the moss flora of the wallowa mountains of oregon

The Moss Flora of the Wallowa Mountains of OregonAuthor(s): Robert H. DuvallSource: The Bryologist, Vol. 41, No. 4 (Aug., 1938), pp. 90-95Published by: American Bryological and Lichenological SocietyStable URL: http://www.jstor.org/stable/3239341 .

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THE BRYOLOGIST THE BRYOLOGIST

FISSIDENS VIRIDULUS var. TAMARINDIFOLIUS (Brid.) Grout. Same habitat as the last, and similar except for the curved capsule. No. PIEDMONT: Durham Co., Blomquist 7383.

(To be continued)

THE MOSS FLORA OF THE WALLOWA MOUNTAINS OF OREGON

ROBERT H. DUVALL

The Wallowa Mountains are an excellent collecting ground for mosses since they contain bogs, mountain streams, rocky cliffs, mountain meadows and wooded areas, all of which are typical habi- tats for certain moss forms. There is also a great altitude range, so that one finds lowland and alpine mosses all within a few hours climb into the mountains. Although a relatively small area was covered, it

was amazing what a wide variety of forms was represented. The area of these mountains as considered in this paper is about 2050 square miles, and is located about Latitude 44? 40' North, Longitude 117?

20' West. The altitude range is 3000-6000 feet.

Topographically the region is quite variable. The highest points are regions of perpetual snow. To the east the Snake River has

ground its canyon through to a depth of over a mile. The lower

stretches grade into the dry interior plateau. We have no record of previous collecting of bryophytes in this

region, although we find a similarity in some of the species to the

report of George N. Jones, "Moss Flora of Southeastern Washington" Volume I, Research Studies of the State College of Washington, Pullman, 1929.

The present report is in no way a complete list of the mosses of the Wallowa Region, but it is hoped that it may be an aid to future work on the moss flora of the region.

Certain difficult species were determined by Dr. T. C. Frye, Mr.

George N. Jones of the University of Washington, and Dr. Winona

Welch of DePauw University. SPHAGNACEAE

SPHAGNUM CAPILLACEUM (Weiss.) Schrank. Cornucopia, Snyder Meadow.

SPHAGNUM GIRGENSOHNII Russ. Wallowa River Region, Ice Lake.

TETRAPHIDACEAE

TETRAPHIS PELLUCIDA (L.) Schimp. Wallowa River Region.

FISSIDENS VIRIDULUS var. TAMARINDIFOLIUS (Brid.) Grout. Same habitat as the last, and similar except for the curved capsule. No. PIEDMONT: Durham Co., Blomquist 7383.

(To be continued)

THE MOSS FLORA OF THE WALLOWA MOUNTAINS OF OREGON

ROBERT H. DUVALL

The Wallowa Mountains are an excellent collecting ground for mosses since they contain bogs, mountain streams, rocky cliffs, mountain meadows and wooded areas, all of which are typical habi- tats for certain moss forms. There is also a great altitude range, so that one finds lowland and alpine mosses all within a few hours climb into the mountains. Although a relatively small area was covered, it

was amazing what a wide variety of forms was represented. The area of these mountains as considered in this paper is about 2050 square miles, and is located about Latitude 44? 40' North, Longitude 117?

20' West. The altitude range is 3000-6000 feet.

Topographically the region is quite variable. The highest points are regions of perpetual snow. To the east the Snake River has

ground its canyon through to a depth of over a mile. The lower

stretches grade into the dry interior plateau. We have no record of previous collecting of bryophytes in this

region, although we find a similarity in some of the species to the

report of George N. Jones, "Moss Flora of Southeastern Washington" Volume I, Research Studies of the State College of Washington, Pullman, 1929.

The present report is in no way a complete list of the mosses of the Wallowa Region, but it is hoped that it may be an aid to future work on the moss flora of the region.

Certain difficult species were determined by Dr. T. C. Frye, Mr.

George N. Jones of the University of Washington, and Dr. Winona

Welch of DePauw University. SPHAGNACEAE

SPHAGNUM CAPILLACEUM (Weiss.) Schrank. Cornucopia, Snyder Meadow.

SPHAGNUM GIRGENSOHNII Russ. Wallowa River Region, Ice Lake.

TETRAPHIDACEAE

TETRAPHIS PELLUCIDA (L.) Schimp. Wallowa River Region.

90 90 [AUGUST [AUGUST



1938] DUVALL: MOSSES OF WALLOWA MOUNTAINS

POLYTHICHACEAE

POLYTRICHADELPHUS LYELLII Mitt. Lostine, Cornucopia, Alder

Springs, Fish Lake, Big Sheep Creek, Mud Springs, Wallowa River Region.

POLYTRICHUM COMMUNE L. Wallowa River Region, Cornucopia. POLYTRICHUM JENSENII Hag. Aneroid Lake Trail, Ice Lake. POLYTRICHUM JUNIPERINUM Willd. Wallowa River Region, Alder

Springs, Fish Lake, Cornucopia, Lostine, Wallowa Lake, Big Sheep Creek, Mud Springs, Aneroid Lake, Joseph.

POLYTRICHUM PILIFERUM Schreb. Wallowa River Region, Cornu-

copia, Big Sheep Creek, Aneroid Lake, Ice Lake. POGONATUM ALPINUM (L.) Roelil. Wallowa River Region, Alder

Springs, Ice Lake Trail. DICRANACEAE

CERATODON PURPUREUS (L.) Sch. Wallowa River Region. DICRANOWEISIA CIRRATA Timm. Alder Springs, Big Sheep Creek,

Mud Springs. DICRANUM STRICTUM Schleich. Alder Springs, Elk Creek, Cornu-

copia. DITRICHUM MONTANUM Leib. Cornucopia, Elk Creek, Snyder

Meadow, Lostine. DITRICHUM TENUIFOLIUM Lindb. Big Sheep Creek. LEPTODICTYON VACILLANS (Sull.) Broth. Cornucopia.

GRIMMIACEAE

BRAUNIA CALIFORNICA Sull. Aneroid Lake, Wallowa River Region. GRIMMIA ALPESTRIS Nees. Cornucopia, Elk Creek, Big Sheep

Creek, Alder Springs, Wallowa River Region. GRIMMIA ALPICOLA Hedw. var. RIVULARIS (Brid.) Broth. Wallowa

River Region, Aneroid Lake, Ice Lake. GRIMMIA APOCARPA (L.) Hedw. Alder Springs, Wallowa River

Region, Big Sheep Creek, Aneroid Lake, Lostine. GRIMMIA APOCARPA (L.) Hedw. var. GRACILIS (Schleich.) Web. &

Mohr. Aneroid Lake, Lostine, Wallowa River Region, Joseph. GRIMMIA COMMUTATA Huebn. Big Sheep Creek, Alder Springs. GRIMMIA MARITIMA Turn. Alder Springs, Big Sheep Creek.

GRIMMIA MONTANA Br. & Sch. Wallowa River Region, Lostine.

GRIMMIA MUHLENBECKII Husn. Alder Springs, Big Sheep Creek,

Joseph. GRIMMIA RAUI Aust. Wallowa River Region. HEDWIGIA CILIATA (Ehrh.) Hedw. Big Sheep Creek, Wallowa

River Region, Aneroid Lake Trail. SCOULERIA AQUATICA Hook. Big Sheep Creek, Aneroid Lake Trail,

Ice Lake Trail, Wallowa River Region.

TORTULACEAE

BARBULA REVOLUTA (Schrad.) Brid. Big Sheep Creek, Alder

Springs.

91



POTTIA TRUNCATA (Hedw.) Fuernr. Var. intermedia Turn. Cor- nucopia.

TORTELLA TORTUOSA (L.) Limpr. Cornucopia. TORTULA ACIPHYLLA (Br. & Sch.) Hartm. Wallowa River Region,

Alder Springs. TORTULA MUELLERI (Br.) Wils. Wallowa River Region, Aneroid

Lake. TORTULA NORVEGICA (Web.) Lindb. Wallowa River Region,

Alder Springs. TORTULA RURALIS (L.) Ehrh. Cornucopia, Lostine, Wallowa

River Region, Alder Springs, Big Sheep Creek.

ENCALYPTACEAE

ENCALYPTA VULGARIS Hedw. Alder Springs.

ORTHOTRICHACEAE

ORTHOTRICHUM LYELLII (Hook.) Taylor. Aneroid Lake Trail. Wallowa River Region, Alder Springs.

ORTHOTRICHUM PUCHELLUM Brunton. Wallowa River Region, Alder Springs.

ORTHOTRICHUM RIVULARE Turn. Wallowa River Region. ULOTA PHYLLANTHA Brid. Alder Springs.

TIMMIACEAE

TIMMIA AUSTRIACA Hedw. Big Sheep Creek, Wallowa River Region, Aneroid Lake Trail, Alder Springs.

AULACOMNIACEAE

AULACOMNIUM ANDROGYNUM Schwaegr. Cornucopia, Aneroid Lake, Lostine.

AULACOMNIUM PALUSTRE (Web. & Mohr.) Schwaegr. Aneroid Lake, Wallowa Lake, Cornucopia, Wallowa River Region, Alder Springs.

BARTRAMIACEAE

BARTRAMIA ITHYPHYLLA Brid. Wallowa River Region, Ice Lake. BARTRAMIA POMIFORMIS (L.) Hedw. Aneroid Lake Trail, Ice Lake,

Wallowa River Region. PHILONOTIS CAPILLARIS Lindb. Cornucopia, Alder Springs. PHILONOTIS FONTANA (L.) Brid. Wallowa River Region, Big Sheep

Creek, Cornucopia, Snyder Meadow, Elk Creek, Aneroid Lake Trail, Alder Springs.

PHILONOTIS FONTANA (L.) Brid. var. LAXA Flowers. Aneroid Lake, Cornucopia, Snyder Meadow.

PHILONOTIS FONTANA (L.) Brid. var. SERIATA Breidl. Big Sheep Creek.

BRYACEAE

BRYUM ARGENTEUM L. Alder Springs, Cornucopia, Lostine. BRYUM ATWATERIAE C. Muell. Wallowa River Region.

92 THE BRYOLOGIST [AUGUST



1938] DUVALL: MOSSES OF WALLOWA MOUNTAINS

BRYUM BIMUM Schreb. Aneroid Lake Trail, Cornucopia. BRYUM CAESPITICIUM L. Alder Springs, Cornucopia. BRYUM CIRRATUM Hoppe & Hornsch. Cornucopia, Snyder Meadow. BRYUM ERYTHROCARPUM Schwaegr. Alder Springs. BRYUM INCLINATUM Br. & Sch. Wallowa River Region, Aneroid

Lake Trail, Ice Lake. BRYUM INTERMEDIUM Brid. Aneroid Lake Trail, Ice Lake, Alder

Springs. BRYUM PERCURRENTINERVE Kindb. Alder Springs. LEPTOBRYUM PYRIFORME (L.) Wils. Big Sheep Creek, Alder

Springs, Lostine, Aneroid Lake Trail, Ice Lake, Cornucopia, Wallowa River Region.

POHLIA ACUMINATA Hoppe & Hornsch. Big Sheep Creek, Mud Springs.

POHLIA ANNOTINA (Hedw.) Loeske. Cornucopia, Snyder Meadow. POHLIA CRUDA (L.) Lindb. Cornucopia, Ice Lake, Aneroid Lake

Trail, Lostine, Alder Springs, Fish Lake, Big Sheep Creek. POHLIA NUTANS (Schreb.) Lindb. Aneroid Lake Trail, Alder

Springs, Cornucopia. POHLIA PROLIGERA Lindb. Big Sheep Creek, Alder Springs, Cornu-

copia. POHLIA PUCHELLA (Hedw.) Lindb. Cornucopia, Aneroid Lake, Ice

Lake. RHODOBRYUM ONTARIENSE (Kindb.) Paris. Big Sheep Creek.

MNIACEAE

MNIUM AFFINE Bland. Cornucopia, Snyder Meadow, Aneroid Lake Trail, Ice Lake.

MNIUM BLYTTII R. & C. Alder Springs, Aneroid Lake, Ice Lake, Big Sheep Creek.

MNIUM HORNUM L. Aneroid Lake Trail, Ice Lake. MNIUM MARGINATUM (Dicks.) Beaver. Alder Springs. MNIUM MEDIUM Br. & Sch. Cornucopia, Elk Creek. MNIUM NUDUM Will. Big Sheep Creek, Alder Springs, Cornucopia. MNIUM ORTHORRHYNCHUM (Brid.) Br. & Sch. Wallowa River

Region, Cornucopia. MNIUM PUNCTATUM (L.) Hedw. Wallowa River Region, Aneroid

Lake Trail, Ice Lake. ROELLIA LUCIDUM (E. G. Britt.) Kindb. Lostine, Alder Springs,

Cornucopia. HYPNACEAE

BESTIA BREWERIANA Lesq. Cornucopia, Alder Springs. BESTIA HOLZINGERI (R. & C.) Broth. Cornucopia, Snyder Meadow. BRACHYTHECIUM ACUTUM (Mitt.) Sulliv. Alder Springs, Aneroid

Lake, Ice Lake. BRACHYTHECIUM ALBICANS (Neck.) Br. & Sch. Alder Springs,

Wallowa River Region, Lostine.

93



BRACHYTHECIUM BOLANDERI (Lesq.) Jaeger & Sauerb. Cornu- copia.

BRACHYTHECIUM CAMPESTRE Br. & Sch. Alder Springs. BRACHYTHECIUM COLLINUM (Schleich) Br. & Sch. Cornucopia,

Alder Springs. BRACHYTHECIUM ERYTHRORRHIZON Br. & Sch. Big Sheep Creek,

Aneroid Lake Trail, Ice Lake, Alder Springs. BRACHYTHECIUM LAMPROCHYSEUM C. Muell. & Kindb. Big Sheep

Creek, Alder Springs, Lostine. BRACHYTHECIUM LEIBERGII Grout. Aneroid Lake, Wallowa River

Region, Alder Springs. BRACHYTHECIUM PLUMULOSUM (Sw.) Br. & Sch. Cornucopia. BRACHYTHECIUM REFLEXUM (Starke) Br. & Sch. Aneroid Lake

Trail. BRACHYTHECIUM RIVULARE Br. & Sch. Cornucopia, Alder Springs. BRACHYTHECIUM WASHINGTONIANUM (Eaton) Grout. Alder Springs,

Cornucopia, Wallowa River Region, Big Sheep Creek. CALLIERGON CORDIFOLIUM (Hedw.) Kindb. Cornucopia. CALLIERGON GIGANTEUM (Schimp.) Kindb. Alder Springs. CALLIERGONELLA SCHREBERI (Willd.) Grout. Alder Springs,

Aneroid Lake Trail, Ice Lake. CAMPTOTHECIUM AENEUM (Mitt.) Jaeger & Sauerb. Cornucopia. CAMPTOTHECIUM AMESIAE R. & C. Wallowa River Region, Alder

Springs, Aneroid Lake, Ice Lake. CAMPTOTHECIUM ARENARIUM (Lesq.) Jaeger & Sauerb. Ice Lake,

Big Sheep Creek, Wallowa River Region, Lostine. CAMPTOTHECIUM LUTESCENS (Huds.) Br. & Sch. Big Sheep Creek,

Alder Springs, Wallowa River Region. DREPANOCLADUS ADUNCUS (Hedw.) Warns. forma AQUATICUS

(Sanio) Moenkem. Cornucopia. DREPANOCLADUS BERGGRENII (C. Jens.) Broth. Alder Springs. EURHYNCHIUM OREGANUM (Sull.) Jaeger & Sauerb. Cornucopia. HOMALOTHECIUM NUTTALLII (Wils.) Grout. Wallowa River

Region, Cornucopia. HYPNUM CURVIFOLIUM Hedw. Aneroid Lake, Ice Lake, Cornucopia,

Wallowa River Region. HYPNUM IMPONENS Hedw. Wallowa River Region. HYPNUM REVOLUTUM (Mitt.) Lindb. Alder Springs. POROTRICHUM ALLEGANIENSE (C. M.) Grout. Cornucopia. PSEUDISOTHECIUM MYOSURIOIDES Dill. Aneroid Lake, Ice Lake. PSEUDISOTHECIUM STOLONIFERUM Hook. Alder Springs. RHYTIDIADELPHUS SQUARROSUS (L). Hedw. Warnst. Alder

Springs. RHYTIDIOPSIS ROBUSTA (Hook.) Broth. Wallowa River Region. SCLEROPODIUM CAESPITOSUM (Wils.) Br. & Sch. Wallowa River

Region, Joseph.

94 THE BRYOLOGIST [AUGUST



ANDREWS: REVIEW ANDREWS: REVIEW

LESKEACEAE ANOMODON VITICULOSUS (L.) Hook. & Tayl. Cornucopia. CLAOPODIUM CRISPIFOLIUM (Hook.) R. & C. Wallowa River

Region. CLAOPODIUM WHIPPLEANUM (Sull.) R. & C. Alder Springs.

HOOKERIACEAE

HOOKERIA LUCENS (L.) Sm. Wallowa River Region.

CRYPHAEACEAE ANTITRICHIA CALIFORNICA C. M. & Kindb. Wallowa River Region.

FABRONIACEAE FABRONIA PUSILLA Raddi. Aneroid Lake Trail.

FONTINALACEAE

DICHELYMA FALCATUM (Hedw.) Myr. Cornucopia. FONTINALIS ANTIPYRETICA (L.) Hedw. var. OREGONENSIS R. & C.

Big Sheep Creek.

REVIEW W. KABIERSCH. Studien fiber die ostasiatischen Arten einiger Laub-

moosfamilien (Mniaceae-Bartramiaceae). Hedwigia 76: 1-94, 1936; 77: 71-136, 1937. The families of mosses here discussed are the not altogether closely

related Mniaccae, Rhizogoniaceac, Bartramiaccac, Aulacomniaceae and Meeseaceae. Most attention, in fact practically the whole 1936 publication, is devoted to the Mniaceae and it is this part that I wish to comment upon.

Such revisions are decidedly more necessary than any other phase of bryological work, and Kabiersch has done well in clearing away some of the useless ballast of synonyms, but could perhaps have gone still farther in this regard with profit to science. In dealing with the eastern Asiatic species he has naturally and rightly considered them against the background of the European and North American species, as also those of the rest of Asia, in fact of the world in general, the Mniaceae not penetrating in many species into the tropics and south- ern hemisphere. In revising the North American species I have similarly considered, as far as material permitted, those of eastern Asia and the world in general, including work at the Herbier Boissier at Geneva in the winter of 1934, and I should like to record consider- able agreement with Kabiersch, but at the same time some little disa- greement on various points, including a few necessary corrections.

In the first place I should take a more synthetic rather than extreme analytic attitude toward divisions within the family. This is admit- tedly not of great importance as long as the divisions are recognizably natural. On this latter point I should like to submit several notes.

LESKEACEAE ANOMODON VITICULOSUS (L.) Hook. & Tayl. Cornucopia. CLAOPODIUM CRISPIFOLIUM (Hook.) R. & C. Wallowa River

Region. CLAOPODIUM WHIPPLEANUM (Sull.) R. & C. Alder Springs.

HOOKERIACEAE

HOOKERIA LUCENS (L.) Sm. Wallowa River Region.

CRYPHAEACEAE ANTITRICHIA CALIFORNICA C. M. & Kindb. Wallowa River Region.

FABRONIACEAE FABRONIA PUSILLA Raddi. Aneroid Lake Trail.

FONTINALACEAE

DICHELYMA FALCATUM (Hedw.) Myr. Cornucopia. FONTINALIS ANTIPYRETICA (L.) Hedw. var. OREGONENSIS R. & C.

Big Sheep Creek.

REVIEW W. KABIERSCH. Studien fiber die ostasiatischen Arten einiger Laub-

moosfamilien (Mniaceae-Bartramiaceae). Hedwigia 76: 1-94, 1936; 77: 71-136, 1937. The families of mosses here discussed are the not altogether closely

related Mniaccae, Rhizogoniaceac, Bartramiaccac, Aulacomniaceae and Meeseaceae. Most attention, in fact practically the whole 1936 publication, is devoted to the Mniaceae and it is this part that I wish to comment upon.

Such revisions are decidedly more necessary than any other phase of bryological work, and Kabiersch has done well in clearing away some of the useless ballast of synonyms, but could perhaps have gone still farther in this regard with profit to science. In dealing with the eastern Asiatic species he has naturally and rightly considered them against the background of the European and North American species, as also those of the rest of Asia, in fact of the world in general, the Mniaceae not penetrating in many species into the tropics and south- ern hemisphere. In revising the North American species I have similarly considered, as far as material permitted, those of eastern Asia and the world in general, including work at the Herbier Boissier at Geneva in the winter of 1934, and I should like to record consider- able agreement with Kabiersch, but at the same time some little disa- greement on various points, including a few necessary corrections.

In the first place I should take a more synthetic rather than extreme analytic attitude toward divisions within the family. This is admit- tedly not of great importance as long as the divisions are recognizably natural. On this latter point I should like to submit several notes.

1938] 1938] 95 95



the moss flora of the wallowa mountains of oregon

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