 10

12

84 1

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20 10 4

2 2

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9 2

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57

13 25

1

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10 8 3 35 6

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8

30

8

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16

21 2

52

-

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52

98

2 31

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24

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1

1

14

1

13

-

-

-

4

98

1 1

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3

1

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39

1

6 5

4 1

8

2

581

10

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12

98

;;<-

7

2

1

27

52"

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:>57 29

1 108

158 10 26 2 21

12 11

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1f

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1



SCIENTIFIC NAMEPetromyzontidae - Lamprey*


Lepisosteus oculatus (Winchell, 1864)Lepisosteus osseus (Linnaeus, 1758)Lepisosteus platostomus Rafinesque, 1820Lepisosteus species

Amiidae - bowfinsAm&calva Linnaeus, 1766


98

6

3

98

-

-

2

1

Hiodon alosoides (Rafinesque, 1819) ^ miBI -Anguillidae - freshwater eels

Anguilla rostrata (Lesueur, 1817)Clupeidae - herrings

Dorosoma cepedianum (Lesueur, 1818)Dorosoma petenense (Gunther, 1867)Dorosoma species

Cyprinidae - carps and minnowsCtenopharyngodon idella (Valenciennes, 1844)Cyprinella camura (Jordan and Meek, 1884)Cyprinella lutrensis (Baird and Girard, 1853)Cyprinella spiloptera (Cope, 1868)Cyprinella venuste Girard, 1856 -: ,;Cyprinus carpio Linnaeus, 1758Cyprinidae species ~^^^IKHybognathus hayi Jordan, 1885Hybognathus nuchalls Agassiz, 1855Lythrurus fumeus (Evermann, 1892)Lythrurus umbrahlis (Girard, 1856)Macrhybopsis aestivalis (Girard, 1856)Macrhybopsis storeriana (Kirtland, 1847)Notemigonus crysoleucas (Mitchill, 1814)Notropis ammophilus Suttkus and Boschung, 1990Notropis amnis Hubbs and Greene, 1951Notropis atherinoides Rafinesque, 1818Notropis buchanani Meek, 1896Notropis chalybaeus (Cope, 1869 iNotropis maculatus (Hay, 1881)Notropis texanus (Girard, 1856)Notropis volucellus (Cope, 1865)Notropis speciesOpsopoeodus em///ae Hay, 1881Phenacobius mirabilis (Girard, 1856)Pimephales notatus (Rafinesque, 1820)Pimephales promelas Rafinesque, 1820Pimephales vigilax (Baird and Girard, 1856)S»'V.T,,'!,"= .,tromaculatus (Mitchill, 1818)

Catostomidae - suckersCarp/odes carp/o (Rafinesque, 1820)Carp/odes cyphnus Lesueur, 1817Erimyzon oblongus (Mitchill, 1814)Erimyzon speciesHypentelium nigricans (Lesueur, 1817)Ictiobus bubalus (Rafinesque, 1818)Ictiobus cypnnellus (Valenciennes 1844Ictiobus niger (Rafinesque. 1819)Ictiobus speciesMinytrema melanops (Rafinesque, 1820)Moxostoma macrolep'dotum (Leuseur, 1817)Moxostoma poecilurum (Jordan, 1877)Moxostoma speciesCatostomidae species

19

-- -

8 - - - 91 - -

3 -------

--

--

916-1-----

50-

-1

5221

5 --

12136 -- 16 --

1-- -

39 - 2

19

--

-

12 8 - ---

98

-

7

4

_

25

--

654 - - - - -

r- - 3 -

1

2-

--

3 - .---

98

-

I12

-.

20

--

8186 14 t

1 5

93

- - -

1112

1 - --

98

1 ~

4

5-

-

1920 20 - -_ ~

-

-

4 13 --

98

-

14111

_

1

73

-

4 - - - -- - 30

3 - -

14619 --

98

4

_

98

14

1

1

SUBBB

179643

--

1999

4 - -

93 -- - 1 -

1

--- -

1 --

^^g

-- -

- -_- -- - 2 - - -- -

--- --- -- 2 --

15

Table 3. Number of individuals for fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98~Continued

SCIENTIFIC NAMEIctaluridae - buiihead catfishM .

Ameiurus me/as (Rafinesque, 1820)Ameiurus natalis (Lesueur, 1819) ^jiSifIctalurus furcatus (Lesueur, 1840)Ictalurus punctatus {Rafinesque. 1818)Noturus gyrinus (Mitchill, 1817)Noturus miurus Jordan, 1877Noturus nocturnus Jordan and Gilbert, 1886Noturus phaeus Taylor, 1969Pylodictis olivaris (Rafinesque, 1818)

Esocidae - pikesEsox americanus vermiculatus Lesueur, 1846T, Aphredoderidae - pirate perchesAphreooderus sayanus (Gilliams, 1824)<T Cyprinodontidae - kiliifishesFunoulus chrysotus (Gunther, 1866)Fundulus notatus {Rafinesque, 1820)Fundulus divaceus (Storer, 1845) ^^^^^Fundulus species ifffff

Poeciiiidae - livebearers Gambusia affims (Baird and Girard, 1853)

Atherinidae - siiversidesLabidesthes sicculus (Cope, 1865) flHHfrMenidia beryllina (Cope, 1866)

Percichthyidae - temperate bassesMorone chrysops (Rafinesque, 1820)Morone mississippiensis Jordan and Eigenmann,Morone saxatilis (Walbaum, 1792)

Centrarcnidae - sunfishesAmbloplites ariommus Viosca, 1936Elassoma zonatum Jordan, 1877 ^^f^^fLepomis cyanelus Rafinesque, 1819Lepomis gulosus (Cuvier, 1829) ^^f^fLepomis humiis (Girard, 1858)Lepomis macrochirus Rafinesque, 1819Lepomis marginatus (Holbrook, 1855)Lepomis megalotis (Rafinesque, 1820)Lepomis microlophus (Gunther, 1859)Lepomis miniatus Evermann, 1899Lepomis hybridLepomis speciesMicropterus punctulatus (Rafinesque, 1819)Micropterus salmoides (Lacepede, 1802) ____Micropterus speciesPomoxis annularis Rafinesque, 1818^^HBPomoxis nigromaculatus (Lesueur, 1829)Pomoxis species

Percidae - perchesAmmocrypta beam Jordan, 1877Ammocrypta Clara Jordan and Meek, 1885Ammocrypta vivax Hay, 1882Etheostoma asprigene (Forbes, 1878)Etheostoma chlorosomum (Hay, 1881)Etheostoma fusiforme (Girard. 1854)Etheostoma gracile (Girard, 1859)Etheostoma histrio Jordan and Gilbert, 1887Etheostoma proeliare {Hay, 1881)Etheostoma stigmaeum (Jordan, 1877)Percina caprodes (Rafinesque, 1818)Percina maculata (Girard, 1859)Percina sciera (Swain, 1883) flBHB

Sciaenidae - drumsAplodinotus gr unniens Rafinesque, 1819

Unclassified fishesUnknown fry

/ co98

-

1 -

6

- -

---f434 1 - 26-1812

- - -

2

-

/ CO

98

-14- - 1

2

- 3

62

6-

36

125 65 1 1317 13

-

- 1

13

-

/ w98

42

I

I

5

- 1- 11 1-

- - -

11

-

/ CO

98

-

1

3

- 21

22

-

1 -

1 12141

56

101-- 12355 31

41

68 1 2- 21

5

-

/ CO

98

-I-2-

3

2

-13

139

-

I116-23 32 6 49 5

1-- 2

4

-

/ CO

98

-2-

584

41

- 17 4824 14

26017 306

- - -----

20

-

/ (^98

- -3--- 4

-

---

94

-

- -

2 2

- 8---2-

~-- - ---

5

-

^98

I

4

3

---

--

- 1173

1362263112---12- 2

--

-3--fa ^H --

-

-


Table 4. Percent relative abundance of fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98 [96, 1996; A, one of three reaches sampled; -, no individuals collected; 0, value less than .005]


Leplsosteldae - garsLepisosteus oculatus (Winchell, 1864) Lepisosteus osseus (Linnaeus, 1758) Lepisosteus platostomus Rafinesque, 1820 Lepisosteus species

Amiidae - bowfins Amia calva Linnaeus, 1766



Clupeidae - herrings Dorosoma cepedianum (Lesueur, 1818) Dorosoma petenense (Gunther. 1867) Doroxmia species


Cyprinella lutrensis (Baird and Girard, 1853) Cyprinella spiloptera (Cope, 1868) Cyprinella venusta Girard, 1856 Cyprinus carpio Linnaeus, 1758 Cyprinidae species Hybognathus hayi Jordan, 1885 Hybognathus nuchahs Agassiz, 1855

Macrhybopsis storeriana (Kirtland, 1847) Notemigonus crysoleucas (Mitchill, 1814)Notropls ammopbilus Suttkus and Boschung, 1990 Notropis amnis Hubbs and Greene, 1951 Notropls atherinoides Rafinesque, 1818 Notropis buchanani Meek. 1896 Notropis chalybaeus (Cope, 1869} Notropis maculatus (Hay, 1881)

Notropis volucellus (Cope, 1865)

Opsopoeodus emillae Hay, 1881 Phenacobius mlrabilis (Girard, 1856)

Pimephales promelas Rafinesque, 1820 Plmephales vigilax (Baird and Girard, 1856) Semotilus atromaculatus (Mitchill, 1818)

Catostomldae - suckers Carp/odes carpio (Rafinesque, 1820)

Erimyzon oblongus (Mitchill, 1814)

Ictiobus bubalus (Rafinesque. 1818) Ictiobus cyprinellus (Valenciennes, 1844) Ictiobus niger (Rafinesque, 1819)

/96 A

5.84

-

21.90 24.62

-

-

-

-

-

4.38

96 B

6.93

7.92

32.67 16.83

-

0.99

-

-

-

4.95

nQQ

#96 C

12.00

3.20

4.00 11.20

-

-

-

-

-

8.00

3.20

*9n

97

4.93

1.06

0.35 0.35

0.35

15.14 1.41

-

5.99

-

-

-

3.52

n7n

98

0.63

0.63

4.00

17.25

0.53 2.63 0.53

-

0.11

0.11

-

-

2.52 0.11 0.42

/96 A

3.57 1.79 2.68

089

7.14 12.50

8.04

-

-

-

-

5.36

4.46

96 B

5.22

1.74

E

22.61 9.57

-

-

-

-

6.96

18.26

O96 C

5.98

4.27

2.56

0.85

1.71 10.26 1.71 1.71

-

0.85

-

-

5.98

3.42

1.71

97

1.27

1.27

0.95

12.97 4.75

3.16

0.32

0.32 24.68

0.32

0.32

5.06

2.22 0.32

98

0.78 0.26 0.26

0.78

4.39

5.43 4.39 0.26 18.60

1.29

0.26

-

-

1.29

0.26

3.62 0.78 1.55

/96

1.44 0.72 2.53

27.80 13.00

10.83

-

-

-

-

-

22.38 1.81 0.36

o"97

1.27 0.84 0.84

14.77

8.86 5.91

-

1.27

-

-

-

20.25 0.42 1.69

98

0.79

1.32

1.06

0.53 3.43

-

-

-

-

1.58

9.50 2.64 0.53 n CQ

/96

0.38

1.88

36.35 1.69

0.56

7.16

3.95

0.19

14.12 0.19

1.69

1.13

0.38

0.19

o97

1.37

0.34

23.89 1.71

0.34

8.87

5.80

-

18.09

0.68 1.02

1.71

98

0.70

0.70

-

28.50 1.40

-

1 17

3.50

0.23

4.21

6.78

3.04

0.23 1.17

0.47

17

Table 4. Percent relative abundance of fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98-Continued

SCIENTIFIC NAMEIctaluridae - bullhead catfishes

Ameiurus natalis (Lesueur, 1819) Ictalurus furcat us (Lesueur, 1840) Ictalurus punctatus (Rafinesque, 1818) Noturus gyrinus (Mitchill, 1817)

Noturus noctumus Jordan and Gilbert. 1886Noturus phaeus Taylor, 1969 Pylodictis olivaris (Rafinesque, 1818)

Esocidae - pikes


Cyprmodontidae - kiliifishes

Fundulus notatus (Rafinesque, 1820) Fundulus olivaceus (Storer, 1845)

Poeciiiidae - iivebearersGambusia affinis (Baird and Girard, 1853)

Atherinidae - siiversides Labidesthes sicculus (Cope, 1865)

Percichthyidae - temperate basses

Centrarchidae - sunfishes

E/assoma zonatum Jordan, 1877 Lepomis cyanellus Rafinesque, 1819 Lepomis gulosus (Cuvier, 1829) Lepomis humilis (Girard, 1858) Lepomis macrochirus Rafinesque, 1819

Lepomis megalotis (Rafinesque, 1820)

Lepomis miniatus Evermann, 1899

Lepomis species Micropterus punctulatus (Rafinesque, 1819) Micropterus salmoides (Lacepede, 1802) Micropterus species Pomoxis annularis Rafinesque, 1818 Pomoxis nigromaculatus (Lesueur, 1829) Pomoxis species

Percidae - perches

Ammocrypta Clara Jordan and Meek, 1885

Etheostoma asprigene (Forbes, 1878) Etheostoma chtorosomum (Hay, 1881)


Etheostoma proeliare (Hay, 1881) Etheostoma stigmaeum (Jordan, 1877)

Percina sciera (Swain, 1883) Sciaenidae - drums

Aplodinotus grunniens Rafinesque, 1819 Unclassified fishes

Unknown fry

96 A

0.73

5.84

-

1.46

-

-

0.73

0.73 0.73 24.82

2.19

-

2.19

-

-

-

-

511

96 B

3.96

-

0.99

-

-

-

3.96 11.88 2.97

-

0.99

-

-

-

-

-

3.96

I

196 C

0.80

-

2.40

-

-

2.40

0.80 3.20 20.00 720

-

-

080

5.60

-

-

-

-

12.00

'

97

1.41

-

0.35

-

-

10.92

0.35 35.92 6.69

0.35

-

0.35

2.11

1.76

-

-

-

-

5.99

98

011

-

-

-

-

5.57

0.53 0.21 2.00 1.37

011

-

55.31

011 0.11 3.89 0.42

-

-

-

0.11

0.74

96 A

1.79 1.79

-

0.89

2.68

0.89

4.46

6.25 6.25

1.79

0.89

2.68 1.79

7.14 3.57

-

1.79

-

-

8.93

96 B

1.74

-

3.48

-

087

087

1.74 2.61 7.83

-

-

0.87

0.87

7.83

-

-

-

-

6S6

k$

C?96 C

5.13 5.98

-

-

1.71

-

-

3.42 2.56

3.42

3.42

6.84 1.71

13.68 0.85

-

1.71

-

-

6.84

-

97

0.95 0.63

-

0.63

0.32

1.58

10.76

0.95 0.63

1.27

0.63

0.63

3.48

-

12.97

-

-

032

5.70

0.63

98

2.33 0.52

-

1.03

2.84

1.81

14.47

2.58 2.58 3.88

6.72

0.78

1.29

6.96

-

0.26

0.78

672

0.26

/96

1.08

-

2.89

-

-

0.36

3.25 036

2.53

-

0.36 0.72

3.25 0.36

-

-

-

-

3.97

J

I97

0.84

-

0.42

-

-

14.77

2.11 0.42 2.95 3.80

0.84

-

0.84

6.33 0.84

-

2.53

-

-

2.11

5.06

ID

98

1.85

-

1.32

-

-

35.88

0.53

3.69

0.53

-

28.50

1.58

-

2.11

-

-

A f)f>

1.06

96

1.51 0.19

0.38

0.19

-

0.75 0.38

0.38

2.45

2.64 094 0.56 3.01

6.59

2.26

0.19 1.69 1.13 1.32

0.19

0.19 n *IQ

-0.75

0.19

1.88

1 if

.1 -j97

2.05

-

0.34

-

1.02

t.02

0.34

3.41

3.75 1.02 0.34 1.71

11.95

2.05

0.34 2.73 1.71

0.34

-

-

-

-

1 71

98

1.40

-

0.70

0.23

3.04 0.93

2.80

1.40

5.37 0.47

4.44

14.95

0.47

1.17 3.74 2.34

0.23

-

-

-

-

0.23

2.57




Petromyzontidae species Lepisosteidae - gars

Lepisosteus oculatus (Wmchell. 1864) Leplsosteus osseus (Linnaeus, 1758) Lepisosteus platostomus Rafinesque, 1820

Amiidae - bowf ins Amia calva Linnaeus, 1766

Hiodontidae - mooneyes Hiodon alosoides (Rafinesque, 1819)

Anguiiiidae - freshwater eeis Anguilla rostrate (Lesueur, 1817)

Ciupeidae - herringsDorosoma cepedianum (Lesueur, 1818) Dorosoma petenense (Gunther, 1867) Dorosoma species


Cyprinella camura (Jordan and Meek, 1884) Cypnnella lutrensis (Baird and Girard, 1853)

Cyprinella venusta Girard, 1858 Cyprinus carpio Linnaeus, 1758 Cyprinidae species

Lythrurus fumeus (Evermann, 1892) Lythrurus umbratilis (Girard, 1858)

Macrhybopsis storeriana (Kirtland, 1847) Notemigonus crysoleucas (Mitchill, 1814)

Notmpis athennoides Rafinesque, 1818 Notropis buchanam Meek. 1896

Votrop/s volucellus (Cope, 1865)

Opsopoeodus emiliae Hay, 1881

Pimephales vigilax (Baird and Girard, 1856)

Catostomidae - suckers

Carpiodes cypnnus Lesueur, 1817

Hypentelium nigricans (Lesueur, 1817) Wiobus bubalus (Rafinesque, 1818) 'ctiobus cyprinellus (Valenciennes. 1844) 'ctiobus niger (Rafinesque, 1819)

Minytrema metenops (Rafinesque, 1820)

Woxostoma poecilurum (Jordan, 1877)

96

0.60 0.30

-

2.40

-

75.15 0.30

-

-

2.99

-

-

8.38

-

0.30 0.30

-

-

J

197

3.61 0.40 0.40

-

0.80

2.41

40.16 2.81

-

0.40

9.24 0.40

-

-

7.23

0.40

0.40 0.40

-

-

i

98

1.00 0.14 0.14

-

1.56

1.28

76.96 0.71 0.14

0.43

0.14

1.28

-

0.14

0.14

3.70 0.14

-

0.14

3.13

--

--

96

7.14

8.16

-

14.29

1.02

5.10 2.04

-

-

1.02

-

-

-

-

15.31 1.02

5.10

-

1 n?

1 <t #

197

4.57

1.71

-

1.14

-

4.57 12.00

-

-

10.29

-

-

-

-

17.14

0.57

-

-

i>

98

1.60

0.80

-

2.00

0.80

0.80

-

-

7.20

-

-

2.80

-

11.60

1.20

-

-

96

0.76

0.25

3.82

14.50

0.25

56.74 0.76

-

0.252.80

-

0.25

0.51

0.25

051

-

1.02

ID

1

197

3.68

-

1.58

31.58

0.53

1.58

-

1.58

-

-

0.53

-

0.53

0.53

2.63

98

0.44

-

2.62

10.04

;;39.74

-

0.44

-

-

-

-

1.75 0.44

0.44

2.62

1.31

96

0.47 0.23 3.27

724

12.85 19.63

-

-

0.23

2.80

-

-

-

-

0.47

-

-

£s

I97

0.39 0.78 0.78

-

7.39 23.35

-

1.56

-

-

545 1.17

0.39

0.39

0.39

-

0.78

-

-

98

0.10 0.10

-

8.96 43.79

0.10

0.16

_

1.15

8.43 0.05

0.26

-

0.05

-

0.10

-

-

/J //

98

2.28 0.15 0.15

-

1.82

1.82

_

4.10 0.30

2.43

-

0.15

1.98

-

0.46

-

4.41 1.37 0.46

-

-

- / *£

198

0.32

2.57

8.04

0.64 0.96

-

-

-

-

-

9.65

-

-

-

-

7J I98

0.48

0.64

-

3.67 2.40 2.40

0.16

4824

2.88 1.76

-

-

0.16 0.16

--

--

4.63

1 OR

6.55 0.16 1.28

-

-

7J //

98

2.18 0.79 0.60

-

556

38.69

0.40

3.57

1.79

-

4.37

-

-

0.40

-

D.40

5.75 1.19

0.40

-

> / 9

It/<?

98

2.60

0.74

0.37

7.81

-

595 4.09

0.74

-

0.74

-

-

5.58

0.37

10.04

1.49

-

-

19


SCIENTIFIC NAMEIctaluridae - bullhead catfishes

Ictalurus furcatus (Lesueur, 1840) Ictalurus punctatus (Raflnesque, 1818) Noturus gyrinus (Mitchill, 1817) Noturus murus Jordan, 1877

Noturus phaeus Taylor, 1969 Pylodictis olivaris (Rafinesque, 1818)

Esocidae - pikes Esox americanus vermiculatus Lesueur, 1846


Cyprinodontidae - kiliifishes

Fundulus olivaceus (Storer, 1845)

Poeciiiidae - livebearersCambusia affmis (Baird and Girard, 1853)

Atherinidae - siiversides Labidesthes sicculus (Cope, 1865) Meriidia berylina (Cope, 1866)

Percichthyidae - temperate basses Morone chrysops (Rafinesque, 1820)

Morone saxatilis (Walbaum, 1792) Centrarchidae - sunfishes

Lepomis cyanellus Rafinesque, 1819 Lepomis gulosus (Cuvier, 1829) Lepomis humilis (Girard, 1858) Lepomis macrochirus Rafinesque, 1819 Lepomis marginatus (Holbrook, 1855) Lepomis megalotis (Rafinesque, 1820)

Lepomis species Micropterus punctulatus (Raflnesque, 1819) Micropterus salmoides (Lacepede, 1602) Micropterus species Pomoxis annularis Rafinesque, 1818 Pomoxis nigromaculatus (Lesueur, 1829)

Percidae - perches

Etheostoma fusiforme (Girard, 1854)

Etheostoma stigmaeum (Jordan, 1877)

Percina maculata (Girard, 1859) Percina sciera (Swain, 1883)

Sciaenidae - drums Aplodinotus grunniens Raflnesque, 1819

Unclassified fishes Unknown fry

/96

-

;3.89

1.50

0.30

-

0.90

0.30 0.30

~

-

-

2.10

5 $

1CO97

2.01

;-

18.47

0.40

-

1.20 040 0.80 1.20

1.61 0.40

0.80

0.40

-

-

3.21

-

98

0.57 0.71

-

-

0.71

0.71

-

0.43

0.43 1.85

0.43

1.28

0.14

-

1.56

96

1.02

1.02

-

306

1.02

11.22 2-04

5.10

3.06 2.04

-

-

9.18

1 £p

/97

3.43

0.57

0.57

1.14

-

571

--

1.14 10.29

1.14

12.57

-

-

686

4.57

;

98

1.60 0.40

0.40

-

31.20

-

0.80 0.40 2.40

1.20

23.60

3.20 0.40

-

-

5.60

96

025

0.25

3.82

0.25

0.25

-

0.25

2.04 1.02 2.04

1.02

0.25 2.80 0.25

0.25

0.51

-

0.51

0.51

0.25

<D

1 1

97

-

0.53

0.53

11.05

1.05

-

1.58

11.58 1.05 2.11

O CO

5.26 1.05

1.05

6.84

3.68

0.53

98

-

0.44 0.44

3.49

-

-

0.44 0.44

5.68

13.10 n o."7

8.73 1.75 1.31

-

0.44

1.75

1.31

96

1.87 1.64

-

1.64

19.16

1.87

1.17

2.10 2.10

3.04

13.79 0.93

-

--

3.50

1 1

97

3.89 3.50

0.39

-

16.34

5.45

0.39

0.39

5.45

0.39

10.12

0.78

5.45

-

-

4.67

98

0.42 2.15

0.10

0.05

0.21

6.44

0.16 0.05

0.05

-

0.31 1.10 0.47 14.51

0.26

042

0.42

6.13 0.26

-

-

320

iIf

98

0.15

122

015

0.91

6.23

-

5.17 2.74 19.45

22.34

1.37

1033 0.61

3 34

030

-

380

7J /I

98

0.64

-

-

26.05

2.89

-

1.61 0.96 3.22 0.96

9.32

2605

1.61

2.89

-

-

1.61

yj /;

98

0.80

-

11.66

-

1.28 0.32 0.16

2.72

2.56

0.16

0.48

-

-

2.88

iI98

020

0.20

7.14

0.40

0.20 0.60 3.17 4.96

1.98

0.60

1.79

7.34 1.39

1.19

0.40

0.20

1.98

> / SF

/ 6

/«?98

0.37

0.74

0.37

1.12

27.51

-

0.37 3.35 0.37 5.95

5.20 0.37

1.86

4.83

9 en

1.12

0.37

2.97




Petromyzontidae species Lepisosteldae - gars

Lepisosteus oculatus (Winchell, 1864) Leplsosteus osseus (Linnaeus, 1758) Lepisosteus p/atostomus Rafinesque, 1820 Lephoit-u" svecies

Amiidae - bowfins Amiacalva Linnaeus, 1766

Hiodontidae - mooneyes Hiodon alosoides (Rafinesque, 1819)

Anguiiiidae - freshwater eels Angullla rostrata (Lesueur, 1817)

Ciupeidae - herrings Dorosoma cepedianum (Lesueur, 1818) Dorosoma petenense (Gunther, 1867) Dorosoma species

Cyprinidae - carps and minnows Ctenopharyngodon idella (Valenciennes, 1844) Cyprinella camura (Jordan and Meek, 1884) Cyprinella lutrensis (Baird and Girard, 1853) Cyprinella spiloptera (Cope, 1868) Cyprinella venusta Girard, 1856 Cypmus carp/'o Linnaeus, 1758 Cyprinidae species Hybognathus hayi Jordan, 1885 Hybognathus nuchalis Agassiz, 1855 Lythrurus fumeus (Evermann, 1892) Lythrurus umbratltis (Girard, 1856) Macrhybopsis aestivalis (Girard, 1856) Macrhybopsis storeriana (Kirtland, 1847) Notemigonus crysoleucas (Mitchill, 1814) Notropis ammophilus Suttkus and Boschung, 1990 Notropis amnis Hubbs and Greene, 1951 Notropis atherinoides Rafinesque 1818 Notropis buchanani Meek, 1896 Notropis chalybaeus (Cope, 1869) Notropis maculatus (Hay, 1881) Notropis texanus (Girard, 1856) Notropis volucellus (Cope, 1865) Notropis species Opsopoeodus emiliae Hay, 1881 Phenacobius mirabilis (Girard, 1856) Pimephales notatus (Rafinesque, 1820) Pimephales promelas Rafinesque, 1820 Pimephales vlgilax (Baird and Girard. 1856) Semotilus atromaculatus (Mitchill, 1818)

Catostomidae - suckers Carp/odes carpio (Rafinesque, 1820) Carp/odes cyprinus Lesueur, 1817 Erimyzon ob/ongus tMitchiil 1814) Enmyzon species Hypentelium nigricans (Lesueur, 1817) Ictiobus bubalus (Rafinesque, 1818) Ictiobus cyprinellus (Valenciennes, 1844) Ictiobus niger (Rafinesque, 1819) Ictiobus species Minytrema melanops (Rafinesque, 1820) Moxostoma macrolepidotum (Leuseur, 1817) Moxostoma poecilurum (Jordan, 1877) Moxostoma species Catostomidae species

/ «

/I s98

2.74

1.00

8.48

-

1.25

0.75 3.49

-

-

-

-

-

-

8.98

::-

0.25

0.25

-

i * isI $ $98

1.38

0.69

3.81

-

~

6.57 4.50 3.46

-

-

-

0.35

-

-

-

-

-

2.08 0.35

-

98

2.07

6.22

021

2.07

-

-

2.07

-

-

-

_

-

-

-

-

-

4.36 2.90 0.21 34.85

-

* i$ S

/ 0° / * / <8 / O

98

4.37

1.87 021

6.65

0.21

19.13

3.74 104

1.25

-

-

1.04

-

-

-

-

2.49 0.21

0.21

-

98

0.21

0.10

-

20.44

-

-

0.10 1.88

-

-

-

-

-

-

-

-

:;

2.50

-

/l/l/o198

0.370.37

0.19

1.85 0.19

-

3.52 1.67

0.19

-

1.67

2.04

20.74 0.74

-

3-33

0.74

-

0.19

0.37

0.56

019

98

0.330.66 1.32

~

0.33

-

1.32 46.86

6.60 9.90

-

0.33

0.33

0.66 2.97

-

-

-

2.64

-

0.66 0.33

-

98

1.56

-

46.09

-

2.08

-

-

0.52

-

-

-

-

-

-

? / c

^

$98

055

0.18

0.37

0.18

-

-

37.94 0.18 1.84

-

-

0.18

0.18

2.39

0.37

-

2.03

2.21

0.18

0.55

-

/ *

s 398

0.28

0.56

028

2.81

-

-

1.69

-

-

-

-

0.84

-

13.76

::

3.09 0.56 0.84

-

/ Cj98

3.700.34 0.67

-

3.70

-

0.34

0.34 10.44 0.67

-

-

0.34

1.68

-

-

-

0.34

0.67

17.51 3.03 2.02

-

//98

0.600.12 0.12

0.24

0.12

-

-

10.91 1.44 6.12

-

-

6.71

1.08

0.12

-

8.75

2.68 0.12

1.68 0.12 0.48

0.24

//98

1.370.29

-

-

-

1.46

23.03 1.17

2.62

0.29

-

12.54

0.87

2.62

0.58

7.87

-

0.58

0.29

//98

101

0.50

0.84

6.88

-

-

12.25

2.35 1.68

-

-

21.31

2.18

-

0.17

1.17

3.02

0.84

/<!98

4.19

2.99

-

3.59

8.38

-

1.80 22.16

-

060

-

-

-

0.60

5.39

0.60 7.19

//98

1.03

1.37

-

31.51 3.42 2.05

-

1.03 2.40

0.34

-

-

_

-

-

3.42

-

0.68

//98

i 1.37

0.19

0.59

-

0.20

-

-

-

-

-

-

~

21


SCIENTIFIC NAMtIctaluridae - bullhead catflshes

Ameiurus me/as (Rafinesque, 1820) Ameiurus natalis (Lesueur, 1819) Ictalurus furcatus (Lesueur, 1840) Ictalurus punctatus (Rafinesque, 1818) Noturus gyrinus (Mitchill, 1817) Noturus miurus Jordan, 1877 Noturus nocturnus Jordan and Gilbert, 1886 Notuws phaeus Taylor, 1969 Pylodictis olivaris (Rafinesque, 1818)

Esocidae - pikes Esox amencanus vermiculatus Lesueur. 1846


Cyprinodontidae - kllilflshes Fundulus chrysotus (Gunther, 1866) Fundulus notatus (Rafinesque, 1820) Fundulus olivaceus (Storer, 1845) Fundulus species

Poeciiiidae - iivebearers Gambusia affinis (Baird and Girard, 1853)

Atherinidae - silversides Labidesthes sicculus (Cope, 1865) Menidia beryllma (Cope, 1866)

Percichthyidae -temperate tis-se« Morone chrysops (Rafinesque, 1820) Morone mississippiensls Jordan and Eigenmann, Morone saxatilis (Walbaum, 1792)

Centrarchidae - sunfishes AmbloplXes ariommus Viosca, 1936 Elassoma zonatum Jordan, 1877 Lepomis cyanellus Rafinesque, 1819 Lepomis gulosus (Cuvier, 1829) Lepomis humilis (Girard, 1858) Lepomis macrochirus Rafinesque, 1819 Lepomis marginatus (Holbrook, 1855) Lepomis megalotis (Rafinesque, 1820) Lepomis micro/ophus (Gunther, 1859) Lepomis minlatus Evermann, 1899 Lepomis hybrid Lepomis species Micropterus punctulatus (Rafinesque, 1819) Micropterus salmoides (Lacepede, 1802) Micropterus species Pomoxis annularis Rafinesque, 1818 Pomoxis nigromaculatus (Lesueur, 1829) Pomona species

Percidae - perches Ammocrypte beam Jordan, 1877 Ammocrypta clara Jordan and Meek. 1885 Ammocrypta vivax Hay, 1882 Etheostoma asprigene (Forbes, 1878) Etheostoma chlorosomum (Hay, 1881) Etheostoma fusiforme (Girard, 1854) Etheostoma gracile (Girard, 1859) Etheostoma histrto Jordan and Gilbert, 1887 Etheostoma proeliare (Hay, 1881) Etheostoma stigmaeum (Jordan, 1877) Percina caprodes (Rafinesque, 1818) Percina maculata (Girard, t859) Percina sciera (Swain, 1883)

Sciaenidae - drums Aplodinotus grunniens Rafinesque, 1819

Unciassified fishes Unknown fry

S"

98

0.50

0.25

-

54.11

025

-

0.50 0.75 150

2.99

1.00 2.24 3.74

3.99 0.25

-

0.50

025

/<§>

98

0.69 0.35

0.69

0.35

-

42.91

-

0.35

0.35 1.38 0.35

20.42

1.73

1.73 0.35

-

5.19

£*98

-

0.41

-

1598

-

:1.24 1.24 3.32 3.32

0.41

13.69

0.21

2.28 0.21

-

-

2.70

*98

0.21

-

44.70

0.21

-

;;1.25 0.62 1.04 2.08

2.70

0.21

1.66

:

-

2.91

/cf98

0.10 0.10 0.42 0.10

-

-

69.97

-

1.67 0.10

0.63

0.10

0.73 0.52

;;-

0.31

/<?98

0.19 1.11

0.56

0.56

0.37

-

0.74

1.85 0.37

3.89

13.33 0.56

24.81 5.93 1.11 1.30 2.41

0.19

-

0.19

1.67

/ <J98

0.33 0.66

0.33

-

2.31

7.26

;;0.66 0.33

4.95

0.33

429 0.33

;;-

2.97

*98

0.26

0.26 0.26

0.26

-

22.40

-

;;1.04 3.65 5.99

3.13

7.29

1.30

3.65

:

-

0.26

/<?98

0.18

1.84 0.37

-

0.18

25.23

-

4.42 0.18 4.24 0.18

9.76

1.66 0.55

0.55

0.18

-

1.10

-j98

1.40

0.56

0.84

0.56

1.69

-

1.69 6.18 0.84 23.88

10.39 0.56 0.84

3.93

13.76 2.25

028

-

5.62

1 598

2.69

1.01

3.70

13.80

-

;;0.34 2.36 7.41

1.35 0.34 0.34

3.03 1.01

4.04 0.34

6.06

1.35

5.05

*98

0.12

1.08 0.12

0.72

2.88

28.42

0.60

-

0.12

6.71 1.20

1.44

7.67 0.12 120

2.40 1.20 0.48

0.24 0.24

0.72

0.12

-

036

/I98

2.62 0.58

-

7.00

0.29

-

058

3.21

0.29

16.62

3.79 7.29 0.29

-

0.29

-

-

146

0.29

§98

1.68 1.34 0.50 5.87 1.01

-

1.34

503

0.34

1.34

0.50

3.02

4.19 0.17

3.02

2.68

3.52 0.34

0.84 0.34

-

-

8.72

98

1.20 1.80 0.60

4.19

-

14.37

-

;;0.60

0.60

8.38

0.60

7.78

-

-

-

2.40

Ij

98

0.34 0.34

1.03

0.34

0.34

-

13.36

0.34

2.05 1.71

1.37 0.34

2.74

0.68

19.86 0.34

3.42

__

-

-

4.11

*98

-

1.37

0.39

0.20

5.29

10.20

11.18 5.69 0.20 21.18

30.98 1.96 5.10 0.39 0.39

2.35 0.20

"-

-

0.20

020





Lepisosteus oculatus (Winchell. 1864)Lepisosteus osseus (Linnaeus, 1758)Lepisosteus platostomus Rafinesque, 1820Lepisosteus species

Amiidae - bowfinsAmia calva Linnaeus, 1766



Clupeidae - herringsDorosoma cepedianum (Lesueur, 1818)Dorosoma petenense (Gunther, 1867)Dorosoma species

Cyprinidae - carps and minnowsCtenopharyngodon idella (Valenciennes, 1844)Cyprinella camura (Jordan and Meek, 1884)Cyprinella lutrensis (Baird and Girard, 1853)Cyprinella spiloptera (Cope, 1868)Cyprinella venusta Girard, 1856Cyprinus carplo Linnaeus, 1758Cyprinidae speciesHybognathus hayi Jordan, 1885Hybognathus nuchalis Agassiz, 1855Lythrurus fumeus (Evermann, 1892)Lythrurus umbratilis (Girard, 1856)Macrhybopsis aestivalis (Girard, 1856)Macrhybopsis storeriana (Kirtland, 1847)Notemigonus crysoleucas (Mitehill, 1814)Notropis ammophilus Suttkus and Boschung, 1990Notropis amnis Hubbs and Greene, 1951Notropis atherinoides Rafinesque, 1818Notropis buchanani Meek, 1896Notropis chalybaeus (Cope, 1869)Notropis maculatus (Hay, 1881)Notropis taxanus (Girard, 1856)Notropis volucellus (Cope, 1865)Notropis speciesOpsopoeodus emiliae Hay, 1881Phenacobius mirabilis (Girard, 1856)Pimephales notatus (Rafinesque. 1820)Pimephales promelas Rafinesque, 1820Pimephales vigilax (Baird and Girard, 1856)Se- LI romaculatus (Mitehill, 1818)

Catostomidae - suckersCarp/odes carpro (Rafinesque, 1820)Carp/odes cyprinus Lesueur, 1817Erimyzon oblongus (Mitchiil, 1814)Erimyzon speciesHypentelium nlgricans (Lesueur, 1817)Ictiobus bubalus (Rafinesque, 1818)Ictiobus cyprinellus (Valenciennes, 1844)Ictiobus niger (Rafinesque, 1819)Ictiobus speciesMinytrema melanops (Rafinesque, 1820)Moxostoma macrolepidotum (Leuseur, 1817)Moxostoma poecilurum (Jordan, 1877)Moxostoma speciesCatostomidae species

98

2.30

1.15

7.28

3.07

34.87

1.15 -

3.456.13

0.38 -

98

0.27

0.14

6.84

0.14

0.6830.230.68 -

1.6418.60

2.19

0.14

0.411.23

0.27

2.60

1.64

1.09 -

S8

--

5.34

3.05

1.533.82

49.623.05-

5.34 -

2.29

0.76

1.53

2.29 -

98

0.170.170.34

3.42-

1.373.081.03

2.39 --

0.17

0.17

0.85

15.90

1.882.050.17 - -

98

0.29

1.17

1.47

5.575.87

5.87

-

1.17

3.81 -

98

1.190.090.090.09

0.09

6.21

0.34 -

2.55

0.26-

1.190.510.090.77 -

98

0.79 -

3.3518.938.48

39.251.78

0.79 - -

1.780.59

0.20 --

0.20

0.20 - -

98

4.81

0.34

0.34

-

0.69 -

0.69 -

23

Table 4. Percent relative abundance of fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98 Continued

SCIENTIFIC NAMEIctaluridae - bullhead catflshes

Ameiurus me/as (Rafinesque, 1820)Ameiurus natalis (Lesueur, 1819)Icta/urus furcatus (Lesueur, 1840)Ictalurus punctatus (Rafinesque, 1818)Noturus gyrinus (Mitchill, 1817)Noturus miurus Jordan, 1877Noturus nocturnus Jordan and Gilbert, 1886Noturus phaeus Taylor, 1969Pylodiclis olivaris (Rafinesque, 1818)

Esocidae - pikesEsox americanus vermiculatus Lesueur, 1846


Cyprinodontidae - kiilif ishesFundulus chrysotus (Gunther, 1866)Furidulus notatus (Rafinesque, 1820)Fundulus olivaceus (Storer, 1845)Fundulus species

Poeciiiidae - iivebearersGambusia affinis (Baird and Girard, 1853)

Atherinidae - siiversidesLabidesthes sicculus (Cope, 1865)Menidia beryllina (Cope, 1S66)

Percichthyidae - temperate bassesMorone chrysops (Rafinesque, 1820)Morone mississippiensls Jordan and Eigenmann,Morone saxatilis (Walbaum, 1792)

Centrarchidae - sunflshmAmbloplites ariommus viosca, 1936Elassoma zonatum Jordan, 1877Lepomis cyanellus Rafinesque, 1819Lepomis gulosus (Cuvier, 1829)Lepomis humilis (Girard, 1858)Lepomis macmchirus Rafinesque, 1819Lepomis marginatus (Hdbrook, 1855)Lepomis megalotis (Rafinesque, 1820)Lepomis microlophus (Gunther, 1859)Lepomis miniatus rmarm, 1899Lepomis hybridLepomis speciesMicropterus punctulatus (Rafinesque, 1819)Micropterus salmoides (Lacepede, 1802)Micropterus speciesPomoxis annularis Rafinesque, 1818Pomoxis nigromaculatus (Lesueur. 1829)Pomoxis species

Percidae - perchesAmmocrypta beam' Jordan, 1877Ammocrypta clara Jordan and Meek, 1885Ammocrypta vlvax Hay, 1882Etheostoma asprigene (Forbes, 1878)Etheostoma chlorosomum (Hay, 1881)Etheostoma fusiforme (Girard, 1854)Etheostoma gracile (Girard, 1859)Etheostoma histrio Jordan and Gilbert, 1887Etheostoma proeliare (Hay, 1 881 )Etheostoma stigmaeum (Jordan, 1877)Percina caprodes (Rafinesque, 1818)Perdna maculata (Girard, 1859)Percina sciera (Swain, 1883)

Sciaenidae - drumsAplodinotus grunniens Rafinesque, 1819

Unclassified fishesUnknown fry

98

--__,_-

~

0.38--

2.30

-

.-__-

0.381.5313.03

0.38 --

9.96

6.904.60-

-- ~ --

0.77

-

98

-~

1.92 __-

0.14

0.27

0.41

8.48

0.82

-

0.410.820.143.42

8.89

0.14

0.140.412.33

0.140.41

-- -~ --

0.14

1.78

-

98

-

3.051.53 _

0.76

0.76

--- -

3.82

-

---

-_- «-

0.76 - -

0.760.76

0.76--

- - -

-

8*0

-

98

- -

0.17 ___ -

0.51

3.59

3.76

0.17--

0.17__

2.052.390.179.57

17.26

2.055.980.85

0.510.17

--

0.680.1711.62

0.17

0.34-

3.59_

085

-

98

-

029-

0.59 __---

0.88

0.59

-

3.81-

40.76

-

---

0.290.290.291.76

6.74

9.38

1.76

1.172.64-~

1.47

- -

0.29 -

0.59

1.17

-

98

- -

0.17--_.---

- -

49.66

0.340.09-

.-__

1.45

4.082.04

1.19

22.110.090.60

2.550.51

-- -~ --

1.70

-

98

- -

0.59 __-

0.79

---~-

18.54

---

__

0.39

0.39-

1.58--

0.39

-- --- -- __

0.99

-

98

-0.34 __- -

1.37

-

1.03-

6.19

---

__

3.782.411.03

46.740.698.9310.654.12 -

412

0.69

-

1.03 -- ~

-

-

c en O§3 I-o33


occurrence of fishes at 38 sites in the mississippi ... · information for 38 fish sampling sites...

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