Sarawak & Sabah, Borneo

26th March – 15th April 2017

Leader: Mike Nelson

Participants: Daniel Bombaert, Rob Fray, Paul Harvey. Sarawak only: John Clark & Tim Doran. Sabah only: Nigel Adlam, Christian Perez and Greg Anderson.

Blue-banded Pitta © Mike Nelson/Birdtour Asia

With our Sarawak extension firmly established as a hotspot for species like Black Oriole, Hose’s Broadbill, Dulit Frogmouth, Hook-billed Bulbul and Scarlet-breasted Flowerpecker, along with many of the islands’ other endemics being readily available, has made this a very desirable destination, indeed this was our 13th tour here. We managed great views of all of those specialties with several days spent in the Crocker Range of Sarawak and it was a perfect combination with the riches of Sabah. Our first foray into the foothills of Sabah included Whitehead’s Broadbill and a pair of endemic Barbets with Mountain and Bornean as well as an obliging Bock’s Hawk-Cuckoo. At the avian rich Mount Kinabalu our list included such mouthwatering numbers as the Whitehead’s trio of Trogon, Spiderhunter and Broadbill, plus Everett’s Thrush, Mountain Blackeye,

Crimson-headed and Red-breasted Partridges and Fruithunter. Sepilok provided us with a nice lowland forest interlude with a surprise group of three Bristleheads along with Black-crowned Pitta, then we were off to the Gomantong caves to check out the ‘nest’ swiftlets before spending a few days on the majestic Kinabatangan River. Although the water levels were unusually high on the river we managed some fantastic hornbill viewing including Black, Oriental Pied, Wreathed, Wrinkled, Rhinoceros and White-crowned. Night birding gave us some stunning views of Large Frogmouth and an obliging Brown Wood Owl. It was then off to one of Borneo’s highlights, the incomparable Borneo Rainforest Lodge in the heart of the Danum Valley. Several days here gave us a suite of babblers including the endemic Black-throated Wren-Babbler and Bornean Ground Babbler. A host of bulbuls reside here and stars among these were Straw-headed and Grey-bellied. A fruiting tree attracted a nice pair of Helmeted Hornbills to complete the hornbill set and a close Cinnamon-rumped Trogon gave us walk away views, a group of noisy Bristlehead followed us down one of the many trails for a couple of hours as we added other species including Moustached Hawk-Cuckoo, Banded and Dusky Broadbill, Large-billed and Bornean Blue Flycatchers as well as other standouts that included, Blue-headed Pitta, Diard’s Trogon, Rufous-tailed Shama and Rufous-chested Flycatcher. All this despite some unseasonable rains that had us soaked on many days. We managed 310 species with 48 endemics and a nice list of 30 mammals.

Dulit Frogmouth © John Clark, and Grey-breasted Babbler

The Sarawak extension began with an early morning visit to a remnant patch of peat swamp. This is Borneo’s most threatened habitat so a trip here always gives us a chance for species seen nowhere else on the tour. Arriving at dawn let us spook a few Large-tailed Nightjars off the road and during breakfast we admired the huge collection of insects that had gathered around the lights and the palm tree at the entrance stirred with many Asian Palm Swifts dropping out forage for the day. After filling up with coffee and breakfast we were eager to begin our first birding in Borneo and set off along the trail where we managed to clean up our targets with fantastic views of Hook-billed Bulbul, Grey-breasted Babbler, Red-crowned Barbet, Scarlet-breasted Flowerpecker and Black-throated Babbler – what a start! Additional goodies came in the form of Red-billed Malkoha, Great Slaty Woodpecker, Black-winged Flycatcher-shrike and an unexpected female Jambu Fruit Dove. From here we spent the rest of the day travelling deeper into Sarawak along the impressive Crocker Range, to a small village in the foothills in preparation for our stay in the mountains.

The following two days were spent around a camp in the western end of the Crocker range looking for one of Borneo’s least-known endemics, the Black Oriole. We managed several views of this bird over the days with a particularly friendly bird around the camp that was joined by his partner on several occasions singing back and forth in the canopy. We also had crippling views of Hose’s Broadbill after it flew back and forth across the trail showing its bright blue belly it then perched up and turned to face us for some outrageous looks. Another

target here was Bornean Banded Pitta and though it always seemed to be just out of visual range, with some persistence we managed to get some views of it as it circled us once running along a log. Pygmy White-eye (or Heleia), Bornean Bulbul and Bornean Leafbird were quite common up here with a supporting cast of Cinereous Bulbul, Black-bellied Malkoha, Maroon-breasted Philentoma and Mountain Barbet as some of the standouts.

Another denizen of the Crocker range is the Dulit Frogmouth, a once enigmatic species seen by very few people but since we discovered it here, it has been seen on a regular basis and the second leg of our Sarawak tour took us farther south along the mountain range to another small village. Here we had several predawn and post dusk sessions for the frogmouth and secured some nice views one evening. Another section of road passes through some wonderful montane forest habitat and a full day birding along here enabled some staggering looks at Blue-banded Pitta sat atop a huge fallen tree giving us prolonged views. Having tried for it in a few spots we could hear one close to the road and creeping down into the cover of the forest we waited at the base of the fallen tree and obligingly it hopped up onto it and began to sing as we all peered down at it. Both Bock’s Hawk-Cuckoo and Sunda Cuckoo were seen here along with Bornean and Whitehead’s Spiderhunter, the latter showing very well uttering it unusual kazoo-like call, Mountain Serpent Eagle was found circling below the clouds and Mountain and Bornean Barbet were constant companions with Temminck’s Sunbird adding a nice splash of color. With most of our targets in the bag we headed back to Kota Kinabalu to begin the tour of Sabah, though due to a cancelled flight it took several more hours than expected, arriving into Kota Kinabalu late and ready for a decent bed!

Red-crowned Barbet and Hose’s Broadbill © Rob Fray

An early morning drive up into the Crocker Range kicked us off with a fantastic Whitehead’s Broadbill bounding back and forth across the road perching restlessly on several occasions. Buoyed by this great bird we continued along the road picking through mixed flocks and solo birds including Blue-and-white Flycatcher, Chestnut-crested Yuhina, Black-and-crimson Oriole, Yellow-bellied Warbler, Bornean Leafbirds, Mugimaki Flycatcher, Chestnut-hooded Laughingthrush including one doing battle with a stick insect the length of a forearm, Bornean Treepie, a pair of wonderfully coloured Long-tailed Broadbills and Blyth’s Hawk-Eagle. Several Mountain Barbets were found and a diminutive Bornean Barbet was picked up in the tail end of a small flock. Our last bird before lunch was a particularly cooperative Bock’s Hawk-Cuckoo which was a nice pick up for those just joining the trip and better views for those from the Sarawak extension.

After a nice morning we ended up at the base of Mount Kinabalu for our five-night stay and got stuck in straight away along one of the many great trails through this moss draped forest. Bornean Whistler was quite

evident along with mixed flocks of Bornean Green Magpie, Chestnut-hooded Laughingthrush and Grey-throated Babbler along with other montane species like Bornean Forktail, Indigo Flycatcher, Black-capped White-eye, Sunda Bush Warbler and Ochraceous Bulbul – the birds in Borneo both a mooted ‘Field Guide’, and Birdlife split, Penan Bulbul.

The following four days spent up here were a mix of rain and overcast skies but even in the pouring rain we still managed some cracking species like a wonderful pair of Fruit-hunter. These two were quite a fortuitous find as we had hurried to find shelter during a heavy downpour when above the roof sat this pair, the chestnut colored female showing particularly well. We all braved the elements for a few seconds to get good looks at this pair before ducking back under to escape getting completely sodden, no easy task. Crimson-headed Partridge were quite numerous with several calling their unpartridge-like songs around us and with patience many crossed the path in front of us (so, unsurprisingly it turns out they aren’t actually a partridge, but a primitive pheasant-pheasant, and renamed Bloodhead in the recent Indonesian field guide). Bare-headed Laughingthrush was a last minute addition with a nice pair over the road, heads bouncing up and down as they sang. For those whose high pitched hearing has gone Bornean Stubtail can be a bit of a challenge to find hiding underneath the mountain vegetation but several goes at this little critter got us all onto it. Sunda Laughingthrush were often seen in raucous, roving bands bounding through the canopy attracting many stragglers looking to take advantage of the spooked prey items., Whitehead’s Broadbill turned up on a few occasions, always a crowd pleaser with its iridescent green plumage and a Whitehead’s Spiderhunter was tracked down for those not on the Sarawak leg plus an outrageous Sunda Cuckoo that landed close by and sat for ages for us.

Our best morning turned out to be our one sunny morning that began with a large mixed flock around the power station with a noisy group of Bornean Green Magpie, Sunda and Chestnut-hooded Laughingthrushs, these were soon joined by a Bornean Whistler, darting out to capture insects drawn to the lights and soon enough Mountain Leaf and Yellow-breasted Warbler joined the fray, White-throated Fantails sat atop the fence looking to pick up a tasty morsel while a pair of Eye-browed Thrush hopped along the path farther down from us. Several Sunda (Ashy) Drongo’s shot past, agile on the wing like fighters swooping down on their prey, Indigo Flycatcher, eyelevel Grey-chinned Minivet, Blyth’s Shrike-Babbler and a pair of Bornean Whistling Thrush added to the frey and Mountain Treeshrew hung out with us waiting for us to “drop” a tidbit for it, all this while trying to sip our coffee and enjoying breakfast.

Wanting to take advantage of the clear weather we hit the trails and were rewarded with a fantastic threesome of Whitehead’s Trogon with a male and two females showing right next to the trail completing the Whitehead’s trio in grand style. One of the viewpoints gave us a nice flock of Giant Swiftlet that were slowly pushed down by the cloud eventually just meters above us singing their rather un-swiftlet song while below us a circling raptor turned out to be a Mountain Serpent Eagle complete with serpent dangling from its talons. Golden-naped Barbet, Maroon Woodpecker, Sunda Cuckooshrike and further views of some very showy Crimson-headed Partridges before the heavens opened up and forced us to take cover. On another day we spent the whole morning along one of the trails picking up a nice group of Red-breasted Partridge as they filed across the trail, first in a group of three then two more followed by another three and a single tail end Charlie. They had been gathering nervously along the side of the trail showing well but when they wanted to cross they wasted no time getting to the other side. Another enjoyable spectacle was farther along when we found a pair of Mountain Wren-Babblers close by the side of the trail feeding and foraging quietly but when they suddenly burst into song seven more sprang out from parts unseen shooting out from cover as if someone had set off a Wren-Babbler grenade scattering vocal birds all around us. They continued to bounce around the understory calling back and forth showing particularly well and we could still hear them singing away as we continued farther down the trail. Another nice show was a captivating pair of Bornean Forktails that allowed us prolonged views as they foraged along the trail in front of us.

On a particularly foggy morning we chanced our luck along the roadside predawn and had a group of three Eye-browed Jungle Flycatchers (actually not a flycatcher at all, Bornean Shade-dweller being an imaginatively newly-coined name for this robin) feeding in the dim light. Making their way down towards us we watched as they darted onto the road to collect prey then sat up on a low perch waiting for another morsel to come along. Once it was light enough we could make out some fine details of this smart looking bird before we headed up to another mountain trial for the rest of the morning and it started well with nice views of Crimson-headed Partridge and a nice flock of Sunda and Chestnut-hooded Laughingthrush but there was no sign of our

intended target. Several forays into the undergrowth gave us nothing but doggedly we continued along the trail. We had all but given up hope and were checking the last stretch of trail and had stopped, looking down a steep ravine we picked up the call of an Everett’s Thrush. We could hear it round a blind curve to our left so waited patiently as it continued to edge closer. Then a brief hint of motion down the slope gave us a shot and there on a bright green mossy log was the bird. We scrambled to get everyone on the bird but it moved before that happened but we could see where it was going. Then Daniel picked up another bird as it was joined by the first bird, a pair together below us, tantalizing through a narrow window and still not seen by all. They moved over another bank and were gone. They fell silent and we peered into the dank depths of the ravine below us for any hint of movement but there was none. We waited a while longer then suddenly came a very close call and all eyes whipped to the left to see one of the pair next to the trail hopping up onto a small stump for some stunning views. Quickly gesticulating to the others who were farther along scanning we quickly bunched up and watched as the bird moved along from branch to branch with the final views out in the open before it dropped down and descended the slope through the moist carpet of leaves quickly disappearing but forever etched in our minds.

Whitehead’s Trogon © Rob Fray and Bornean Leafbird

Having managed a good haul of Mount Kinabalu’s specialties we set off into the lowlands at Sepilok and its famous canopy tower where we spent the afternoon. A whole new suite of birds awaited us here with Asian Drongo-Cuckoo, Diard’s Trogon, Grey-rumped Treeswift, Sunda Woodpecker, Dark-throated Oriole, Fiery Minivet, Streaked Bulbul, Black-and-yellow Broadbill and Silver-rumped Spinetails. A nice group of very noisy Bushy-crested Hornbills paid the tree above us a visit for some nice close looks and after sifting through many Mugimaki Flycatchers we found a nice yellowish eye-browed Narcissus Flycatcher – a scarce wintering migrant. We also had several mammal encounters; with a mother and young Orangutan seen well foraging up a huge tree across the river from us, a Sunda Colugo fleeing from a Wallace’s Hawk-Eagle that was itself being pursued by a pair of noisy Greater Racket-tailed Drongos and the flying carpet Red Giant Flying Squirrel which glided from its high canopy perch down to a distant tree to forage.

A morning along the forest track here gave us our first great looks at Black-crowned Pitta sitting about ten meters up and singing away, a nice pair of Black-capped Babblers, Rufous Piculet, White-chested Babblers, Raffles’s and Chestnut-breasted Malkoha, Hairy-backed and Buff-vented Bulbul, Long-billed Spiderhunter, and several lowland babblers with Chestnut-winged, Fluffy-backed and Bold-striped Tit-Babbler making noisy groups. A very close calling male Rufous-collared Kingfisher sat for a prolonged period allowing us to eye its magnificent colours. We also had a nice pair of Copper-throated Sunbirds feeding on flowers in the car park but best of all though was a surprise group of three Bristleheads that showed quite well next to the trail for a few minutes giving us one of Borneo’s most desired birds and monotypic family, a real bonus as we haven’t

seen it at this site for the past few tours. With that huge find, and a relief to many, we headed to Gomantong caves for the afternoon here picking up the nest swiftlets of Black and Mossy-nest Swiftlet but harvesting of the Edible-nest Swiftlets meant there were none to be found on the cave walls but were seen in good numbers hawking for insects outside.

Continuing on from the caves we headed to Sukau and the Kinabatangan River for the remainder of the afternoon. Patrolling one of the tributaries we slowly cruised the murky waters picking up Little Green Pigeon, White-bellied Fish Eagle and Lesser Fish Eagle, a slew of hornbills including Oriental Pied, Bushy-crested, Wrinkled, Black and Rhinoceros. A nice perched Storm’s Stork was the first of several we found, Green Imperial Pigeons were conspicuously perched out while a noisy and gaudy Stork-billed Kingfisher showed well. Black-and-red Broadbills added their outrageous hues to the wall of green around them and a nice pair of Buff-rumped Woodpeckers rounded off the day time birding. After dinner some night birding gave us a very confiding (Bornean) Brown Wood Owl perched above us and after a little searching we tracked down the calling Large Frogmouth right next to one of the bungalows.

A full day on the river allowed us to pick up some new species with White-crowned Hornbill, Oriental Darter, Changeable Hawk-Eagle, Crested Goshawk, Malaysian Blue Flycatcher, Scarlet-rumped Trogon, Bornean Brown Barbet, Rufous-backed Kingfisher, an iridescent lavender-coloured Ruddy Kingfisher very close, a nice White-fronted Falconet devouring a dragonfly, circling Lesser Adjutant, one of many vocal Hooded Pittas was also located along with Green and Common Iora and just before the heavens open up and poured down on us we watched as thirty-plus Wrinkled Hornbills came flooding out of the nearby oil palms in threes and fours. They all landed close by for some excellent looks at their flamboyant colours and just seemed to keep coming till the rain got so strong that we were soon joined by another boat full of tourists under the eaves of a riverside shelter. Once the rain had subsided we got onto Sunda Pied Fantail, Yellow-bellied Prinia and White-breasted Waterhen before heading back onto the river.

Another feature of the river is the primates that always seem to hang out in droves in the trees lining the waterways. Several large groups of Proboscis Monkeys were seen foraging next to the river. A few spectacular males with their huge plate like floppy noses were seen surveying their domains while feisty little ones cavorted in the branches around them. Long-tailed Macaques were also very common and no less boisterous but the calm order was maintained by a few more solemn Sunda Silvered Langurs that seemed much calmer as they perched high in the boughs watching the world go by.

The next morning, we did some further birding down another tributary where we found an Indian Cuckoo as well as a lovely Chestnut-winged Cuckoo that flew across and perched very briefly. Another nice find along the river was a pair of very vocal and musical Bornean Black Magpies, found in the same tree two days running. With their bell-like honks and fluty whistles they perched out giving some low clicking croaks before flying across the river to continue their odd natural symphony in the forest around us. One of the most trying but rewarding birds along the river were a pair of Chestnut-necklaced Partridges which we managed to lure to the edge of one of the only unflooded banks down a narrow creek where we all managed to get a view of them.

The final stop on our Borneo tour is the jewel in the crown, Danum Valley. We had three full days here in the bird rich patch of lowland forest bookended by an afternoon and evening so we wasted no time after our arrival to get stuck in to the closest forest trail. Though slow in the early afternoon we eventually began to pick up birds with the star-of-the-show a lovely Blue-headed Pitta that bounded across the track in front of us twice. With this nice endemic target in the bag we headed back for our first night drive and Thomas’s Flying Squirrel, Greater Mouse Deer, Sambar Deer, Sunda Colugo and the rather uncommon Long-tailed Porcupine were all seen well.

The following three days was spent walking the fantastic trail system here and great birds were numerous. Malaysian and Moustached Hawk-Cuckoo’s were found with the Moustached being particularly vocal, there were additional Cuckoo’s with Indian, Violet and Little Bronze also seen. Raptors included Jerdon’s Baza, Oriental Honeybuzzard, a dusk flying Bat Hawk, Rufous-bellied Eagle, Wallace’s Hawk-Eagle and Brahminy Kite. Down by the river at lunch one day we watched a Grey-headed Fish Eagle perched atop a large boulder eating a fish. Trogon’s were abundant and we picked up multiple views of Scarlet-rumped, Diard’s and Red-naped as well as a lone Cinnamon-rumped next to the road one morning allowing some brilliant views of this rather uncommon and enigmatic trogon. Just after such fine views of the trogon we then encountered a noisy

group of Crested Jays (now known to be a basal shrike, with the new name Jay Shrike recently coined for it) for some long anticipated views after hearing several but not getting connected with them previously, it was nice to get some good, long, solid looks as they vaulted round the understory close at hand.

Large Frogmouth and Whitehead’s Trogon © Rob Fray

Rhinoceros Hornbill were abundant here with fifteen seen in one huge group one morning but the standout here was a pair of Helmeted Hornbills feeding in a fruiting tree, the species was recently uplisted to Critically Endangered due to the massive hunting pressure, primarily in Indonesia for the ivory trade as it’s the only species of hornbill with an ivory-filled casque. There are some really nice woodpecker’s here and we saw Rufous, Grey-and-Buff, Buff-rumped, White-bellied, Banded and a lovely pair of Orange-backed foraging close to the trail.

Flycatchers are always fun to track down and we had no problem with Malaysian, Bornean and Large-billed Blue here seen on several occasions. We also managed the more migrant Mugimaki and Narcissus as well as the drabber Grey-streaked Flycatcher. A deep forest trail gave us some brilliant looks at a close Rufous-chested Flycatcher perched above us singing away as well as a preening Grey-chested Jungle Flycatcher that had been dipping into a small creek to wash.

Some noisy Dusky Broadbills showed up one morning, soon joined by a pair of Banded Broadbills allowing us to clean up the Broadbills. The fiesta of spatulate bills had us craning canopywards but only the Banded Broadbills allowed us scope views as the spritely group of five Dusky Broadbills continued to move vigorously through the trees above us never settled in one place for more than a few seconds. High up with these were a Bornean Blue Flycatcher and a pair of Straw-headed Bulbul’s that let us know their position from their amazingly intricate song. Both Rufous-winged and Maroon-breasted Philentomas were picked up with Fiery Minivets adding a splash of color and Lesser Cicadabird calling regularly from the canopy. There is always a

slew of bulbuls here and though the aforementioned Straw-headed Bulbuls were the stars we found Grey-bellied, Cream-vented, Red-eyed, Spectacled, Grey-cheeked, Yellow-bellied and Buff-vented.

Another group that is always well represented here are the babblers and though split between a few genera we rounded up a big collection with the four Malacapteron babblers being the most numerous with Rufous-crowned, Scaly-crowned, Sooty-capped and Moustached all seen well. Bold-striped and Fluffy-backed Tit-Babblers were numerous as well as Chestnut-winged and on several occasions we found noisy groups of Chestnut-rumped Babblers, once near the lodge in an open area where we could see the blue sacs so many of these babblers possess, inflated during vocalizations. Also noted were a pair of Grey-headed, Ferruginous, Short-tailed and Black-capped Babblers. Horsfield’s Wren-Babbler and the incomparable Striped Wren-Babbler also showed well and the two endemic babblers that are usually a highlight here didn’t disappoint with first an alluring Bornean Ground Babbler uttering its two note whistle giving it away and its variegated white on black striping along the throat and breast made for a striking scene. Lastly a trio of Black-throated Wren-Babblers right next to the trail showed particularly well. First picked up calling below us, they moved up towards the trail with the level of noise increasing in volume as three birds began to play off each other duetting and bounding around in the understory. Back and forth across the trail the three of them moved often right over our heads for a fantastic and cacophonous display by this rather bulky babbler.

The canopy walkway can always turn up some goodies and though half was closed off for repairs the main section with the tallest platform was open and it was here we spent some time watching for high canopy denizens like Pale-blue Flycatcher, Yellow-eared and Spectacled Spiderhunter; the latter chasing the more diminutive Yellow-eared from a small pool of water that gathers in a depression on the trunk of a huge Koompassia excelsa. Velvet-fronted Nuthatch, Bornean Brown Barbet and Greater and Lesser Green Leafbirds were all noted and in the lee of a broken trunk Greg found a Red Giant Flying Squirrel poking its head out. It ducked back down after a short period and curled around its long bushy tail.

As if the daytime birds weren’t enough, besides the wonderful night mammals we also encountered some night birds with a very obliging Blyth’s Frogmouth that perched for some time allowing us a long period of study of this bantam frogmouth.

All good things must come to an end and after our final morning walk we headed back to catch our flights to Kota Kinabalu, passing a rather unobstructed Mount Kinabalu, for our farewell meal to celebrate a most successful tour.

Systematic List

GALLIFORMES: Phasianidae

Great Argus Argusianus argus heard only

Red-breasted Partridge Arborophila hyperythra e

Chestnut-necklaced Partridge Tropicoperdix charltonii graydoni

Crimson-headed Partridge Haematortyx sanguiniceps e

Bornean [Crested] Fireback Lophura ignita nobilis e

COLUMBIFORMES: Columbidae

Spotted Dove Streptopelia chinensis

Little Cuckoo Dove Macropygia ruficeps nana

Asian Emerald Dove Chalcophaps indica indica

Zebra Dove Geopelia striata I

Little Green Pigeon Treron olax

Pink-necked Green Pigeon Treron vernans

Jambu Fruit Dove Ptilinopus jambu

Green Imperial Pigeon Ducula aenea

Mountain Imperial Pigeon Ducula badia

CUCULIFORMES: Cuculidae

Chestnut-winged Cuckoo Clamator coromandus

Bock's Hawk Cuckoo Hierococcyx [sparverioides] bocki

Moustached Hawk Cuckoo Hierococcyx vagans

Malaysian Hawk Cuckoo Hierococcyx fugax

Sunda Cuckoo Cuculus lepidus

Indian Cuckoo Cuculus micropterus

Banded Bay Cuckoo Cacomantis sonneratii

Plaintive Cuckoo Cacomantis merulinus

Little Bronze Cuckoo Chrysococcyx minutillus aheneus

Violet Cuckoo Chrysococcyx xanthorhynchus

Asian Drongo Cuckoo Surniculus lugubris brachyurus

Raffles's Malkoha Rhinortha chlorophaeus

Black-bellied Malkoha Phaenicophaeus diardi

Red-billed Malkoha Phaenicophaeus javanicus

Chestnut-breasted Malkoha Phaenicophaeus curvirostris

Bornean Ground Cuckoo Carpococcyx radiatus e heard only

Short-toed Coucal Centropus rectunguis heard only

Greater Coucal Centropus sinensis bubutus

Lesser Coucal Centropus bengalensis javanensis heard only

APODIFORMES: Hemiprocnidae

Grey-rumped Treeswift Hemiprocne longipennis harterti

Whiskered Treeswift Hemiprocne comata comata

APODIFORMES: Apodidae

Giant Swiftlet Hydrochous gigas

Glossy Swiftlet Collocalia esculenta cyanoptila

Bornean Swiftlet Collocalia dodgei e

Mossy-nest Swiftlet Aerodramus salangana natunae

Black-nest Swiftlet Aerodramus maximus lowi

Edible-nest Swiftlet Aerodramus fuciphagus vestita

Silver-rumped Spinetail Rhaphidura leucopygialis

Brown-backed Needletail Hirundapus giganteus

Asian Palm Swift Cypsiurus balasiensis

House Swift Apus [affinis] nipalensis

CAPRIMULGIFORMES: Podargidae

Large Frogmouth Batrachostomus auritus

Dulit Frogmouth Batrachostomus harterti e

Bornean Frogmouth Batrachostomus mixtus e heard only

Blyth's Frogmouth Batrachostomus affinis affinis

CAPRIMULGIFORMES: Caprimulgidae

Large-tailed Nightjar Caprimulgus macrurus salvadorii

Grey Nightjar Caprimulgus jotaka heard only

Malaysian Eared Nightjar Lyncornis temminckii

GRUIFORMES: Rallidae

White-breasted Waterhen Amaurornis phoenicurus

Black-backed Swamphen Porphyrio indicus

Common Moorhen Gallinula chloropus

CHARADRIIFORMES: Scolopacidae

Common Sandpiper Actitis hypoleucos

PELECANIFORMES: Anhingidae

Oriental Darter Anhinga melanogaster

CICONIIFORMES: Ciconiidae

Storm's Stork Ciconia stormi

Lesser Adjutant Leptoptilos javanicus

CICONIIFORMES: Ardeidae

Purple Heron Ardea purpurea

Eastern Great Egret Ardea modesta

Intermediate Egret Ardea intermedia intermedia

Eastern Cattle Egret Ardea coromandus

Little Egret Egretta garzetta

Striated Heron Butorides striata

Black-crowned Night Heron Nycticorax nycticorax

Yellow Bittern Ixobrychus sinensis

ACCIPITRIFORMES: Accipitridae

Jerdon's Baza Aviceda jerdoni

Oriental Honeybuzzard Pernis ptilorhynchus torquatus

Bat Hawk Macheiramphus alcinus alcinus

Brahminy Kite Haliastur indus

White-bellied Fish Eagle Ichthyophaga leucogaster

Lesser Fish Eagle Ichthyophaga humilis

Grey-headed Fish Eagle Ichthyophaga ichthyaetus

Mountain Serpent Eagle Spilornis kinabaluensis e

Crested Serpent Eagle Spilornis cheela

Crested Goshawk Lophospiza trivirgatus

Black Eagle Ictinaetus malayensis

Rufous-bellied Eagle Lophotriorchis kienerii

Changeable Hawk Eagle Nisaetus limnaetus

Blyth's Hawk Eagle Nisaetus alboniger

Wallace's Hawk Eagle Nisaetus nanus

STRIGIFORMES: Strigidae

Mountain Scops Owl Otus spilocephalus luciae

Brown Wood Owl Strix leptogrammica vaga

'Sunda' Collared Owlet Glaucidium [brodiei] borneense heard only

TROGONIFORMES: Trogonidae

Red-naped Trogon Harpactes kasumba impavidus

Diard's Trogon Harpactes diardii diardii

Whitehead's Trogon Harpactes whiteheadi e

Cinnamon-rumped Trogon Harpactes orrhophaeus vidua

Scarlet-rumped Trogon Harpactes duvaucelii

Orange-breasted Trogon Harpactes oreskios dulitensis heard only

CORACIIFORMES: Bucerotidae

Oriental Pied Hornbill Anthracoceros albirostris convexus

Black Hornbill Anthracoceros malayanus

Rhinoceros Hornbill Buceros rhinoceros borneoensis

Helmeted Hornbill Rhinoplax vigil

Bushy-crested Hornbill Anorrhinus galeritus

White-crowned Hornbill Berenicornis comatus

Wrinkled Hornbill Aceros corrugatus

Wreathed Hornbill Rhyticeros undulatus

PICIFORMES: Capitonidae

Gold-whiskered Barbet Psilopogon chrysopogon chrysopsis

Red-throated Barbet Psilopogon mystacophanos heard only

Mountain Barbet Psilopogon monticola e

Yellow-crowned Barbet Psilopogon henricii

Golden-naped Barbet Psilopogon pulcherrima e

Blue-eared Barbet Psilopogon australis duvaucelii

Bornean Barbet Psilopogon eximia e

'Bornean' Brown Barbet Calorhamphus [fuliginosus] tertius

PICIFORMES: Picidae

Rufous Piculet Sasia abnormis

Sunda Woodpecker Dendrocopos moluccensis

Rufous Woodpecker Celeus brachyurus badiosus

White-bellied Woodpecker Dryocopus javensis javensis

Banded Yellownape Chrysophlegma mineaceus malaccensis

Checker-throated Yellownape Chrysophlegma mentalis humii

Maroon Woodpecker Blythipicus rubiginosus

Orange-backed Woodpecker Reinwardtipicus validus xanthopygius

Buff-rumped Woodpecker Meiglyptes tristis grammithorax

Buff-necked Woodpecker Meiglyptes tukki tukki

Grey-and-buff Woodpecker Hemicircus concretus sordidus

CORACIIFORMES: Alcedinidae

Common Kingfisher Alcedo atthis bengalensis

Blue-eared Kingfisher Alcedo meninting verreauxii

Blue-banded Kingfisher Alcedo euryzona peninsulae

Rufous-backed Kingfisher Ceyx rufidorsa motleyi

Banded Kingfisher Lacedo pulchella melanops heard only

Stork-billed Kingfisher Pelargopsis capensis inominata

Ruddy Kingfisher Halcyon coromanda minor

Collared Kingfisher Todirhamphus chloris laubmannianus

Rufous-collared Kingfisher Actenoides concretus borneanus

CORACIIFORMES: Meropidae

Red-bearded Bee-eater Nyctyornis amictus

Blue-throated Bee-eater Merops viridis viridis

CORACIIFORMES: Coraciidae

Common Dollarbird Eurystomus orientalis

FALCONIFORMES: Falconidae

White-fronted Falconet Microhierax latifrons e

PSITTACIFORMES: Psittacidae

Long-tailed Parakeet Psittacula longicauda

Blue-crowned Hanging Parrot Loriculus galgulus

PASSERIFORMES: Calyptomenidae

Green Broadbill Calyptomena viridis heard only

Hose's Broadbill Calyptomena hosii e

Whitehead's Broadbill Calyptomena whiteheadi e

PASSERIFORMES: Eurylaimidae

Dusky Broadbill Corydon sumatranus

Black-and-red Broadbill Cymbirhynchus macrorhynchos

Banded Broadbill Eurylaimus javanicus

Black-and-yellow Broadbill Eurylaimus ochromalus

Long-tailed Broadbill Psarisomus dalhousiae

PASSERIFORMES: Pittidae

Bornean Banded Pitta Hydrornis schwaneri e

Blue-headed Pitta Hydrornis baudii e

Hooded Pitta Erythropitta sordida mulleri

Blue-banded Pitta Erythropitta arquata e

Black-crowned Pitta Erythropitta ussheri e

PASSERIFORMES: Pardalotidae

Golden-bellied Gerygone Gerygone sulphurea sulphurea

PASSERIFORMES: Vireonidae

Blyth's Shrike Babbler Pteruthius aeralatus robinsoni

Erpornis Erpornis zantholeuca

PASSERIFORMES: Oriolidae

Dark-throated Oriole Oriolus xanthonotus consobrinus

Black Oriole Oriolus hosii e

Black-and-crimson Oriole Oriolus cruentus vulneratus

PASSERIFORMES: Pachycephalidae

Bornean Whistler Pachycephala hypoxantha e

PASSERIFORMES: Vangidae

Bar-winged Flycatcher-shrike Hemipus picatus intermedius

Black-winged Flycatcher-shrike Hemipus hirundinaceus

Rufous-winged Philentoma Philentoma pyrhopterum

Maroon-breasted Philentoma Philentoma velatum

PASSERIFORMES: Pityriaseidae

Bristlehead Pityriasis gymnocephala e

PASSERIFORMES: Aegithinidae

Common Iora Aegithina tiphia aequanimis

Green Iora Aegithina viridissima

PASSERIFORMES: Artamidae

White-breasted Woodswallow Artamus leucorynchus

PASSERIFORMES: Campephagidae

Sunda Cuckooshrike Coracina larvata normani

Lesser Cicadabird Lalage fimbriata schierbrandii

Pied Triller Lalage nigra nigra

Fiery Minivet Pericrocotus igneus igneus

Grey-chinned Minivet Pericrocotus solaris cinereigula

PASSERIFORMES: Rhipiduridae

White-throated Fantail Rhipidura albicollis kinabalu

Pied Fantail Rhipidura javanica

Spotted Fantail Rhipidura perlata

PASSERIFORMES: Dicruridae

Sunda [Ashy] Drongo Dicrurus leucophaeus stigmatops

Hair-crested Drongo Dicrurus [hottentottus] borneensis

Greater Racket-tailed Drongo Dicrurus paradiseus brachyphorus

PASSERIFORMES: Monarchidae

Black-naped Monarch Hypothymis azurea prophata

Asian Paradise Flycatcher Terpsiphone paradisi borneensis

PASSERIFORMES: Laniidae

Crested Jay Platylophus galericulatus coronatus

PASSERIFORMES: Corvidae

Bornean Black Magpie Platysmurus aterrimus e

Bornean Green Magpie Cissa jefferyi e

Bornean Treepie Dendrocitta cinerascens e

Slender-billed Crow Corvus enca compilator

PASSERIFORMES: Stenostiridae

Grey-headed Canary-flycatcher Culicicapa ceylonensis

PASSERIFORMES: Hirundinidae

Barn Swallow Hirundo rustica gutturalis

Pacific Swallow Hirundo tahitica

PASSERIFORMES: Pycnonotidae

Straw-headed Bulbul Pycnonotus zeylanicus

Black-headed Bulbul Pycnonotus atriceps atriceps

Bornean Bulbul Pycnonotus montis e

Grey-bellied Bulbul Pycnonotus cyaniventris

Yellow-vented Bulbul Pycnonotus goiavier gourdini

Cream-vented Bulbul Pycnonotus simplex perplexus

Red-eyed Bulbul Pycnonotus brunneus brunneus

Spectacled Bulbul Pycnonotus erythropthalmos

Ochraceous Bulbul Alophoixus ochraceus ruficrissus

Grey-cheeked Bulbul Alophoixus bres gutturalis

Yellow-bellied Bulbul Alophoixus phaeocephalus

Hairy-backed Bulbul Tricholestes criniger

Buff-vented Bulbul Iole olivacea charlottae

Streaked Bulbul Ixos malaccensis

Cinereous Bulbul Hemixos cinereus connectens

Hook-billed Bulbul Setornis criniger

PASSERIFORMES: Timaliidae

Bold-striped Tit Babbler Mixornis bornensis

Fluffy-backed Tit Babbler Macronous ptilosus reclusus

Chestnut-winged Babbler Cyanoderma erythropterum bicolor

Rufous-fronted Babbler Cyanoderma rufifrons sarawacensis

Chestnut-backed Scimitar Babbler Pomatorhinus montanus bornensis

Grey-throated Babbler Stachyris nigriceps borneensis

Grey-headed Babbler Stachyris poliocephala

Black-throated Babbler Stachyris nigricollis

Chestnut-rumped Babbler Stachyris maculata maculata

PASSERIFORMES: Pellorneidae

Moustached Babbler Malacopteron magnirostre cinereocapilla

Sooty-capped Babbler Malacopteron affine phoeniceum

Scaly-crowned Babbler Malacopteron cinereum cinereum

Rufous-crowned Babbler Malacopteron magnum saba

White-chested Babbler Pellorneum rostratum macropterum

Ferruginous Babbler Pellorneum bicolor

Short-tailed Babbler Pellorneum malaccense sordidum

Temminck's Babbler Pellorneum pyrrogenys canicapillus

Black-capped Babbler Pellorneum capistratum morrelli

Striped Wren Babbler Kenopia striata

Horsfield's Wren Babbler Turdinus sepiarius harterti

Black-throated Wren Babbler Turdinus atrigularis e

Mountain Wren Babbler Turdinus crassus e

Bornean Ground Babbler Ptilocichla leucogrammica e

PASSERIFORMES: Leiothrichidae

Brown Fulvetta Alcippe brunneicauda

Sunda Laughingthrush Garrulax palliatus

Bare-headed Laughingthrush Garrulax calvus e

Chestnut-hooded Laughingthrush Ianthocincla treacheri treacheri e

PASSERIFORMES: Zosteropidae

Chestnut-crested Yuhina Yuhina everetti e

Pygmy Ibon Heleia squamifrons e

Black-capped White-eye Zosterops atricapillus

Mountain Black-eye Zosterops emiliae e

PASSERIFORMES: Phylloscopidae

Arctic Warbler Seicercus borealis

Mountain Leaf Warbler Seicercus trivirgatus kinabaluensis

Yellow-breasted Warbler Seicercus montis floris

PASSERIFORMES: Cettidae

Yellow-bellied Warbler Abroscopus superciliaris schwaneri

Bornean Stubtail Urosphena whiteheadi e

Aberrant (Sunda) Bush Warbler Horornis flavolivaceus oreophila

Mountain Tailorbird Phyllergates cuculatus cinereicollis

PASSERIFORMES: Locustellidae

Striated Grassbird Megalurus palustris

PASSERIFORMES: Cisticolidae

Dark-necked Tailorbird Orthotomus atrogularis atrogularis

Rufous-tailed Tailorbird Orthotomus sericeus sericeus

Ashy Tailorbird Orthotomus ruficeps borneoensis

Yellow-bellied Prinia Prinia flaviventris latrunculus

PASSERIFORMES: Sittidae

Velvet-fronted Nuthatch Sitta frontalis corallipes

PASSERIFORMES: Sturnidae

Asian Glossy Starling Aplonis panayensis

Common Hill Myna Gracula religiosa

Crested Myna Acridotheres cristatellus I

Javan Myna Acridotheres javanicus I

PASSERIFORMES: Turdidae

Orange-headed Thrush Geokichla citrina aurata

Everett's Thrush Zoothera everetti e

Eye-browed Thrush Turdus obscurus

Fruit-hunter Chlamydochaera jefferyi e

PASSERIFORMES: Muscicapidae

Oriental Magpie Robin Copsychus saularis adamsi

White-crowned Shama Copsychus stricklandii e

White-rumped Shama Copsychus malabaricus

Rufous-tailed Shama Copsychus pyrropyga

Grey-streaked Flycatcher Muscicapa griseisticta

Dark-sided Flycatcher Muscicapa sibirica

Asian Brown Flycatcher Muscicapa dauurica dauurica

Grey-chested Jungle Flycatcher Cyornis umbratilis

Pale Blue Flycatcher Cyornis unicolor herterti

Hill Blue Flycatcher Cyornis [banyumas] coeruleatus

Large-billed Blue Flycatcher Cyornis caerulatus caerulatus

Malaysian Blue Flycatcher Cyornis turcosus

Bornean Blue Flycatcher Cyornis superbus e

Blue-and-white Flycatcher Cyanoptila cyanomelana

Verditer Flycatcher Eumyias thalassina

Indigo Flycatcher Eumyias indigo

Eye-browed Jungle Flycatcher Vauriella gularis e

White-browed Shortwing Brachypteryx montana erythrogyna

Chestnut-naped Forktail Enicurus ruficapillus

White-crowned Forktail Enicurus leschenaulti frontalis

Bornean Forktail Enicurus borneensis e

Bornean Whistling Thrush Myophonus borneensis e

Narcissus Flycatcher Ficedula narcissina narcissina

Mugimaki Flycatcher Ficedula mugimaki

Snowy-browed Flycatcher Ficedula hyperythra sumatrana

Rufous-chested Flycatcher Ficedula dumetoria

Little Pied Flycatcher Ficedula westermanni

PASSERIFORMES: Chloropseidae

Greater Green Leafbird Chloropsis sonnerati zosterops

Lesser Green Leafbird Chloropsis cyanopogon cyanopogon

Bornean Leafbird Chloropsis kinabaluensis e

PASSERIFORMES: Irenidae

Asian Fairy Bluebird Irena puella crinigera

PASSERIFORMES: Dicaeidae

Yellow-breasted Flowerpecker Prionochilus maculatus maculatus

Yellow-rumped Flowerpecker Prionochilus xanthopygius e

Scarlet-breasted Flowerpecker Prionochilus thoracicus

Orange-bellied Flowerpecker Dicaeum trigonostigma dayakanum

Black-sided Flowerpecker Dicaeum monticolum e

Scarlet-backed Flowerpecker Dicaeum cruentatum nigrimentum

PASSERIFORMES: Nectariniidae

Ruby-cheeked Sunbird Chalcoparia singalensis

Plain Sunbird Anthreptes simplex

Brown-throated Sunbird Anthreptes malacensis

Red-throated Sunbird Anthreptes rhodolaema heard only

Olive-backed Sunbird Cinnyris jugularis

Copper-throated Sunbird Leptocoma calcostetha

Van Hasselt's Sunbird Leptocoma brasiliana

Crimson Sunbird Aethopyga siparaja

Temminck's Sunbird Aethopyga temminckii

Purple-naped Spiderhunter Arachnothera hypogrammicum

Thick-billed Spiderhunter Arachnothera crassirostris

Spectacled Spiderhunter Arachnothera flavigaster

Long-billed Spiderhunter Arachnothera robusta

Little Spiderhunter Arachnothera longirostra

Yellow-eared Spiderhunter Arachnothera chrysogenys

Bornean Spiderhunter Arachnothera everetti e

Whitehead's Spiderhunter Arachnothera juliae e

PASSERIFORMES: Passeridae

Eurasian Tree Sparrow Passer montanus malaccensis

PASSERIFORMES: Estrildidae

Dusky Munia Lonchura fuscans e

Scaly-breasted Munia Lonchura punctulata cabanisi

Black-headed Munia Lonchura atricapilla jagori

PASSERIFORMES: Motacillidae

Grey Wagtail Motacilla cinerea cinerea

Paddyfield Pipit Anthus rufulus

MAMMALS

Mountain Treeshrew Tupaia montana baluensis e

Large Flying Fox Pteropus vampyrus

Sunda Colugo Cynocephalus variegatus

Maroon Leaf Monkey Presbytis rubicunda e

Sunda Silvered Leaf Monkey Presbytis cristata

Proboscis Monkey Nasalis larvatus e

Long-tailed Macaque Macaca fascicularis

Sunda Pig-tailed Macaque Macca nemestrina

Bornean Gibbon Hylobates muelleri e heard only

Bornean Orang-utan Pongo pygmaeus e

'Bornean' Giant Squirrel Ratufa affinis

Prevost's Squirrel Callosciurus prevostii pluto

Kinabalu Squirrel Callosciurus baluensis e

Plantain Squirrel Callosciurus notatus

Ear-spot Squirrel Callosciurus adamsi

Bornean Black-banded Squirrel Callosciurus orestes e

Low's Squirrel Sundasciurus lowii

Jentink's Squirrel Sundasciurus jentinki e

Brooke's Squirrel Sundasciurus brookei

Plain Pygmy Squirrel Exilisciurus exilis e

Whitehead's Pygmy Squirrel Exilisciurus whiteheadi e

Thomas's Flying Squirrel Aeromys thomasi e

Red Giant Flying Squirrel Petaurista petaurista

Long-tailed Porcupine Trichys fasciculata

Malay Civet Viverra tangalunga

Small-toothed Palm Civet Arctogalidia trivirgata

Bearded Pig Sus barbatus

Greater Mouse-Deer Tragulus napu

Sambar Deer Cervus unicolor