the special senses - linn–benton community collegecf.linnbenton.edu/mathsci/bio/jacobsr/upload/12...
TRANSCRIPT
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HW page 9: Matching Eye Disorders
PreLab 5
HOMEWORK DUE IN LAB 5
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Hearing and Equilibrium
THE SPECIAL SENSES
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The organ of hearing and equilibrium
Cranial nerve VIII - Vestibulocochlear
Regions
External ear
Middle ear
Internal ear (labyrinth)
THE EAR
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Figure 15.25a
External
acoustic
meatus
Auricle
(pinna)
(a) The three regions of the ear
Helix
Lobule
Pharyngotympanic
(auditory) tube
Tympanic
membrane
External
ear
Middle
ear
Internal ear
(labyrinth)
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Copyright © 2010 Pearson Education, Inc. Figure 15.26
Pharyngotym-
panic tube
Tensor
tympani
muscle
Tympanic
membrane
(medial view)
Stapes
Malleus
View
Superior
Anterior
Lateral
Incus Epitympanic
recess
Stapedius
muscle
Middle Ear
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Auditory tube
Connects the middle ear to the nasopharynx
Equalizes pressure
Opens during swallowing and yawning
Otitis media
MIDDLE EAR
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Contains functional organs for hearing & equilibrium
Bony labyrinth
Membranous labyrinth
INNER EAR
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Figure 15.27
Anterior
Semicircular
ducts in
semicircular
canals
Posterior
Lateral
Cristae ampullares
in the membranous
ampullae
Utricle in
vestibule
Saccule in
vestibule Stapes in
oval window
Temporal bone
Facial nerve
Vestibular
nerve
Superior vestibular ganglion
Inferior vestibular ganglion
Cochlear
nerve
Maculae
Spiral organ
(of Corti) Cochlear
duct
in cochlea
Round
window
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Three distinct regions
Vestibule
Gravity
Head position
Linear acceleration and deceleration
Semicircular canals
Angular acceleration and deceleration
Cochlea
Vibration
INNER EAR - BONY LABYRINTH
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Figure 15.27
Anterior
Semicircular
ducts in
semicircular
canals
Posterior
Lateral
Cristae ampullares
in the membranous
ampullae
Utricle in
vestibule
Saccule in
vestibule Stapes in
oval window
Temporal bone
Facial nerve
Vestibular
nerve
Superior vestibular ganglion
Inferior vestibular ganglion
Cochlear
nerve
Maculae
Spiral organ
(of Corti) Cochlear
duct
in cochlea
Round
window
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The cochlea
A spiral, conical, bony chamber Vestibular canal (scala vestibuli)
Cochlear duct (scala media)
Tympanic canal (scala tympani)
INNER EAR
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Figure 15.28a
(a) Helicotrema
Modiolus Cochlear nerve, division of the vestibulocochlear nerve (VIII)
Cochlear duct
(scala media)
Spiral ganglion
Osseous spiral lamina
Vestibular membrane
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Cavity of the cochlea is divided into 3 chambers
Vestibular canal (scala vestibuli)
Vestibular membrane
Cochlear duct (scala media)
Basilar membrane supporting Organ of Corti
Tympanic canal (scala tympani)
INNER EAR
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Copyright © 2010 Pearson Education, Inc. Figure 15.28b
(b)
Cochlear duct (scala media; contains endolymph)
Tectorial membrane
Vestibular membrane
Scala vestibuli (contains perilymph)
Scala tympani
(contains
perilymph) Basilar
membrane
Spiral organ
(of Corti)
Stria
vascularis
Spiral
ganglion
Osseous spiral lamina
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Figure 15.28c
(c)
Tectorial membrane Inner hair cell
Outer hair cells
Hairs (stereocilia) Afferent nerve
fibers
Basilar
membrane
Fibers of cochlear nerve
Supporting cells
Organ of Corti
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Transduction of sound
Mechanical energy in middle ear
Fluid pressure wave in inner ear
Nerve impulse
PHYSIOLOGY OF HEARING
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Sounds set up vibrations in air that beat against the eardrum
that pushes a chain of tiny bones that press fluid in the
internal ear against membranes that set up shearing forces
that pull on the tiny hair cells that stimulate nearby neurons
that give rise to the impulses that travel to the brain – and
you hear.
(This is from your textbook)
PHYSIOLOGY OF HEARING IN A NUTSHELL
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Figure 15.29
Area of
high pressure
(compressed
molecules)
Crest
Trough
Distance Amplitude
Area of
low pressure
(rarefaction)
A struck tuning fork alternately compresses
and rarefies the air molecules around it,
creating alternate zones of high and
low pressure.
(b) Sound waves
radiate outward
in all directions.
Wavelength
Air p
re
ssu
re
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Copyright © 2010 Pearson Education, Inc. Figure 15.30
Time (s)
(a) Frequency is perceived as pitch.
High frequency (short wavelength) = high pitch Low frequency (long wavelength) = low pitch
(b) Amplitude (size or intensity) is perceived as loudness.
High amplitude = loud Low amplitude = soft
Time (s)
Pre
ssu
re
P
re
ssu
re
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Scala
vestibuli
Cochlear duct
(contains endolymph)
Scala
tympani
Perilymph
Basilar
membrane
Cochlea
Sound waves
Helicotrema Stapes vibrating
in oval window Malleus Incus
External auditory
canal
Tympanic
membrane
Secondary tympanic
membrane vibrating
in round window Auditory tube
Vestibular membrane
Middle ear
Tectorial membrane
Spiral organ
(organ of Corti)
1
Scala
vestibuli
Cochlear duct
(contains endolymph)
Scala
tympani
Perilymph
Basilar
membrane
Cochlea
Sound waves
Helicotrema Stapes vibrating
in oval window Malleus Incus
External auditory
canal
Tympanic
membrane
Secondary tympanic
membrane vibrating
in round window Auditory tube
Vestibular membrane
Middle ear
Tectorial membrane
Spiral organ
(organ of Corti)
1 2
Scala
vestibuli
Cochlear duct
(contains endolymph)
Scala
tympani
Perilymph
Basilar
membrane
Cochlea
Sound waves
Helicotrema Stapes vibrating
in oval window Malleus Incus
External auditory
canal
Tympanic
membrane
Secondary tympanic
membrane vibrating
in round window Auditory tube
Vestibular membrane
Middle ear
Tectorial membrane
Spiral organ
(organ of Corti)
1 2
3
Scala
vestibuli
Cochlear duct
(contains endolymph)
Scala
tympani
Perilymph
Basilar
membrane
Cochlea
Sound waves
Helicotrema Stapes vibrating
in oval window Malleus Incus
External auditory
canal
Tympanic
membrane
Secondary tympanic
membrane vibrating
in round window Auditory tube
Vestibular membrane
Middle ear
Tectorial membrane
Spiral organ
(organ of Corti)
1 2
3 4
Scala
vestibuli
Cochlear duct
(contains endolymph)
Scala
tympani
Perilymph
Basilar
membrane
Cochlea
Sound waves
Helicotrema Stapes vibrating
in oval window Malleus Incus
External auditory
canal
Tympanic
membrane
Secondary tympanic
membrane vibrating
in round window Auditory tube
Vestibular membrane
Middle ear
Tectorial membrane
Spiral organ
(organ of Corti)
1 2
3 4
5 Scala
vestibuli
Cochlear duct
(contains endolymph)
Scala
tympani
Perilymph
Basilar
membrane
Cochlea
Sound waves
Helicotrema Stapes vibrating
in oval window Malleus Incus
External auditory
canal
Tympanic
membrane
Secondary tympanic
membrane vibrating
in round window Auditory tube
Vestibular membrane
Middle ear
Tectorial membrane
Spiral organ
(organ of Corti)
1 2
3 4
5
6
Scala
vestibuli
Cochlear duct
(contains endolymph)
Scala
tympani
Perilymph
Basilar
membrane
Cochlea
Sound waves
Helicotrema Stapes vibrating
in oval window Malleus Incus
External auditory
canal
Tympanic
membrane
Secondary tympanic
membrane vibrating
in round window Auditory tube
Vestibular membrane
Middle ear
Tectorial membrane
Spiral organ
(organ of Corti)
1 2
3 4
5
6
7
Scala
vestibuli
Cochlear duct
(contains endolymph)
Scala
tympani
Perilymph
Basilar
membrane
Cochlea
Sound waves
Helicotrema Stapes vibrating
in oval window Malleus Incus
External auditory
canal
Tympanic
membrane
Secondary tympanic
membrane vibrating
in round window Auditory tube
Vestibular membrane
Middle ear
Tectorial membrane
Spiral organ
(organ of Corti)
1 2
3 4
5
6
7 8
8
Scala
vestibuli
Cochlear duct
(contains endolymph)
Scala
tympani
Perilymph
Basilar
membrane
Cochlea
Sound waves
Helicotrema Stapes vibrating
in oval window Malleus Incus
External auditory
canal
Tympanic
membrane
Secondary tympanic
membrane vibrating
in round window Auditory tube
Vestibular membrane
Middle ear
Tectorial membrane
Spiral organ
(organ of Corti)
1 2
3 4
5
6
7 8
8
9
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Copyright © 2010 Pearson Education, Inc. Figure 15.31a
Scala tympani
Cochlear duct
Basilar
membrane
1 Sound waves vibrate
the tympanic membrane.
2 Auditory ossicles vibrate.
Pressure is amplified.
3 Pressure waves created by
the stapes pushing on the oval
window move through fluid in
the scala vestibuli.
Sounds with frequencies
below hearing travel through
the helicotrema and do not
excite hair cells.
Sounds in the hearing range
go through the cochlear duct,
vibrating the basilar membrane
and deflecting hairs on inner
hair cells.
Malleus Incus
Auditory ossicles
Stapes
Oval
window
Scala vestibuli
Helicotrema
Cochlear nerve
3 2
1
Round
window
Tympanic
membrane (a) Route of sound waves through the ear
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Copyright © 2010 Pearson Education, Inc. Figure 15.31b
Fibers of basilar membrane
(b) Different sound frequencies cross the
basilar membrane at different locations.
Medium-frequency sounds displace
the basilar membrane near the middle.
Low-frequency sounds displace the
basilar membrane near the apex.
Base
(short,
stiff
fibers)
Frequency (Hz)
Apex
(long,
floppy
fibers)
Basilar membrane
High-frequency sounds displace
the basilar membrane near the base.
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Copyright © 2010 Pearson Education, Inc. Figure 15.28c
(c)
Tectorial membrane Inner hair cell
Outer hair cells
Hairs (stereocilia) Afferent nerve
fibers
Basilar
membrane
Fibers of cochlear nerve
Supporting cells
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Figure 15.33
Medial geniculate
nucleus of thalamus
Primary auditory
cortex in temporal lobe
Inferior colliculus
Lateral lemniscus
Superior olivary nucleus
(pons-medulla junction)
Spiral organ (of Corti)
Bipolar cell
Spiral ganglion of cochlear nerve
Vestibulocochlear nerve
Medulla
Midbrain
Cochlear nuclei
Vibrations
Vibrations
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Timing comparison
Side nearest sound detects sound first
Comparison of volume
High pitched = blocked by head
Low pitched = curve around head
LOCALIZATION OF SOUND
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Conductive deafness Interference in movement of middle ear bones
Impacted earwax, perforated eardrum or otosclerosis of the ossicles
Sensorineural deafness Damage to the neural structures
Aging, prolonged exposure to loud sounds
ABNORMALITIES OF HEARING
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Otitis media
ABNORMALITIES OF HEARING
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Otosclerosis
ABNORMALITIES OF HEARING
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Meniere’s Disease
ABNORMALITIES OF HEARING
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Tinnitus
ABNORMALITIES OF HEARING
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Information about position and movement of head
Vestibular apparatus
Utricle
Saccule
Semi-circular canals
PHYSIOLOGY OF EQUILIBRIUM
Vesibule
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Figure 15.34
Macula of
saccule
Otoliths
Hair bundle
Kinocilium
Stereocilia Otolithic
membrane
Vestibular
nerve fibers
Hair cells
Supporting
cells
Macula of
utricle
Vestibular Apparatus
Maculae are
perpendicular
to one another
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Macula
Utricle
Horizontal movements
Tilting the head side to side
Saccule
Vertical movements
VESTIBULE
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Crista ampullaris
Sensory receptor for dynamic equilibrium
One in the ampulla of each semicircular canal
Major stimuli are rotatory movements
SEMI-CIRCULAR CANALS
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Copyright © 2010 Pearson Education, Inc. Figure 15.36a–b
Fibers of vestibular nerve
Hair bundle (kinocilium
plus stereocilia)
Hair cell
Supporting
cell
Membranous
labyrinth
Crista
ampullaris
Crista
ampullaris
Endolymph
Cupula
Cupula
(a) Anatomy of a crista ampullaris in a
semicircular canal
(b) Scanning electron
micrograph of a
crista ampullaris
(200x)
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Conflicts between eye movements and equilibrium
Nystagmus
Slow component
Fast component
MOTION SICKNESS