signal coding definitions types of coding schemes inferring sender and receiver coding signal...

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Signal coding • Definitions • Types of coding schemes • Inferring sender and receiver coding • Signal function and coding

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Page 1: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Signal coding

• Definitions

• Types of coding schemes

• Inferring sender and receiver coding

• Signal function and coding

Page 2: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Conflict resolution example

Page 3: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Signal definitions

Page 4: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Assumptions

• Senders produce signals in order to provide honest (accurate) information to receivers– Dishonest signalling will be considered in the

third section of the course

• Communication involves signal production, transmission, and reception. All three processes influence the accuracy of any coding scheme

Page 5: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Coding matrix

Probability of giving a signal in each condition. Sender matrixmust be similar to Receiver matrix for communication to occur.

Page 6: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Coding conventions

• None - all probabilities equal

• Perfect - each signal coocurs with each condition

• Imperfect– Specific - one signal per condition, but

conditions may not be discriminable– Unique - one or more signals per condition

Page 7: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Reasons for imperfect coding

• Sender– Errs in assessment of condition– Mistakenly assigns signals to conditions

• Receiver– Signal is modified by transmission– Errs in identifying signal– Has not yet learned correct signal

Page 8: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Coding schemes

• Codes require signal diversity

• Variation can be created by – Modifying signal element attributes

• Sound: amplitude, frequency, duration

• Light: color, size, location

– Combining signal elements in series

• Signal elements must be perceptually distinct

Page 9: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Hierarchical syntax

Page 10: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Stereotypy

Stereotypic signals may represent constraints on senders

Page 11: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Coding options

Page 12: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Coding scheme examples

Page 13: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Iconic aggressive signals

Page 14: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Inferring sender coding schemes

• For discrete conditions and discrete signals, use contingency table analysis

• For discrete conditions and continuous signals, use discriminant function analysis

• For continuous signals and uncertain conditions, use clustering or principle component analysis

Page 15: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Phyllostomus hastatus

Page 16: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Females form stable groups

Page 17: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Departing bats give screech calls

Bouts of screech calling often coincide with group departures

Page 18: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Calling rate and diet varies with season

Page 19: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Calling females recruit group mates

Page 20: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Groups have distinct calls

Page 21: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Inferring receiver coding schemes

• Determine how receiver categorizes the set of signal variants– In captivity, present alternative forms to

determine which are perceived as same or different using habituation-dishabituation expt or operant conditioning paradigm

• Determine which condition is associated with each category by the receiver

Page 22: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Females can discriminate group mates by call

Page 23: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Compound coding schemes

• Combination mapping– Combining information into a single signal can save

energy and reduce exposure

• Parameter mapping– Body size covaries with pitch, energy reserves

covaries with calling rate

• Hierarchical mapping– Mean differences denote group, individual

differences denote individual

Page 24: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Tamarin group calls

Page 25: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Signal ontogeny

• Heritable• Favored when coding scheme is predictable, e.g.

individual or kinship differences

• Learned– Individual or trial-and-error– Social

• Critical period vs. open-ended

• Believed to be faster and avoids costly errors

Page 26: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Females learn to match calls

Group 1 Group 2

Before

5 monthsafter move

Page 27: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Signal function and coding

• Binary assignment– E.g. sex labels– need only 2 signals, can be heritable

• Binary recognition– need many signals– decide own vs other

Page 28: Signal coding Definitions Types of coding schemes Inferring sender and receiver coding Signal function and coding

Signal function and coding

• Binary comparison - often continuous signals– opponent fighting ability, threshold mate

choice, best-of-n mate choice

• Manifold decisions– Iconic rule - Honeybee language– pairwise associations