vascular plant species checklist of bighorn canyon national

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Vascular Plant Species Checklist of Bighorn Canyon National Recreation Area, Montana and Wyoming Prepared for the National Park Service - Bighorn Canyon National Recreation Area and the Greater Yellowstone Network By Bonnie Heidel and Walter Fertig Wyoming Natural Diversity Database University of Wyoming PO Box 3381, Laramie, WY 82071 January 2002 . Insert 4 pictures of "new" species discoveries – available upon request

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Page 1: Vascular Plant Species Checklist of Bighorn Canyon National

Vascular Plant Species Checklist

of Bighorn Canyon National Recreation Area,

Montana and Wyoming

Prepared for the National Park Service - Bighorn Canyon National Recreation Area

and the Greater Yellowstone Network

By Bonnie Heidel and Walter Fertig

Wyoming Natural Diversity Database

University of Wyoming PO Box 3381, Laramie, WY 82071

January 2002

.

Insert 4 pictures of "new" species discoveries – available upon request

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Table of Contents

Page # Abstract . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 Acknowledgments . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 Study Area . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 Methods . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 Results . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 Summary of Plant Inventory Work at Bighorn Canyon National Recreation Area . . 9

Flora of Bighorn Canyon National Recreation Area . . . . . . . . . . . . . . . . . . . . . . . . . 11 Noxious Weed Documentation . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 12

Additional Rare Plant Records in Bighorn Canyon National Recreation Area . . . . . . 12

Discussion . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14 Literature Cited . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 16

Cover caption: Additions to the Bighorn Canyon National Recreation Area flora include Castilleja exilis (Big Horn County, WY), Dodecatheon conjugens (Big Horn and Carbon cos., MT), Oxytropis besseyi var. ventosa (Carbon County, Montana), and Rosa arkansana (Big Horn County, Montana; starting upper left, moving clockwise.)

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Figures, Tables, and Appendices

Page # Figure 1. Bighorn Canyon National Recreation Area . . . . . . . . . . . . . . . . . . . . . . . . . 7 Figure 2. Increase in Number of Plant Taxa Recorded at Bighorn Canyon National Recreation Area, 1924-2001 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 Table 1. Chronology of Major Plant Collecting Efforts at Bighorn Canyon National Recreation Area . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10 Table 2. Summary of the Flora of Bighorn Canyon National Recreation Area . . . . . . . . 12 Table 3. Summary of the Flora of Bighorn Canyon National Recreation Area by County Subunits . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 12 Table 4. State Noxious Weeds Confirmed or Reported from Bighorn Canyon National Recreation Area . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 13 Table 5. Increase in Number of Plant Taxa Recorded at Bighorn Canyon National Recreation Area, 1924-2001 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14 Appendix A. Annotated Vascular Plant Checklist of the Bighorn Canyon National Recreation Area . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 18 Appendix B. Rejected Taxa . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 59 Appendix C. Unconfirmed Taxa . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 63 Appendix D. Potential Vascular Plant Flora of Bighorn Canyon National Recreation Area . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 66

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ABSTRACT Under congressional mandate, each unit of the National Park Service is required to develop comprehensive species lists that capture at least 90% of their total flora and fauna. Bighorn Canyon National Recreation Area was surveyed extensively from 1976-1984, resulting in the documentation of over 650 vascular plant taxa for the park. Rare plant surveys in 1998-99, however, resulted in the discovery of 60 additional species, calling into question whether Bighorn Canyon NRA had achieved the 90% threshold. In 2001 we were contracted to revise the existing checklist of the park’s flora and conduct field surveys in under-sampled habitats to identify potentially missing species. After extensive review of the existing literature and herbarium collections at Bighorn Canyon and the Rocky Mountain Herbarium, we identified 81 species that had been falsely reported for the park and 45 additional taxa that could not be verified. Our herbarium search and fieldwork also identified over 120 additional species that had not previously been identified for Bighorn Canyon, including 22 additional weed species. Our revised checklist now contains 739 vascular plant taxa, all of which have been verified with voucher specimens or reliable observation data. We also compiled a list of 148 potential species that could occur in the study area due to the presence of suitable habitat. If we add the 44 potential species with the highest probability of occurring in the park, the current checklist would still represent over 94% of the total flora. Some relatively inaccessible areas of the National Recreation Area and cryptic or taxonomically difficult species groups remain undersampled, but overall, the known flora of Bighorn Canyon is probably at or near the 90% target level. ACKNOWLEDGMENTS We thank National Park Service staff for their interest, help and support. Lane Cameron (Biological Inventory Coordinator for the Greater Yellowstone Network) provided coordination throughout the project. Field plans were coordinated with Rick Lasko (Bighorn Canyon National Recreation Area). The planning contributions of Martina Keil, and the benefit of information on BICA herbarium additions compiled by Suzanne Morstad in addition to her observations were indispensible. Boat transporation was coordinated through Chris Ryan. We appreciate the opportunities to discuss this project with Darrell Cook, Theo Hugs, and other NPS staff. The herbarium and data resources of the University of Wyoming - Rocky Mountain Herbarium (RM) were vital and are acknowledged with deepest gratitude. B. E. Nelson, manager of the RM, shared information from his collections at Bighorn Canyon in 1979-1980. Dr. Dennis Knight of the University of Wyoming provided a box of old data forms, photos, and original notes from the floristic and ecological studies of Robert Lichvar, Ellen Collins, and Yoshiko Akashi at Bighorn Canyon in the early 1980s. We thank Scott Laursen and Melanie Arnett for help in compiling herbarium information at the RM, and Martin Miller (Montana Natural Heritage Program) for help processing the 2001 rare plant information in Montana. Stuart Markow helped greatly in specimen review. The report benefited by the reviews of Robert Dorn, B. E. Nelson, Lane Cameron and Suzanne Morstad. This project was conducted in collaboration with the Montana Natural Heritage Program. It was supported by the Inventory and Monitoring Initiative of the National Park Service - Greater Yellowstone Network.

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INTRODUCTION The National Park Service established Bighorn Canyon National Recreation Area* in 1966 to promote recreational opportunities on Yellowtail Reservoir and to protect historical sites, such as the Mason-Lovell Ranch and Bad Pass Trail. Due to its location at the confluence of the Great Plains, Central Rocky Mountains, and Wyoming Basins ecoregions, Bighorn Canyon harbors an unusually rich diversity of vegetation types and over two dozen locally endemic or rare plant species (Knight et al. 1987; Heidel and Fertig 2000). From a conservation perspective, Bighorn Canyon National Recreation Area is significant because it is one of the few areas in the northern Bighorn Basin and adjacent foothills of the Bighorn/Pryor Mountains that is permanently protected and managed with an emphasis on maintaining biological processes (Merrill et al. 1996). An important mission of the National Park Service is to conserve the natural, recreational and cultural resources of each park unit for current and future generations. In spite of this emphasis, many parks have lacked comprehensive inventories of their flora and fauna or monitoring programs to assess impacts of management actions on target species. In 1998, Congress directed the Park Service to develop a program to inventory and monitor the biological resources of the entire park system in order provide better informed management decisions. Since 1999, the Park Service has been conducting regional expert workshops to determine the state of existing knowledge on the biota of each park and to identify data gaps and research needs. The ultimate goal of this effort is to attain a 90% complete species checklist and develop monitoring programs for each park unit (National Park Service 2000). An initial floristic survey of Bighorn Canyon by Lichvar et al. (1985) documented 656 vascular plant taxa for the National Recreation Area. In conjunction with a rare plant inventory in 1998-1999, Heidel and Fertig (2000) documented 60 additional plant taxa for the NRA, calling into question whether the existing checklist represented at least 90% of the actual flora. In April 2001, the Greater Yellowstone Network of the National Park Service contracted with the University of Wyoming’s Natural Diversity Database (WYNDD) to revise the existing checklist and conduct new field surveys to assess whether the 90% representation goal had been achieved. In addition, WYNDD was charged with documenting new localities of noxious weeds and rare plants. The results of this study are summarized on the following pages. This report includes an annotated checklist of the known flora of Bighorn Canyon, lists of falsely reported and unconfirmed species, and a list of plants that potentially occur in the area. STUDY AREA Bighorn Canyon National Recreation Area parallels Yellowtail Reservoir (Bighorn Lake) for 71 miles from the vicinity of Kane, Wyoming to Fort Smith, Montana (Figure 1). It encompasses 22,499 ha (55,595 acres) and ranges in elevation from 1100-2450 meters (3600-8040 ft). It also includes the Yellowtail Reservoir that has completely inundated the original Bighorn Canyon. * The NPS code that is used to denote Bighorn Canyon National Recreation Area is “BICA”. This code is used in the report to refer to both the study area and the NPS herbarium facility in particular.

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Downcutting by the Bighorn River and regional faulting and erosion have exposed thick Paleozoic limestone and dolomite cliffs, soft red Cretaceous sandstones, and Quaternary alluvium and pediments (Love and Christensen 1985). Knight et al. (1987) documented 21 main vegetation types in the study area, ranging from lakeshore marshes and floodplain woodlands to prairie grasslands and coniferous forests at higher elevations. The most common vegetation type by far, however, is Utah juniper/Curlleaf mountain mahogany, which covers over 40% of the NRA. Based on vegetation, geology, and landforms, the Bighorn Canyn National Recreation Area can be subdivided into three distinct biogeographic units that roughly correspond with the boundaries of the three counties that intersect the park. The southernmost segment lies in Big Horn County, Wyoming. Much of this area consists of rocky outcrops of red sandstone, and desert shale barrens dominated by Utah juniper, curlleaf mountain-mahogany, greasewood, shadscale, Gardner’s saltbush, and other desert species with affinities to the flora of the Great Basin. This section is the driest portion of the study area and averages just 6.7 inches of annual precipitation (based on NOAA records from Lovell, WY). The southern unit also contains riparian woodlands, extensive marshes along the floodplain of Yellowtail Reservoir and uplands that are in the Yellowtail Wildlife Habitat Management Area, co-managed with the Wyoming Game and Fish Department. The middle segment of the study area is mainly in Carbon County, Montana. It has the highest elevation of the three segments because it includes the face of the Pryor Mountains escarpment, but is primarily a dry foothills landscape, dissected by steep limestone and dolomite canyons. The vegetation consists of arid foothills grasslands, Utah juniper and curlleaf mountain mahogany shrublands, and limited coniferous forests of Douglas-fir, limber pine, and Engelmann spruce and subalpine fir (Knight et al. 1987). This area also contains numerous cold water seeps that provide habitat for a number of rare wetland plants, including Sullivantia hapemanii (Heidel and Fertig 2000). Due to its diversity of habitats and topography, the middle segment has the highest species richness of any of the three segments comprising the study area. Lichvar et al. (1985) reported a number of subalpine and alpine taxa from this area, but most of these species were actually found on one section of state land just west of the present National Recreation Area boundaries. The southern and middle segments of the NRA contain most of its local and regional endemic species, such as Eriogonum brevicaule var. canum, Erigeron allocotus, and Rorippa calycina. The Montana portion also contains a high percentage of rare desert plants at the northern margins of their range that are found nowhere else in that state (Dorn 1978; Heidel and Fertig 2000). The northernmost segment of the study area lies within Big Horn County, Montana. It is the wettest segment of the study area. The area around Fort Smith has an annual precipitation averaging 18.9 inches. It is a separate unit connected to the other segments by the Yellowtail Reservoir. The land area of this northernmost unit is over 10 times smaller than the other two segments. However, it includes well-developed mesic valley bottom vegetation and perennial coldwater streams within 30 ft. of the reservoir highwater level under NPS administration, unlike the other segments. In addition, the north unit contains Great Plains vegetation, foothills woodlands, and a short reach of the free-flowing Big Horn River. The Three Mile Fishing Access area located 3 miles north of the northeastern end of the National Recreation Area was not included in this study.

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Figure 1. Bighorn Canyon National Recreation Area

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BIGHORN CANYONNATIONALRECREATION AREA No Trespassing

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Pryor MountainWild Horse Range

YellowtailWildlife HabitatManagement Area

Picnic area

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Site of Historic Fort C. F. SmithPrivate Property

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METHODS Lichvar et al. (1985) assembled a vascular plant species checklist for Bighorn Canyon National Recreation Area based on collections made during floristic and vegetation surveys in 1983-84. Heidel and Fertig (2000) amended this list to include species records from their rare plant survey work in 1998-99, as well as additional species reported by Knight et al. (1987), and additions among the collections of the Bighorn Canyon National Recreation Area Herbarium (BICA). Prior to survey work in 2001, we reevaluated every entry in our preliminary checklist to ensure that all species were corroborated by a valid voucher specimen or observation. An initial sweep of the Rocky Mountain Herbarium (RM) revealed a number of specimens that were either misidentified or were collected outside the boundaries of Bighorn Canyon National Recreation Area. In May 2001, Fertig reexamined the entire BICA collection to confirm the identity of all voucher specimens, verify their presence within the study area, and annotate the preliminary checklist with county-level distribution data. From this new information we developed a working checklist that highlighted those species still in need of confirmation in the field or from the collections at RM. Field surveys in 2001 focused on six undersampled vegetation types that we identified during our rare plant inventory in 1998-99: riparian corridors of coldwater streams, cushion plant communities, marsh and forest riparian habitat, cliffs, montane grassland, and seeps (Heidel and Fertig 2000). We also inventoried those sites that had more difficult access and areas that required repeat visits to capture the phenological array of species. In particular, we emphasized early and late-flowering graminoids, exotics, and composites that appeared to be undersampled in previous studies. Survey locations were scattered across the three segments of the study area to ensure geographic breadth of inventory, in keeping with National Park Service inventory guidelines (USDI National Park Service 2000). We did not include species planted by the National Park Service or at the landscaped headquarters in Lovell, Wyoming. We did include species that persisted from plantings that pre-date BICA establishment in 1966. Our herbarium work and fieldwork required four visits to Bighorn Canyon between 29 May and 11 August 2001. In the field we made collections of species that were either additions to the study area flora or which had not previously been vouchered. Collecting was done only if plant populations were sufficiently large. When necessary, we substituted detailed observation data for physical specimens. In the case of rare species or noxious weeds, we documented new location and habitat data to augment existing records within the study area. For all species we recorded county-of-distribution information within the National Recreation Area. Voucher specimens were taken to the Rocky Mountain Herbarium (RM) for identification and deposited there. Duplicate sheets were prepared for deposit at BICA. At the conclusion of the field season, we created a revised checklist incorporating the additional species collected or observed in 2001. We also rechecked the collections of the RM for all remaining species on the initial checklist that lacked vouchers at BICA or were not relocated in the field. Using the RM’s digital specimen database, we were able to locate several dozen historical records from the park, including 49 collections made by B. Ernie Nelson in 1979-80 that were not included by Lichvar et al. (1985). The entire checklist was revised to follow the standard nomenclature of Kartesz and Meacham (1999) as used in NPSpecies database for scientific and common names. Synonyms were provided if these names differed from those used in the current statewide floras of Montana and Wyoming (Dorn 1984, 2001).

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Our final checklist contains all vascular plant taxa known to occur in Bighorn Canyon National Recreation Area based on verified voucher specimens or observations (Appendix A). This list has been annotated to include global heritage rank (NatureServe 2001, http://natureserve.abi.org/), native or exotic status (Fertig 1999a), distribution within the National Recreation Area by county, a representative voucher collection (with collector and repository), date of first documentation in the study area, and other pertinent comments. Appendix B and C contain lists of species that were previously reported for the park but have either been rejected or cannot be confirmed at present. To test whether the flora of Bighorn Canyon has been adequately sampled, we developed an additional list of species that may potentially occur within the area, but which have not yet been documented (Appendix D). This list was created by comparing our confirmed species list with the Rocky Mountain Herbarium’s digital “Atlas of the Vascular Plants of Wyoming” (www.esb.utexas.edu/chumley/wyomap/atlas.htm) and checklists of the flora of the Pryor Mountains, Bighorn Range, and Bighorn Basin (Lesica and Achuff 1992, Fertig 1999b). The probability of occurrence for each species was given a rank of high, medium, or low depending on its global abundance and the presence of suitable habitat in the study area. Additional comments are provided on where these species are most likely to be found within Bighorn Canyon. RESULTS Summary of Plant Inventory Work at Bighorn Canyon National Recreation Area Prior to 1976, botanical studies in the Bighorn Canyon area were limited to sporadic collections or baseline ecological surveys of dominant community types (Table 1). The earliest known collection from the area is a specimen of Sarcobatus vermiculatus made by R.C. Snelling in 1924 (Rocky Mountain Herbarium records). During the next 50 years, only three additional species were collected and deposited at the RM from the Wyoming portion of the NRA (early records from Montana are poorly known, although W.E. Booth made a small number of collections in the vicinity of Bighorn Canyon in the 1950s and 1960s, deposited at Montana State University.) All previous botany and vegetation studies cited by Lichvar et al. (1985) and Knight et al. (1987) are from outside the BICA study area. They did not mention and we were not able to determine the source of two unpublished plant checklists for Bighorn Canyon in 1968 (110 species) and 1975 (225 species) (Anonymous 1968, 1975). The source and authenticity of these floristic records has not been confirmed. Robert Dorn documented 24 additional plant taxa for the NRA in 1976-77 while doing general collecting in Carbon County, Montana (Dorn 1978). M.L. Roberts, a University of Wyoming graduate student, added one species in 1976. Collections were also made by B. E. Nelson, University of Wyoming (RM) in 1979-80 as part of a multi-year floristic survey of the Bighorn Range and surrounding basins (Nelson and Hartman 1984). Nelson documented 49 additional species for the NRA at the mouth of John Blue Canyon and along Yellowtail Reservoir in Big Horn County, Wyoming. In 1981-82, Park Service employees Terry Peters, Kathy Jobe, William Messerschmidt, and Arlene Wolf made over 100 collections from Horseshoe Bend, Trail Creek, and Barrys Landing in Wyoming and Montana and added 56 additional species to the park’s flora (BICA records).

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Table 1. Chronology of Major Plant Collecting Efforts at Bighorn Canyon NRA Year(s) Collector

(main repository)

Number of Additional Species

Collection Area Comments

1924-1966

R.C. Snelling, W.E. Booth, D. Despain, and E. Clausen (RM)

4 Bighorn Canyon, WY. Identified from Rocky Mountain Herbarium digital location database.

1976-1977

Robert Dorn (RM, MONTU) and M.L. Roberts (RM)

25 Pryor Desert and Bighorn Mountains in Montana

Cited in Lichvar et al. (1985) and highlighted in a Madrono article on significant new records of the Great Basin flora for Montana (Dorn 1978).

1979-1980

B. E. Nelson (RM)

49 Mouth of John Blue Canyon in WY on east side of Yellowtail Reservoir (part of floristic survey the Bighorn Range) and locally on the west side

The collections include additions to the study area flora. They were not cited in Lichvar et al. (1985).

1981-1982

Terry Peters, Kathy Jobe, William Messerschmidt, and Arlene Wolf (BICA)

56 Mostly near Horseshoe Bend Campground, Trail Creek Campground, and Barry’s Landing in Wyoming and Montana

The collections include additions to the study area flora. They were not cited in Lichvar et al. (1985).

1983 Robert Lichvar, Ellen Collins, and Dennis Knight (RM, BICA, NY)

384 Throughout Bighorn Canyon NRA in Wyoming and Montana as part of floristic survey and vegetation study

Principle source of information presented in Lichvar et al. (1985)

1984 George Jones, Dale Ditolla, and Richard Myers (RM)

47 Throughout Bighorn Canyon NRA in Wyoming and Montana as part of vegetation study

The collections include additions to the study area flora. Most are reported in Knight et al. (1987), but not all are in Lichvar et al. (1985).

1994-1996

Walter Fertig and Robert Dorn (RM)

3 Horseshoe Bend and Yellowtail Wildlife Habitat Management Unit, WY

Cited in Heidel and Fertig (2000)

1998-1999

Bonnie Heidel, Walter Fertig, and Laura Welp (RM, MONTU, BICA)

60 Throughout Bighorn Canyon NRA in Wyoming and Montana as part of rare plant survey

Additions to the study area flora were reported in the appendix of Heidel and Fertig (2000).

2001 Bonnie Heidel, Walter Fertig, and Suzanne Morstad (RM, BICA)

111 Throughout Bighorn Canyon NRA in Wyoming and Montana as part of floristic survey

Results in this report.

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The first truly systematic inventory of the flora of Bighorn Canyon National Recreation Area was conducted in 1983 by Robert Lichvar and Ellen Collins of the Wyoming Natural Heritage Program in collaboration with Dr. Dennis Knight of the University of Wyoming. Lichvar and Collins made over 600 collections (deposited at BICA, RM, and the New York Botanical Garden) and reported nearly 500 additional species for study area. In 1984, Knight and his student collaborators (George Jones, Dale Dittola, Richard Myers, and Yoshiko Akashi) added several hundred additional vouchers and at least 47 additional taxa (BICA and RM records) in the course of documenting the study area vegetation. Based on their own collections and the records of Dorn (1978), Lichvar et al. (1984, 1985) reported 656 vascular plant taxa for Bighorn Canyon. This list did not include many of the earlier records of B. E. Nelson deposited at the RM or specimens deposited at BICA by National Park Service staff. The Lichvar et al. (1985) checklist also contained at least 60 taxa that were misidentified or collected outside of Bighorn Canyon National Recreation Area and 45 species that cannot be confirmed today with vouchers. Based on our reassessment, the 1983 and 1984 collections of Lichvar, Collins, Knight, Jones, and associates added 431 additional species to the flora. Only three species were added to the flora between 1985 and 1998. Bonnie Heidel, Walter Fertig, and Laura Welp of the Montana and Wyoming natural heritage programs documented 60 additional taxa at Bighorn Canyon National Recreation Area during rare plant surveys in 1998-99. Heidel and Feritg (2000) compiled a revised checklist containing all additional records collected since 1985 and reports from Knight et al. (1987), and the collections at BICA that were omitted by Lichvar et al. (1985). This list identified 774 vascular plant taxa for the NRA. Flora of Bighorn Canyon National Recreation Area Based on our field surveys in 2001 and review of existing voucher specimens at RM and BICA, the flora of Bighorn Canyon National Recreation Area currently consists of 739 vascular plant taxa (Table 2, Appendix A). We observed or collected 76 additional species for the area in 2001 and relocated 33 taxa previously reported by Lichvar et al. (1985) that had been misidentified or lacked corroborating voucher specimens from within Bighorn Canyon. Suzanne Morstad (NPS) discovered 2 additional species in 2001 and we located 8 additional species records from previously overlooked specimens at RM and BICA. Our surveys and herbarium work added 407 county distribution records within the NRA, increasing the size of each subunit’s flora by 28-36% (Table 3, Appendix A). Our revised checklist does not include 81 species from Heidel and Fertig (2000) or Lichvar et al. (1985) that have been shown to be falsely reported (Appendix B). Of these taxa, 51 were misidentified and 30 were collected just outside of Bighorn Canyon National Recreation Area. An additional 45 species reported from previous checklists still cannot be confirmed by authenticated voucher specimens or observations (Appendix C). According to Robert Dorn (personal communication), some of these “missing” specimens may have been deposited at the New York Botanical Garden, rather than RM or BICA (we have been unable to confirm this). We believe that at least 34 of the 81 rejected species and 37 of the 45 unconfirmed species still have some potential to be found in the study area based on their occurrence in similar habitats in the vicinity (Appendix D).

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Table 2. Summary of the flora of Bighorn Canyon National Recreation Area1 Taxonomic Level Tally in Lichvar et al. (1985) Current Tally (2001) Family 73 83 Genus 320 372 Species/ varieties 656 739 Table 3. Summary of the Flora of Bighorn Canyon National Recreation Area by County Subunits County Total # of Taxa # of Additional Taxa

Documented in 2001 Percentage Increase in # of Taxa in 2001

Big Horn, WY 342 99 + 28.9% Carbon, MT 493 138 + 28.0% Big Horn, MT 463 170 + 36.7% Noxious Weed Documentation We documented 22 additional species of non-native plants on Bighorn Canyon National Recreation Area in 2001, bringing the total number of exotic plants in the park to 105 (an increase of 21.0%). Non-natives now account for 14.2% of the total flora. By comparison, exotic plants represent only 12.6% of the flora of Wyoming (Fertig 1999a). Twelve species considered noxious weeds in Montana or Wyoming occur within the study area and one additional species has been reported, but could not be confirmed in 2001 (Table 4). Although weeds were not the primary focus of our 2001 fieldwork, we documented several new populations of salt-cedar (Tamarix chinensis) and Russian knapweed (Acroptilon [Centaurea] repens) that were not previously recorded in Bighorn Canyon National Recreation Area noxious weed GIS database. Salt-cedar was located along several side drainages of Bighorn Canyon but was most abundant at the southern end of the study area in the Yellowtail Wildlife Habitat Management Area. Low-water conditions in 2001 exposed extensive areas of mudflats that are being rapidly colonized by Salt-cedar seedlings. Discrete patches of Russian knapweed were located along the Ok-a-Beh road and Frozen Leg picnic site, widely-distributed at the highwater margins, and colonizing on the newly-exposed mudflats at the southern end of Yellowtail Reservoir. Additional Rare Plant Records in Bighorn Canyon National Recreation Area No additional rare plant species were discovered during floristic surveys in 2001, although we did identify several new locations for Sweetwater milkvetch (Astragalus aretioides), Lesica's bladderpod (Lesquerella lesicii), persistant-sepal yellowcress (Rorippa calycina) and Hapeman's sullivantia (Sullivantia hapemanii var. hapemanii). Rorippa calycina was documented at several

1 Some differences in the number of families and genera between 1985 and 2001 reflect changes in taxonomic concepts rather than the documentation of new species. Following Dorn (2001), the number of families recognized in 2001 would increase to 91.

Page 13: Vascular Plant Species Checklist of Bighorn Canyon National

13

new locations on newly exposed mudflats at the south end of Yellowtail Reservoir, demonstrating the capacity of this species to rapidly colonize new habitat given a local seedbank. One new population of Sullivantia hapemannii was discovered and an extensive habitat complex just upstream and downstream of Yellowtail Dam was surveyed. Printouts and electronic files of all rare plant records documented in the 1998-99 and 2001 studies will be submitted separately. Table 4. State Noxious Weeds Confirmed or Reported from Bighorn Canyon National Recreation Area. List derived from Fertig (1999a) and Montana Weed Control Association (www.mtweed.org). Species Common Name State Noxious

Weed List Distribution in Bighorn Canyon NRA

Acroptilon repens [Centaurea repens]

Hardheads [Russian knapweed]

WY and MT WY-Big Horn, MT- Big Horn

Arctium minus Lesser burdock WY WY-Big Horn, MT-Carbon, MT-Big Horn

Cardaria chalepensis Lens-pod hoary-cress MT Reported for Bighorn Canyon NRA by Lichvar et al. (1985), but no vouchers have been relocated for confirmation and the species was not observed in the field in 2001.

Cardaria pubescens Globe-pod hoarycress WY WY-Big Horn Centaurea biebersteinii [Centaurea maculosa]

Spotted knapweed WY and MT WY-Big Horn, MT-Carbon, MT-Big Horn

Centaurea diffusa White knapweed [Bushy knapweed, Diffuse knapweed]

WY and MT MT-Big Horn

Cirsium arvense Canadian thistle WY and MT WY-Big Horn, MT- Big Horn

Convolvulus arvensis Field bindweed WY and MT WY-Big Horn, MT- Big Horn

Cynoglossum officinale Gypsy-flower [Common hound's-tongue]

WY and MT WY-Big Horn, MT-Carbon, MT-Big Horn

Elymus repens [Agropyron repens]

Creeping wild rye [Quackgrass]

WY MT-Carbon, MT-Big Horn

Euphorbia esula var. uralensis

Wolf's-milk [Leafy spurge]

WY and MT MT-Carbon, MT-Big Horn

Leucanthemum vulgare [Chrysanthemum leucanthemum]

Ox-eye daisy WY and MT MT-Big Horn

Tamarix chinensis Five-stamen tamarisk [Salt cedar]

WY and MT WY-Big Horn, MT-Carbon, MT-Big Horn

Page 14: Vascular Plant Species Checklist of Bighorn Canyon National

14

DISCUSSION Based on our reassessment of all previous reports and review of herbarium specimens (RM and BICA), the flora of Bighorn Canyon National Recreation Area consisted of 628 confirmed taxa as of 1999 (Table 5). Fieldwork in 2001 resulted in the discovery or relocation of 111 additional species for the park, increasing the total flora to 739 species. The net increase of 15.0% confirms our suspicions that the documented flora of Bighorn Canyon National Recreation Area was below the 90% target goal of the National Park Service. The large increase in number of additional species relative to time and collection effort (Figure 2) suggests that additional taxa remain to be found at the study area. Our potential species list (Appendix D) includes 148 taxa from adjacent areas of Montana and Wyoming that could occur within the study area. Forty-four of the potential species are considered “highly probable” because of their region-wide abundance and preference for common habitat types present in the area. If these 44 taxa were added to the Bighorn Canyon checklist, the current species count of 739 taxa would still represent 94.4% of the total flora, well over the 90% threshold. This percentage drops to 87.0% if an additional 66 species of moderate probability are included in the checklist and 83.3% if 38 low probability species are recognized. Since many of these low to moderate probability taxa are unlikely to be documented, the current checklist of Bighorn Canyon NRA is probably very close to the Park Service’s goal of 90% completeness. Table 5. Increase in Number of Plant Taxa Recorded at Bighorn Canyon National Recreation Area, 1924-2001

Years

Additional # of Species Documented

Cumulative # of Species Documented

1924 1 1 1954 1 2 1964 1 3 1966 1 4 1976 24 28 1977 1 29 1979 26 55 1980 23 78 1981 21 99 1982 35 134 1983 384 518 1984 47 565 1994 2 567 1996 1 568 1998 31 599 1999 29 628 2001 111 739

Page 15: Vascular Plant Species Checklist of Bighorn Canyon National

15

Figure 2. Increase in Number of Plant Taxa Recorded at Bighorn Canyon National Recreation Area, 1924-2001.

0

100

200

300

400

500

600

700

800

1924 1954 1964 1966 1976 1977 1979 1980 1981 1982 1983 1984 1994 1996 1998 1999 2001

Year

Num

ber o

f Tax

a

Cumulative Number of Taxa

Some additional geographic areas, habitats, and taxonomic groups may be targeted for future survey to ensure that Bighorn Canyon National Recreation Area meets the park service goal of a 90% complete flora. The high-elevation western boundary of the National Recreation Area on the midslopes of East Pryor Mountain was partially surveyed for rare species in 1998-99 but has not been revisited for more common plants. Some species that extend downslope from higher elevation subalpine habitat types may have been missed in our survey. Several sites along Yellowtail Reservoir are inaccessible by land and may still be undercollected. The sand dune deposits below the mouth of Big Bull Elk Creek were surveyed for rare species in 1998-99 but have not been revisited for more common plants. Disturbed habitats and wetlands yielded high numbers of additional taxa in 2001 and may still harbor undocumented species, especially in taxonomically difficult families (such as Asteraceae, Poaceae, and Cyperaceae). A number of cryptic plants, as well as species with early or very late phenologies, may still be undersampled. Some species that we consider “undocumented” may actually have valid vouchers in the collections of the New York Botanical Garden, but to confirm this will require either a visit to the facility or a specimen loan. Finally, we were not able to work at the Montana State University Herbarium, where there may be additional specimens from the study area collected by W.E. Booth. The highest priorities would be to review the list of already confirmed species and those with a high probability of occurring in the Bighorn Canyon National Recreation Area.

Page 16: Vascular Plant Species Checklist of Bighorn Canyon National

16

LITERATURE CITED Akashi, Y. 1988. Riparian vegetation dynamics along the Bighorn River, Wyoming. Master Thesis,

University of Wyoming, Laramie. Anonymous, 1968. Table V. A systematic list of the plant taxa occurring in the study areas of the

Big Horn [sic] Canyon National Recreation Area. Revised 1968. Copied from unknown report document.

Anonymous, 1975. A partial list of plants of the Bighorn Canyon Area. Xerox copy. Dorn, R. D. 1978. Great Basin vegetation in Carbon County, Montana. Madroño 25(2): 105-106. Dorn, R.D. 1984. Vascular Plants of Montana. Mountain West Publishing, Cheyenne, WY. Dorn, R.D. 2001. Vascular Plants of Wyoming, third edition. Mountain West Publishing,

Cheyenne, WY. Fertig, W. 1999a. Non-native plants of Wyoming. Castilleja (A Publication of the Wyoming

Native Plant Society) 18(2): 7-10. Fertig, W. 1999b. Protection status and checklist of the vascular plant flora of the Bighorn

Landscape. Report prepared for the Nature Conservancy Wyoming Field Office by the Wyoming Natural Diversity Database, Laramie. Heidel, B. and W. Fertig. 2000. Rare plants of Bighorn Canyon National Recreation Area. Report

prepared for the National Fish and Wildlife Foundation and Bighorn Canyon National Recreation Area by the Montana Natural Heritage Program and Wyoming Natural Diversity Database.

Kartesz, J.T. and C.A. Meacham. 1999. Synthesis of the North American Flora, version 1.0. North

Carolina Botanical Garden, Chapel Hill, NC. Knight, D.H., G. P. Jones, Y. Akashi, and R. W. Myers. 1987. Vegetation ecology in the Bighorn

Canyon National Recreation Area, Wyoming and Montana. Report prepared for the US National Park Service and University of Wyoming-National Park Service Research Center.

Lesica, P. and P. Achuff. 1992. Distribution of vascular plant species of special concern and limited

distribution in the Pryor Mountains Desert, Carbon County, Montana. Report to Bureau of Land Management. Montana Natural Heritage Program, Helena.

Lichvar, R.W., E. I. Collins, and D.H. Knight. 1984. Checklist of vascular plants for the Bighorn

Canyon National Recreation Area. Unpublished report for the Wyoming National Park Service Research Center and the Bighorn Canyon National Recreation Area.

Page 17: Vascular Plant Species Checklist of Bighorn Canyon National

17

Lichvar, R.W., E. I. Collins, and D.H. Knight. 1985. Checklist of vascular plants for the Bighorn Canyon National Recreaton Area, Wyoming and Montana. Great Basin Naturalist 45(4):734-746.

Love, J.D. and A.C. Christiansen. 1985. Geologic Map of Wyoming. US Geological Survey. Merrill, E.H., T.W. Kohley, M.E. Herdendorf, W.A. Reiners, K.L. Driese, R.W. Marrs, and S.H.

Anderson. 1996. The Wyoming Gap Analysis Project Final Report. Department of Zoology and Physiology, Department of Botany, and Wyoming Cooperative Fish and Wildlife Research Unit, University of Wyoming, Laramie, WY.

Nelson, B.E. and R.L. Hartman. 1984. Flora of the Big Horn Mountains, checklist. Report

prepared by the Rocky Mountain Herbarium, University of Wyoming, Laramie, WY. USDI National Park Service. 2000. Guidance for the design of sampling schemes for inventory and

monitoring of biological resources in National Parks. Appendix 1. In: The National Park Service’s Greater Yellowstone Area Vertebrate and Vascular Plant Inventory Workshop. Bozeman, MT.

Page 18: Vascular Plant Species Checklist of Bighorn Canyon National

18

App

endi

x A

. A

nnot

ated

Che

cklis

t of t

he V

ascu

lar P

lant

Flo

ra

of B

igho

rn C

anyo

n N

atio

nal R

ecre

atio

n A

rea.

Th

e fo

llow

ing

spec

ies l

ist i

s mod

ified

from

Lic

hvar

et a

l. (1

985)

and

Hei

del a

nd F

ertig

(200

0) a

nd is

bas

ed o

n ne

w fi

eld

surv

eys i

n 20

01 a

nd a

th

orou

gh re

-exa

min

atio

n of

spec

imen

s at t

he U

nive

rsity

of W

yom

ing’

s Roc

ky M

ount

ain

Her

bariu

m (R

M) a

nd B

igho

rn C

anyo

n N

atio

nal R

ecre

atio

n A

rea

Her

bariu

m (B

ICA

). N

omen

clat

ure

follo

ws K

arte

sz a

nd M

each

am (1

999)

for s

cien

tific

nam

es a

nd c

omm

on n

ames

. Pe

rtine

nt sy

nony

ms h

ave

been

incl

uded

for n

ames

that

diff

er fr

om th

ose

in th

e m

ost c

urre

nt st

atew

ide

flora

s of M

onta

na a

nd W

yom

ing

(Dor

n 19

84, 2

001)

. Th

e lis

t is

orga

nize

d al

phab

etic

ally

by

fam

ily, f

ollo

win

g th

e cl

assi

ficat

ion

adop

ted

by K

arte

sz a

nd M

each

am (1

999)

, with

alte

rnat

e fa

mily

nam

es in

clud

ed

whe

re th

ese

diff

er fr

om D

orn

(200

1).

For b

revi

ty, f

amily

nam

es d

o no

t inc

lude

the

stan

dard

suff

ix ‘a

ceae

’. G

loba

l ran

k is

a m

easu

re o

f ran

gew

ide

abun

danc

e of

eac

h ta

xon

usin

g a

scal

e fr

om 1

(rar

est)

to 5

(mos

t com

mon

) and

is d

eriv

ed fr

om th

e N

atur

eSer

ve w

ebsi

te (h

ttp://

natu

rese

rve.

abi.o

rg/).

N

ativ

e an

d ex

otic

(int

rodu

ced)

stat

us w

as d

eter

min

ed fr

om F

ertig

(199

9a) a

nd K

arte

sz a

nd M

each

am (1

999)

. Pr

esen

ce in

eac

h of

the

thre

e co

untie

s w

ithin

Big

horn

Can

yon

Nat

iona

l Rec

reat

ion

Are

a is

indi

cate

d by

an

“X”

for s

peci

es w

ith a

doc

umen

ted

herb

ariu

m v

ouch

er a

nd “

O”

if th

e re

cord

is

base

d on

a re

liabl

e ob

serv

atio

n. X

or O

indi

cate

s a n

ew c

ount

y di

strib

utio

n re

cord

with

in th

e st

udy

area

in 2

001.

A “

?” in

dica

tes s

ome

unce

rtain

ty

rega

rdin

g th

e id

entif

icat

ion

of a

spec

ies o

r its

con

firm

ed p

rese

nce

with

in a

cou

nty.

Sam

ple

vouc

her s

peci

men

s are

cite

d fo

r eac

h sp

ecie

s and

con

tain

in

form

atio

n on

the

colle

ctor

, col

lect

ion

num

ber,

and

repo

sito

ry (i

f kno

wn)

. Th

e fir

st y

ear c

olum

n in

dica

tes t

he y

ear o

f the

ear

liest

kno

wn

vouc

here

d he

rbar

ium

reco

rd o

r con

firm

ed o

bser

vatio

n w

ithin

Big

horn

Can

yon

Nat

iona

l Rec

reat

ion

Are

a. T

his y

ear d

oes n

ot n

eces

saril

y co

rres

pond

with

the

year

that

the

repr

esen

tativ

e vo

uche

r was

col

lect

ed.

Add

ition

al c

omm

ents

are

pro

vide

d as

nec

essa

ry to

cla

rify

our t

reat

men

t of a

par

ticul

ar ta

xon;

in

this

col

umn

the

BIC

A a

bbre

viat

ion

refe

rs to

eith

er th

e B

igho

rn C

anyo

n N

atio

nal R

ecre

atio

n A

rea

stud

y ar

ea o

r its

her

bariu

m fa

cilit

y.

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Acer

Ac

er g

labr

um v

ar.

glab

rum

R

ocky

Mou

ntai

n m

aple

G

5T5

N

O

X

Lich

var 5

685

(BIC

A)

1983

Acer

Ac

er n

egun

do v

ar.

inte

rius

Ash-

leaf

map

le

G5T

5 N

O

X C

ollin

s 57

0 (B

ICA)

19

83

Agav

Yu

cca

glau

ca

Soap

wee

d yu

cca

G5

N

X O

O

C

ollin

s 69

5 (R

M)

1983

Amar

anth

Am

aran

thus

blit

oide

s M

at a

mar

anth

G

? N

X

X O

C

ollin

s 85

4 (R

M)

1983

In

clud

es A

. alb

us

repo

rted

in K

nigh

t et

al.

1987

. Am

aran

th

Amar

anth

us re

trofle

xus

Red

-root

am

aran

th

G?

E O

X

H

eide

l 207

1 (R

M)

2001

Anac

ardi

R

hus

glab

ra

Smoo

th s

umac

G

5 N

O

X Li

chva

r 640

2 (R

M)

1983

Anac

ardi

R

hus

trilo

bata

[Rhu

s ar

omat

ica

var.

trilo

bata

]

Ill-s

cent

ed s

umac

G

5 N

O

O

X

Col

lins

552

(RM

) 19

82

Anac

ardi

To

xico

dend

ron

rydb

ergi

i W

este

rn p

oiso

n-iv

y G

5 N

O

X

X Li

chva

r 592

9 (R

M)

1983

Api

Beru

la e

rect

a C

ut-le

af w

ater

-pa

rsni

p G

4G5

N

O

X O

Li

chva

r 691

1 (R

M)

1983

Page 19: Vascular Plant Species Checklist of Bighorn Canyon National

19

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Api

Bupl

euru

m

amer

ican

um

Amer

ican

thor

ow-w

ax

G5

N

X

H

eide

l 199

6 (R

M)

2001

Ea

rlier

repo

rts

wer

e ba

sed

on

spec

imen

s co

llect

ed o

utsi

de

of B

CN

RA

Api

Cic

uta

mac

ulat

a va

r. an

gust

ifolia

Sp

otte

d w

ater

-he

mlo

ck

G5T

5 N

X

X

Hei

del 2

124

(RM

) 19

99

Api

Con

iose

linum

sc

opul

orum

R

ock

lova

ge

G4

N

X

Bo

oth

5463

9 (R

M)

1954

Fi

rst r

epor

t for

M

onta

na

Api

Con

ium

mac

ulat

um

Pois

on-h

emlo

ck

G4G

5 E

O

X O

Kn

ight

908

(RM

) 19

83

Ap

i C

ymop

teru

s ac

aulis

Pl

ains

spr

ing-

pars

ley

G5

N

X X

Li

chva

r 560

1 (R

M)

1983

Api

Her

acle

um m

axim

um

[H. s

phon

dyliu

m v

ar.

lana

tum

]

Amer

ican

cow

-pa

rsni

p G

5 N

O

O

X

Lich

var 6

140

(RM

) 19

83

Api

Lom

atiu

m d

isse

ctum

va

r. m

ultif

idum

C

hoco

late

-tips

G

3G4

T?

N

X Li

chva

r 639

8 (R

M)

1983

Api

Lom

atiu

m

foen

icul

aceu

m

Car

rot-l

eaf d

eser

t-pa

rsle

y G

5 N

O

X Li

chva

r 568

3 (R

M)

1983

Api

Lom

atiu

m o

rient

ale

Orie

ntal

des

ert-

pars

ley

G5

N

X

Li

chva

r 566

4 (R

M)

1983

Api

Mus

ineo

n di

varic

atum

Le

afy

wild

par

sley

G

5 N

X

X O

C

ollin

s 55

8 (R

M)

1983

Api

Mus

ineo

n va

gina

tum

Sh

eath

ed w

ild p

arsl

ey

G3?

N

X X

Hei

del 1

854

(MO

NTU

) 19

83

Api

Osm

orhi

za b

erte

roi

[O

. chi

lens

is]

Mou

ntai

n sw

eet-

cice

ly

G5

N

X C

ollin

s 80

7 (R

M)

1983

Api

Osm

orhi

za

depa

uper

ata

Blun

t-fru

it sw

eet-

cice

ly

G5

N

X H

eide

l 209

6 (B

ICA)

20

01

Earli

er re

port

(Lic

hvar

et a

l. 19

85) w

as b

ased

on

a

mis

iden

tific

atio

n Ap

i O

smor

hiza

long

isty

lis

Anis

eroo

t G

5 N

X

Lich

var 6

114

(RM

) 19

83

Ap

i Pe

rider

idia

gai

rdne

ri va

r. bo

real

is

[P. m

onta

na]

Com

mon

yam

pah

G5

N

O

X

Lich

var 6

898

(RM

) 19

83

Api

Pter

yxia

tere

bint

hina

va

r. al

biflo

ra

[C

ymop

teru

s te

rebi

nthi

nus

var.

albi

floru

s]

Turp

entin

e w

avew

ing

G5

T4T5

N

X

O

Fe

rtig

1838

5 (R

M)

1983

Api

Sani

cula

mar

iland

ica

Mar

ylan

d bl

ack

snak

eroo

t G

5 N

O

X Li

chva

r 616

2b (R

M)

1983

Apoc

yn

Apoc

ynum

an

dros

aem

ifoliu

m

Spre

adin

g do

gban

e G

5 N

X

Col

lins

816

(RM

) 19

83

Page 20: Vascular Plant Species Checklist of Bighorn Canyon National

20

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Apoc

yn

Apoc

ynum

can

nabi

num

In

dian

-hem

p G

5 N

O

X

O

Wol

f 88

(BIC

A)

1982

va

riety

not

de

term

ined

As

clep

iad

Ascl

epia

s in

carn

ata

Swam

p m

ilkw

eed

G5

N

X

N

elso

n 70

20 (R

M)

1980

Ascl

epia

d As

clep

ias

spec

iosa

Sh

owy

milk

wee

d G

5 N

O

X

X C

ollin

s 83

4 (R

M)

1982

Ascl

epia

d As

clep

ias

virid

iflor

a G

reen

com

et

milk

wee

d G

5 N

X

O

X H

eide

l 203

5 (R

M)

2001

Aste

r Ac

hille

a m

illefo

lium

C

omm

on y

arro

w

G5

N

O

O

X Jo

nes,

Dito

lla, a

nd

Mey

ers

s.n.

(BIC

A)

1984

Aste

r Ac

ropt

ilon

repe

ns

[Cen

taur

ea re

pens

] H

ardh

eads

[R

ussi

an k

napw

eed]

G

? E

X

O

Ferti

g 18

410

(RM

) 19

82

Nox

ious

wee

d in

M

T an

d W

Y As

ter

Agos

eris

gla

uca

var.

dasy

ceph

ala

Pale

goa

t-chi

cory

G

4G5

T?

N

X Li

chva

r 612

2 (B

ICA)

19

83

Aste

r Ag

oser

is g

lauc

a va

r. gl

auca

Pa

le g

oat-c

hico

ry

G4G

5 T?

N

X

Lich

var 5

572

(BIC

A)

1983

Aste

r Ag

oser

is g

lauc

a va

r. la

cini

ata

Pale

goa

t-chi

cory

G

4G5

T?

N

X

C

ollin

s 64

5 (R

M)

1983

D

uplic

ate

spec

imen

at B

ICA

was

mis

iden

tifie

d As

ter

Ambr

osia

aca

nthi

carp

a Fl

at-s

pine

bur

r-ra

gwee

d G

5 N

X

X

Hei

del 2

076

(RM

) 20

01

Aste

r Am

bros

ia a

rtem

isiif

olia

An

nual

ragw

eed

G5

N

X

N

elso

n 70

22 (R

M)

1980

Aste

r Am

bros

ia p

silo

stac

hya

Pere

nnia

l rag

wee

d G

5 N

O

X Li

chva

r 692

8 (R

M)

1983

Aste

r Am

bros

ia tr

ifida

G

reat

ragw

eed

G5

N

O

X

Col

lins

900

(RM

) 19

83

As

ter

Ante

nnar

ia m

icro

phyl

la

Smal

l-lea

f pus

syto

es

G4G

5 N

X X

Jone

s, D

itolla

, and

M

eyer

s s.

n. (B

ICA)

19

84

Incl

udes

m

isid

entif

ied

A.

cory

mbo

sa

repo

rted

by K

nigh

t et

al.

(198

7)

Aste

r An

tenn

aria

rose

a R

osy

puss

ytoe

s G

5 N

X

Col

lins

649

(RM

) 19

83

Spec

imen

or

igin

ally

m

isid

entif

ied

as A

. pa

rvifo

lia

Aste

r Ar

ctiu

m m

inus

Le

sser

bur

rdoc

k G

? E

X X

O

Col

lins

849

(BIC

A)

1982

N

oxio

us w

eed

in

WY

Aste

r Ar

nica

cor

difo

lia

Hea

rt-le

af

leop

ardb

ane

[H

eartl

eaf a

rnic

a]

G5

N

X Li

chva

r 617

4 (B

ICA)

19

83

Aste

r Ar

nica

fulg

ens

Shin

ing

leop

ardb

ane

G5

N

X

X Li

chva

r 593

7 (B

ICA)

19

83

Aste

r Ar

nica

latif

olia

D

affo

dil l

eopa

rdba

ne

G5

N

X

Li

chva

r 593

6 (B

ICA)

19

83

Aste

r Ar

nica

sor

oria

Tw

in le

opar

dban

e G

5 N

X

Lich

var 6

138

(RM

) 19

83

As

ter

Arte

mis

ia b

ienn

is v

ar.

bien

nis

Bien

nial

wor

mw

ood

G5T

5 N

X

O

O

Nel

son

5077

(RM

) 19

79

Page 21: Vascular Plant Species Checklist of Bighorn Canyon National

21

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Aste

r Ar

tem

isia

cam

pest

ris

var.

scou

leria

na

Paci

fic w

orm

woo

d G

5 T4

T5

N

X X

X Jo

nes,

Ditt

ola,

and

M

eyer

s s.

n. (B

ICA)

19

84

Aste

r Ar

tem

isia

can

a va

r. ca

na

Coa

ltow

n sa

gebr

ush

[Silv

er s

age]

G

5T5

N

X

X M

esse

rsch

mid

t 39

(BIC

A)

1981

Aste

r Ar

tem

isia

dra

cunc

ulus

D

rago

n w

orm

woo

d

[Tar

rago

n]

G5

N

X

X Li

chva

r 688

7 (R

M)

1983

Aste

r Ar

tem

isia

frig

ida

Prai

rie s

ageb

rush

[F

ringe

d sa

ge]

G5?

N

X

X X

Col

lins

891

(BIC

A)

1982

Aste

r Ar

tem

isia

ludo

vici

ana

var.

ludo

vici

ana

Whi

te s

ageb

rush

G

5T?

N

X

X Li

chva

r 689

2 (R

M)

1983

Aste

r Ar

tem

isia

mic

haux

iana

M

icha

ux's

wor

mw

ood

G4G

5 N

X

Hei

del 1

814

(BIC

A)

1999

Aste

r Ar

tem

isia

nov

a Bl

ack

sage

brus

h G

5 N

O

X

Jo

nes,

Ditt

ola,

and

M

eyer

s s.

n. (B

ICA)

19

84

Aste

r Ar

tem

isia

ped

atifi

da

Bird

sfoo

t sag

ebru

sh

G4

N

X

W

elp

7971

(RM

) 19

98

As

ter

Arte

mis

ia tr

iden

tata

va

r. tri

dent

ata

Big

sage

brus

h

[Bas

in b

ig s

ageb

rush

]

G5T

4T5

N

X X

Jo

nes,

Dito

lla, a

nd

Mey

ers

s.n.

(BIC

A)

1984

Aste

r Ar

tem

isia

trid

enta

ta

var.

vase

yana

Bi

g sa

gebr

ush

[M

ount

ain

big

sage

brus

h]

G5T

4T5

N

X

H

eide

l 206

6 (R

M)

2001

Aste

r Ar

tem

isia

trid

enta

ta

var.

wyo

min

gens

is

Big

sage

brus

h

[Wyo

min

g bi

g sa

gebr

ush]

G5T

5 N

X

O

O

Cla

usen

66-

237

(RM

) 19

66

Aste

r Ba

lsam

orhi

za in

cana

H

oary

bal

sam

root

G

4 N

X

Col

lins

653

(BIC

A)

1983

Aste

r Ba

lsam

orhi

za s

agitt

ata

Arro

w-le

af b

alsa

mro

ot

G4G

5 N

X X

Lich

var 5

654

(BIC

A)

1983

Aste

r Bi

dens

cer

nua

Nod

ding

bur

r-m

arig

old

G5

N

X

X N

elso

n 50

64 (R

M)

1979

Aste

r Bi

dens

trip

artit

a

[Bid

ens

com

osa]

Th

ree-

lobe

be

ggar

ticks

G

5 N

X

Nel

son

5067

(RM

) 19

79

Aste

r Br

icke

llia e

upat

orio

ides

va

r. co

rym

bulo

sa

Fals

e bo

nese

t G

5T?

N

X

X H

eide

l 213

9 (R

M)

1998

Aste

r Br

icke

llia g

rand

iflor

a Ta

ssel

-flow

er

bric

kellb

ush

G5

N

X

H

eide

l 211

9 (R

M)

2001

Ea

rlier

repo

rts

wer

e ba

sed

on

spec

imen

s co

llect

ed o

utsi

de

of B

CN

RA

Aste

r C

enta

urea

bie

bers

tein

ii [C

enta

urea

mac

ulos

a]

Spot

ted

knap

wee

d G

? E

X X

X Li

chva

r 641

8 (B

ICA)

19

83

Nox

ious

wee

d in

M

T an

d W

Y

Aste

r C

enta

urea

diff

usa

Whi

te k

napw

eed

[Bus

hy k

napw

eed;

D

iffus

e kn

apw

eed]

G?

E

X

Hei

del 2

159

(RM

) 20

01

Nox

ious

wee

d in

M

T an

d W

Y

Page 22: Vascular Plant Species Checklist of Bighorn Canyon National

22

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Aste

r C

haen

actis

dou

glas

ii va

r. do

ugla

sii

[C. d

ougl

asii

var.

mon

tana

]

Dus

ty-m

aide

n G

5 N

X

X

Hei

del 1

831

(RM

) 19

83

Aste

r C

hrys

otha

mnu

s lin

ifoliu

s Sp

ear-l

eaf

rabb

itbru

sh

G5

N

X X

Fe

rtig

1840

4 (R

M)

1983

Aste

r C

hrys

otha

mnu

s vi

scid

iflor

us v

ar.

visc

idifl

orus

Gre

en ra

bbitb

rush

G

5T5

N

X

X Li

chva

r 688

6 (B

ICA)

19

83

Aste

r C

icho

rium

inty

bus

Chi

cory

G

? E

X C

ollin

s 89

4 (R

M)

1983

Aste

r C

irsiu

m a

rven

se

Can

adia

n th

istle

G

5 E

X

X C

ollin

s 89

9 (B

ICA)

19

83

Nox

ious

wee

d in

M

T an

d W

Y As

ter

Cirs

ium

flod

man

ii Fl

odm

an's

this

tle

G5

N

X

O

Jobe

(s.n

.) (B

ICA)

19

80

As

ter

Cirs

ium

hoo

keria

num

Hoo

ker t

hist

le

G4

N

X

H

eide

l 170

7 (M

ON

TU)

1998

Ea

rlier

repo

rts

wer

e ba

sed

on

spec

imen

s co

llect

ed o

utsi

de

of B

ICA

Aste

r C

irsiu

m u

ndul

atum

W

avy-

leaf

this

tle

G5

N

X

X C

ollin

s 73

8 (B

ICA)

19

83

As

ter

Cirs

ium

vul

gare

Bu

ll th

istle

G

5 E

O

X

Lich

var 6

417

(BIC

A)

1983

Aste

r C

onyz

a ca

nade

nsis

C

anad

ian

hors

ewee

d G

5 N

X

X O

N

elso

n 70

37 (R

M)

1980

Aste

r C

repi

s ac

umin

ata

Long

-leaf

haw

k's-

bear

d G

5 N

X

Jone

s, D

itolla

, and

M

eyer

s (B

ICA)

19

84

Aste

r C

repi

s at

ribar

ba

Slen

der h

awk'

s-be

ard

G5

N

X Li

chva

r 617

5 (R

M)

1983

Aste

r C

repi

s in

term

edia

Li

mes

tone

haw

k's-

bear

d G

5 N

X X

Ferti

g 19

448

(RM

) 19

83

Aste

r C

repi

s m

odoc

ensi

s Si

skiy

ou h

awk'

s-be

ard

G4G

5 N

X

Lich

var 5

924

(BIC

A)

1983

Aste

r C

repi

s ru

ncin

ata

var.

glau

ca

Fidd

le-le

af h

awk'

s-be

ard

G5T

? N

X X

Hei

del 1

725

(MO

NTU

) 19

83

Aste

r D

ysso

dia

papp

osa

Fetid

-mar

igol

d G

5 N

X X

Jone

s, D

itolla

, and

M

eyer

s s.

n. (B

ICA)

19

84

Aste

r Ec

hina

cea

angu

stifo

lia

[E

. pal

lida

var.

angu

stifo

lia]

Blac

ksam

son

[N

arro

wle

af

cone

flow

er]

G4

N

X C

ollin

s 82

4 (B

ICA)

19

83

Aste

r Er

icam

eria

nau

seos

a va

r. gr

aveo

lens

[Chr

ysot

ham

nus

naus

eosu

s va

r. gr

aveo

lens

]

Rub

ber r

abbi

tbru

sh

G5T

5 N

X

X

Nel

son

7038

(RM

) 19

80

Page 23: Vascular Plant Species Checklist of Bighorn Canyon National

23

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Aste

r Er

icam

eria

nau

seos

a va

r. na

useo

sa

[C

hrys

otha

mnu

s na

useo

sus

var.

naus

eosu

s]

Rub

ber r

abbi

tbru

sh

G5T

5 N

X

X O

Jo

nes,

Dito

lla, a

nd

Mey

ers

s.n.

(BIC

A)

1984

Aste

r Er

iger

on a

lloco

tus

Big

Hor

n fle

aban

e G

3 N

X

X

Dor

n 26

25

(MO

NTU

) 19

76

Aste

r Er

iger

on c

aesp

itosu

s Tu

fted

fleab

ane

G5

N

X X

C

ollin

s 82

4 (B

ICA)

19

76

As

ter

Erig

eron

cor

ymbo

sus

Long

-leaf

flea

bane

G

5 N

X

Lich

var 6

357

(RM

) 19

83

As

ter

Erig

eron

gla

bellu

s va

r. gl

abel

lus

Stre

amsi

de fl

eaba

ne

G5

T4T5

N

X

Lich

var 6

117

(RM

) 19

83

Aste

r Er

iger

on o

chro

leuc

us

var.

ochr

oleu

cus

Buff

fleab

ane

G5T

3 N

X

X X

Lich

var 6

192

(RM

) 19

83

Aste

r Er

iger

on o

chro

leuc

us

var.

scrib

neri

Buff

fleab

ane

G5T

5 N

X

Lich

var 5

711

(RM

) 19

83

Earli

er re

ports

w

ere

base

d on

sp

ecim

ens

colle

cted

out

side

of

BC

NR

A As

ter

Erig

eron

pum

ilus

var.

pum

ilus

Shag

gy fl

eaba

ne

G5T

5 N

X

X

Lich

var 5

599

(RM

) 19

83

Aste

r Er

iger

on s

trigo

sus

Prai

rie fl

eaba

ne

G5

N

O

X

Lich

var 6

132

(RM

) 19

83

As

ter

Eupa

toriu

m m

acul

atum

va

r. br

uner

i

[Eup

ator

iade

lphu

s m

acul

atus

var

. bru

neri]

Spot

ted

Joe-

Pye-

wee

d G

5 T4

T5Q

N

X

X

Hei

del 2

132

(RM

) 19

83

Aste

r Eu

rybi

a co

nspi

cua

[Ast

er c

onsp

icuu

s]

East

ern

show

y w

ood-

aste

r G

5 N

O

2001

O

bser

ved

in

vege

tativ

e co

nditi

on b

y H

eide

l in

T6S

R31

E Se

c 19

As

ter

Eury

bia

glau

ca

[A

ster

gla

ucod

es,

Euce

phal

us g

lauc

us]

Gra

y w

ood-

aste

r G

4G5

N

X

H

eide

l 192

7 (M

ON

TU)

1999

Aste

r Eu

tham

ia o

ccid

enta

lis

[S

olid

ago

occi

dent

alis

] W

este

rn g

olde

ntop

G

5 N

X

Hei

del 2

155

(RM

) 20

01

Aste

r G

ailla

rdia

aris

tata

G

reat

bla

nket

-flow

er

G5

N

X

X H

eide

l 205

2 (B

ICA)

20

01

Prev

ious

ly

repo

rted,

but

no

vouc

her

spec

imen

s lo

cate

d fo

r cor

robo

ratio

n.

Aste

r G

naph

aliu

m p

alus

tre

Wes

tern

mar

sh

cudw

eed

G5

N

X

X N

elso

n 50

65 (R

M)

1979

Page 24: Vascular Plant Species Checklist of Bighorn Canyon National

24

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Aste

r G

rinde

lia s

quar

rosa

va

r. sq

uarro

sa

Cur

ly-c

up g

umw

eed

G5T

5 N

X

X X

Knig

ht K

-16

(BIC

A)

1983

Aste

r G

utie

rrezi

a sa

roth

rae

Kind

lingw

eed

[B

room

sna

kew

eed]

G

5 N

X

X X

Lich

var 6

925

(BIC

A)

1981

Aste

r H

elia

nthu

s an

nuus

C

omm

on s

unflo

wer

G

5 N

X

O

N

elso

n 50

79 (R

M)

1979

Aste

r H

elia

nthu

s nu

ttallii

N

utta

ll's s

unflo

wer

G

5 N

O

X

X N

elso

n 70

29 (R

M)

1980

Aste

r H

elia

nthu

s pa

ucifl

orus

va

r. su

brho

mbo

ideu

s

[H

. rig

idus

var

. s.]

Stiff

sun

flow

er

G5T

5?

N

X Li

chva

r 692

4 (R

M)

1983

Aste

r H

elia

nthu

s pe

tiola

ris

Prai

rie s

unflo

wer

G

4G5

N

X X

O

Col

lins

794

(RM

) 19

83

As

ter

Het

erot

heca

villo

sa v

ar.

villo

sa

Hai

ry fa

lse

gold

en-

aste

r G

5T?

N

X O

O

Jo

nes,

Dito

lla, a

nd

Mey

ers

s.n.

(BIC

A)

1984

Aste

r H

iera

cium

alb

iflor

um

Whi

te-fl

ower

ha

wkw

eed

G4G

5 N

X

Lich

var 6

390

(RM

) 19

83

Aste

r H

iera

cium

um

bella

tum

N

arro

w-le

af

haw

kwee

d G

5?

N

X H

eide

l 214

1 (R

M)

2001

Aste

r H

ymen

opap

pus

filifo

lius

var.

lute

us

Fine

-leaf

woo

lly-w

hite

G

5T?

N

X X

Li

chva

r 559

7 (R

M)

1983

O

rigin

ally

de

term

ined

as

var.

filifo

lius

Aste

r H

ymen

opap

pus

filifo

lius

var.

poly

ceph

alus

[H

. pol

ycep

halu

s]

Fine

-leaf

woo

lly-w

hite

G

5 T4

T5

N

X

Fe

rtig

1945

7 (R

M)

2001

Ea

rlier

repo

rts

wer

e ba

sed

on

mis

iden

tifed

sp

ecim

ens

Aste

r Iv

a ax

illaris

var

. ro

bust

ior

Dee

r-roo

t

[Pov

erty

-wee

d]

G5

N

X X

X Li

chva

r 599

2 (R

M)

1982

Aste

r Iv

a xa

nthi

folia

C

arel

essw

eed

[M

arsh

-eld

er]

G

5 N

X

Nel

son

5076

(RM

) 19

79

Aste

r La

ctuc

a lu

dovi

cian

a Lo

uisi

ana

lettu

ce

G4G

5 N

X

Lich

var 6

902

(RM

) 19

83

As

ter

Lact

uca

serri

ola

Pric

kly

lettu

ce

G?

E O

O

X

Col

lins

892

(RM

) 19

83

As

ter

Lact

uca

tata

rica

var.

pulc

hella

[L

. obl

ongi

folia

]

Rus

sian

blu

e le

ttuce

G

5T5

N

X

X C

ollin

s 82

9 (R

M)

1982

Aste

r Le

ucan

them

um v

ulga

re

[Chr

ysan

them

um

leuc

anth

emum

]

Ox-

eye

dais

y G

? E

O

19

99

Nox

ious

wee

d in

M

T an

d W

Y.

Obs

erve

d at

T7S

R

31E

Sec.

7 b

y H

eide

l As

ter

Liat

ris p

unct

ata

Dot

ted

gayf

eath

er

G5

N

X X

X Li

chva

r 636

1 (R

M)

1983

Aste

r Lo

gfia

arv

ensi

s

[Fila

go a

rven

sis]

Fi

eld

cotto

n-ro

se

G?

E

X

Lich

var 6

128

(RM

) 19

83

Fals

ely

repo

rted

as n

ativ

e &

rare

by

Lic

hvar

et a

l. (1

985)

As

ter

Lygo

desm

ia ju

ncea

R

ush

skel

eton

-pla

nt

G5

N

X X

X Li

chva

r 613

3 (R

M)

1983

Page 25: Vascular Plant Species Checklist of Bighorn Canyon National

25

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Aste

r M

acha

eran

ther

a ca

nesc

ens

var.

cane

scen

s

Hoa

ry ta

nsy-

aste

r G

5T5

N

X X

O

Jone

s, D

ittol

a, a

nd

Mey

ers

s.n.

(BIC

A)

1984

Aste

r M

acha

eran

ther

a gr

inde

lioid

es

[Hap

lopa

ppus

nut

tallii

]

Ray

less

tans

y-as

ter

G5

N

X O

X

Lich

var 5

998

(RM

) 19

83

Aste

r M

acha

eran

ther

a ta

nace

tifol

ia

Taho

ka-d

aisy

[Tan

sy-a

ster

] G

5 N

X

O

Li

chva

r 603

1 (R

M)

1982

Aste

r M

alac

othr

ix s

onch

oide

s Ye

llow

-sau

cers

G

5 N

X

Lich

var 6

047

(RM

) 19

83

As

ter

Mic

rose

ris n

utan

s N

oddi

ng s

ilver

puffs

G

5 N

X

Col

lins

651

(RM

) 19

83

As

ter

Not

hoca

lais

cus

pida

ta

Wav

y-le

af p

rairi

e-da

ndel

ion

G5

N

O

X

Lich

var 5

709

(RM

) 19

83

Aste

r O

ligon

euro

n rig

idum

va

r. hu

mile

[Sol

idag

o rig

ida

var.

hum

ilis]

Har

d-le

af fl

at-to

p go

lden

rod

G5T

5 N

X

Lich

var 6

904

(BIC

A)

1983

Aste

r Pa

cker

a ca

na

[Sen

ecio

can

us]

Silv

er-w

oolly

gr

ound

sel

G5

N

X X

X Li

chva

r 590

7 (R

M)

1981

Aste

r Pa

cker

a pa

uper

cula

[S

enec

io p

aupe

rcul

us]

Bals

am g

ound

sel

G4G

5 N

X

Lich

var 6

105

(RM

) 19

83

Aste

r Pa

cker

a pl

atte

nsis

[Sen

ecio

pla

ttens

is]

Prai

rie g

roun

dsel

G

4 N

X

Hei

del 1

853

(MO

NTU

) 19

99

Aste

r Pa

cker

a ps

euda

urea

va

r. ps

euda

urea

[S

enec

io p

seud

aure

us

var.

pseu

daur

eus]

Stre

amba

nk

grou

ndse

l G

5T?

N

O

X

Hei

del 1

922

(MO

NTU

) 19

99

Aste

r Pa

cker

a st

rept

anth

ifolia

va

r. ru

bric

aulis

[Sen

ecio

st

rept

anth

ifoliu

s va

r. ru

bric

aulis

]

Roc

ky M

ount

ain

grou

ndse

l G

5T?

N

X H

eide

l 200

1 (R

M)

1998

Pr

evio

usly

re

porte

d, b

ut n

o vo

uche

r sp

ecim

ens

loca

ted

for c

orro

bora

tion.

As

ter

Picr

otha

mnu

s de

serto

rum

[Arte

mis

ia s

pine

scen

s]

Buds

age

G5

N

X

Li

chva

r 560

7 (R

M)

1983

Aste

r Pl

atys

chku

hria

in

tegr

ifolia

Ba

sin-

dais

y G

5 N

X

X

Lich

var 6

032

(RM

) 19

83

Aste

r R

atib

ida

colu

mni

fera

R

ed-s

pike

Mex

ican

-ha

t G

5 N

X

Lich

var 6

927

(RM

) 19

82

Aste

r R

udbe

ckia

laci

niat

a va

r. am

pla

Gre

en-h

ead

cone

flow

er

G5T

? N

X

Col

lins

815

(BIC

A)

1980

Aste

r Se

neci

o er

emop

hilu

s va

r. er

emop

hilu

s

Des

ert r

agw

ort

G5T

5 N

X

Lich

var 6

383

(RM

) 19

83

Page 26: Vascular Plant Species Checklist of Bighorn Canyon National

26

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Aste

r Se

neci

o in

tege

rrim

us

var.

exal

tatu

s La

mb-

tong

ue ra

gwor

t G

5T5

N

X Li

chva

r 638

3 (R

M)

1983

Aste

r So

lidag

o ca

nade

nsis

va

r. sa

lebr

osa

Can

adia

n go

lden

rod

G5T

5?

N

X

X Kn

ight

K-3

4 (R

M)

1983

Aste

r So

lidag

o gi

gant

ea

Late

gol

denr

od

G5

N

X

X N

elso

n 70

34 (R

M)

1980

Aste

r So

lidag

o m

isso

urie

nsis

va

r. m

isso

urie

nsis

M

isso

uri g

olde

nrod

G

5T5

N

O

X

Lich

var 6

354

(RM

) 19

84

Aste

r So

lidag

o ne

mor

alis

var

. lo

ngip

etio

lata

G

ray

gold

enro

d G

5T5

N

X H

eide

l 214

6 (R

M)

2001

Aste

r So

nchu

s ar

vens

is s

sp.

ulig

inos

us

[S. u

ligin

osus

]

Fiel

d so

w-th

istle

G

? E

X X

W

olf 1

42 (B

ICA)

19

82

Aste

r So

nchu

s as

per

Spin

y-le

af s

ow-th

istle

G

? E

X

X Li

chva

r 690

3 (B

ICA)

19

83

Aste

r So

nchu

s ol

erac

eus

Com

mon

sow

-this

tle

G?

E X

Lich

var 6

047

(RM

) 19

83

As

ter

Spha

erom

eria

cap

itata

C

lust

er-h

ead

chic

ken-

sage

G

3 N

X

X

Lich

var 5

627

(RM

) 19

76

Aste

r St

enot

us a

rmer

ioid

es

[Hap

lopa

ppus

ar

mer

ioid

es]

Thrif

t moc

k-go

lden

wee

d G

4G5

N

X X

Li

chva

r 559

6 (R

M)

1983

Aste

r St

epha

nom

eria

min

or

[S

. ten

uifo

lia]

Less

er w

ire-le

ttuce

G

5 N

X

Hei

del 1

928

(BIC

A)

1999

Aste

r St

epha

nom

eria

ru

ncin

ata

Flow

erin

g-st

raw

G

5 N

X

O

O

Jone

s, D

itolla

, and

M

eyer

s s.

n. (B

ICA)

19

84

Aste

r Sy

mph

yotri

chum

ci

liatu

m

[A

ster

bra

chya

ctis

]

Alka

li Am

eric

an-a

ster

G

5 N

X

Hei

del 1

929

(MO

NTU

) 19

98

Aste

r Sy

mph

yotri

chum

ea

toni

i

[Ast

er b

ract

eola

tus,

A.

eato

nii]

Brac

ted

Amer

ican

-as

ter

G5

N

X C

ollin

s 87

4 (B

ICA)

19

83

Aste

r Sy

mph

yotri

chum

er

icoi

des

var.

pans

um

[A

ster

eric

oide

s va

r. pa

nsus

]

Whi

te h

eath

Am

eric

an-a

ster

G

5T5

N

X X

X N

elso

n 70

30 (R

M)

1980

Aste

r Sy

mph

yotri

chum

fro

ndos

um

[A

ster

fron

dosu

s]

Leaf

y Am

eric

an-a

ster

G

4 N

X

Hei

del 2

138

(RM

) 20

01

Aste

r Sy

mph

yotri

chum

la

nceo

latu

m v

ar.

hesp

eriu

m

[A

ster

lanc

eola

tus

var.

hesp

eriu

s, A

. h.]

Whi

te p

anic

led

Amer

ican

-ast

er

G5T

5?

N

X H

eide

l 214

4 (R

M)

1999

Pr

evio

usly

re

porte

d, b

ut n

o vo

uche

r sp

ecim

ens

loca

ted

for c

orro

bora

tion.

Page 27: Vascular Plant Species Checklist of Bighorn Canyon National

27

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Aste

r Ta

nace

tum

vul

gare

C

omm

on ta

nsy

G?

E

X

Hei

del 1

926

(MO

NTU

) 19

99

Aste

r Ta

raxa

cum

laev

igat

um

Red

-see

d da

ndel

ion

G?

E

X

Lich

var 5

633

(BIC

A)

1983

Aste

r Ta

raxa

cum

offi

cina

le

Com

mon

dan

delio

n G

5 E

X

O

Ferti

g 19

442

(RM

) 20

01

As

ter

Tetra

dym

ia c

anes

cens

Sp

inel

ess

hors

ebru

sh

G5

N

X X

Jo

nes,

Dito

lla, a

nd

Mey

ers

s.n.

(BIC

A)

1984

Aste

r Te

trane

uris

aca

ulis

var

. ac

aulis

[H

ymen

oxys

aca

ulis

va

r. ac

aulis

]

Stem

less

four

-ner

ve

dais

y G

5T5

N

X X

X W

olf 1

05 (B

ICA)

19

82

Aste

r Te

trane

uris

torre

yana

[Hym

enox

ys to

rreya

na]

Torre

y's fo

ur-n

erve

da

isy

G4

N

X

Li

chva

r 562

8 (R

M)

1983

Aste

r To

wns

endi

a ho

oker

i H

ooke

r's T

owns

end-

dais

y G

5 N

O

X

Li

chva

r 606

9 (B

ICA)

19

83

Aste

r To

wns

endi

a in

cana

H

oary

Tow

nsen

d-da

isy

G5

N

X X

D

orn

2628

(RM

) 19

76

Aste

r To

wns

endi

a pa

rryi

Parry

's T

owns

end-

dais

y G

4?

N

X

Li

chva

r 596

0 (R

M)

1983

Aste

r To

wns

endi

a sp

athu

lata

Sw

ord

Tow

nsen

d-da

isy

G3

N

O

X X

Lich

var 5

631

(RM

) 19

83

Aste

r Tr

agop

ogon

dub

ius

Mea

dow

goa

t's-b

eard

G

? E

X X

X M

esse

rsch

mid

t s.n

. BI

CA

1981

Aste

r W

yeth

ia s

cabr

a Ba

dlan

d m

ule'

s-ea

rs

G4

N

X O

Dor

n 26

29 (R

M)

1976

Aste

r Xa

nthi

um s

trum

ariu

m

var.

cana

dens

e R

ough

coc

kleb

urr

G5T

5 N

X

X

Col

lins

915

(RM

) 19

79

Aste

r Xy

lorh

iza

glab

riusc

ula

Smoo

th w

oody

-ast

er

G4

N

X X

Jo

nes,

Dito

lla, a

nd

Mey

ers

s.n.

(BIC

A)

1984

Berb

erid

M

ahon

ia re

pens

C

reep

ing

Ore

gon-

grap

e G

5 N

X

Lich

var 5

679

(RM

) 19

83

Betu

l Be

tula

occ

iden

talis

W

ater

birc

h G

4G5

N

X X

X Kn

ight

68

(RM

) 19

82

Bo

ragi

n As

peru

go p

rocu

mbe

ns

Ger

man

mad

wor

t G

? E

X

X Li

chva

r 564

8 (B

ICA)

19

83

Bora

gin

Cry

ptan

tha

ambi

gua

Basi

n ca

t's-e

ye

G4

N

X

D

orn

2623

(RM

) 19

76

Bo

ragi

n C

rypt

anth

a ca

na

Mou

ntai

n ca

t's-e

ye

G5

N

X X

Li

chva

r 600

0 (R

M)

1976

In

clud

es re

ports

of

C. c

aesp

itosa

from

Kn

ight

et a

l. 19

97

Bora

gin

Cry

ptan

tha

celo

sioi

des

Butte

cand

le

G5

N

X X

X Li

chva

r 566

9 (R

M)

1982

Bora

gin

Cry

ptan

tha

flavo

cula

ta

Rou

gh-s

eed

cat's

-eye

G

5 N

X

X

Lich

var 5

591

(RM

) 19

76

Bo

ragi

n C

rypt

anth

a ke

lsey

ana

Kels

ey's

cat

's-e

ye

G4

N

X O

Col

lins

710

(RM

) 19

83

Col

lins

710

or

igin

ally

de

term

ined

as

C.

wat

soni

i

Page 28: Vascular Plant Species Checklist of Bighorn Canyon National

28

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Bora

gin

Cyn

oglo

ssum

offi

cina

le

Gyp

sy-fl

ower

[Com

mon

hou

nd's-

tong

ue]

G?

E O

O

X

Lich

var 6

118

(RM

) 19

83

Nox

ious

wee

d in

M

T an

d W

Y

Bora

gin

Eritr

ichi

um h

owar

dii

How

ard'

s al

pine

fo

rget

-me-

not

G4

N

O

19

98

Obs

erve

d ab

ove

Layo

ut C

reek

(T8S

R

28E

S33)

by

Hei

del

Bora

gin

Hac

kelia

def

lexa

var

. am

eric

ana

Nod

ding

stic

ksee

d G

5TU

N

X

Lich

var 6

368

(BIC

A)

1983

Bora

gin

Hac

kelia

flor

ibun

da

Man

y-flo

wer

st

icks

eed

G5?

N

X

Lich

var 6

096

(BIC

A)

1983

Bora

gin

Lapp

ula

occi

dent

alis

va

r. cu

pula

ta

[L. r

edow

skii

var.

cupu

lata

, L. t

exan

a]

Flat

-spi

ne s

heep

burr

G5T

5 N

X

X X

Lich

var 6

030

(RM

) 19

83

Bora

gin

Lapp

ula

occi

dent

alis

va

r. oc

cide

ntal

is

[L. r

edow

skii

var.

redo

wsk

ii]

Flat

-spi

ne s

heep

burr

G5T

5 N

X

X

Col

lins

722

(RM

) 19

83

Bora

gin

Lapp

ula

squa

rrosa

var

. er

ecta

Br

istly

she

epbu

rr G

? E

X X

Fe

rtig

1942

8 (R

M)

1984

Bora

gin

Lith

ospe

rmum

inci

sum

Fr

inge

d gr

omw

ell

G5

N

X X

X Li

chva

r 556

3 (R

M)

1983

Bora

gin

Lith

ospe

rmum

rude

rale

C

olum

bian

puc

coon

G

5 N

O

X

O

Lich

var 5

653

(RM

) 19

83

Bo

ragi

n M

erte

nsia

cilia

ta

Tall

fring

e bl

uebe

lls

G5

N

X Li

chva

r 616

4 (R

M)

1983

Bora

gin

Mer

tens

ia o

blon

gifo

lia

Lang

uid-

lady

[Lea

fy b

lueb

ells

] G

5 N

X

Lich

var 5

573

(RM

) 19

83

Bora

gin

Ono

smod

ium

mol

le

var.

occi

dent

ale

Soft-

hair

mar

bles

eed

G4G

5 T4

? N

O

X Li

chva

r 613

9 (B

ICA)

19

83

incl

udes

var

. mol

le

fals

ely

repo

rted

by

Lich

var e

t al.

(198

5)

Bora

gin

Tiqu

ilia n

utta

llii

[C

olde

nia

nutta

llii]

Ros

ette

crin

klem

at

G5

N

X

D

orn

2950

(RM

) 19

77

Bras

sic

Alys

sum

aly

ssoi

des

Pale

mad

wor

t G

? E

X Li

chva

r 568

1 (R

M)

1983

Bras

sic

Alys

sum

des

erto

rum

D

eser

t mad

wor

t G

? E

X X

O

Ferti

g 19

419

(RM

) 20

01

Br

assi

c Ar

abid

opsi

s th

alia

na

Thal

ecre

ss

G?

E

X

Hei

del 2

050

(RM

) 20

01

Br

assi

c Ar

abis

dem

issa

var

. la

ngui

da

[Boe

cher

a de

mis

sa v

ar.

lang

uida

]

Nod

ding

rock

cres

s G

5T4

N

X X

H

eide

l 199

5 (R

M)

1983

Bras

sic

Arab

is fe

ndle

ri va

r. sp

atifo

lia

[B

oech

era

fend

leri

var.

spat

ifolia

]

Fend

ler's

rock

cres

s G

5T4

N

X

X

Ferti

g 18

387

(RM

) 19

98

Page 29: Vascular Plant Species Checklist of Bighorn Canyon National

29

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Bras

sic

Arab

is g

labr

a To

wer

-mus

tard

G

5 N

X X

Hei

del 2

040

(BIC

A)

1983

Bras

sic

Arab

is h

olbo

ellii

var.

collin

sii

[Boe

cher

a ho

lboe

llii

var.

collin

sii]

Hol

boel

l's ro

ckcr

ess

G5T

5 N

O

X Li

chva

r 568

7 (R

M)

1983

Li

chva

r 568

7 w

as

orig

inal

ly

mis

iden

tifie

d as

va

r. re

trofra

cta

Bras

sic

Arab

is h

olbo

ellii

var.

pend

uloc

arpa

[A

. pen

dulo

carp

a va

r. pe

ndul

ocar

pa,

Boec

hera

exi

lis]

Hol

boel

l's ro

ckcr

ess

G5T

5?

N

X

H

eide

l 182

7 (M

ON

TU)

1999

Bras

sic

Arab

is h

olbo

ellii

var.

pine

toru

m

[Boe

cher

a ho

lboe

llii

var.

pine

toru

m]

Hol

boel

l's ro

ckcr

ess

G5T

5?

N

X

Li

chva

r 592

3 (R

M)

1983

Li

chva

r 592

3 w

as

orig

inal

ly

mis

iden

tifie

d as

va

r. re

trofra

cta

Bras

sic

Arab

is h

olbo

ellii

var.

retro

fract

a

[A

. hol

boel

lii va

r. se

cund

a, B

oech

era

holb

oellii

var

. sec

unda

]

Hol

boel

l's ro

ckcr

ess

G5T

5 N

X X

Lich

var 5

693

(RM

) 19

83

Dup

licat

e sp

ecim

en a

t BIC

A m

ay b

e m

isid

entif

ied

Bras

sic

Arab

is n

utta

llii

[B

oech

era

nutta

llii]

Nut

tall's

rock

cres

s G

5 N

X X

Lich

var 5

705

(RM

) 19

83

Bras

sic

Arab

is s

pars

iflor

a va

r su

bvillo

sa

[Boe

cher

a sp

arsi

flora

]

Eleg

ant r

ockc

ress

G

5T?

N

X

D

orn

2556

(p

erso

nal

herb

ariu

m)

1976

Bras

sic

Barb

area

orth

ocer

as

Amer

ican

yel

low

-ro

cket

G

? N

X

Lich

var 6

161

(BIC

A)

1983

Bras

sic

Cam

elin

a m

icro

carp

a Li

ttle-

pod

fals

e fla

x G

? E

X

C

ollin

s 58

7 (R

M)

1983

Bras

sic

Cap

sella

bur

sa-p

asto

ris

Shep

herd

's-p

urse

G

? E

X

X Li

chva

r 571

8 (R

M)

1983

Bras

sic

Car

daria

pub

esce

ns

Glo

be-p

od

hoar

ycre

ss

G?

E X

Lich

var 6

052

(RM

) 19

79

Nox

ious

wee

d in

W

Y Br

assi

c C

horis

pora

tene

lla

Cro

ssflo

wer

G

? E

X

Li

chva

r 564

8 (R

M)

1983

Bras

sic

Des

cura

inia

inca

na v

ar.

inca

na

Mou

ntai

n ta

nsy-

mus

tard

G

5T5?

N

X

X

Ferti

g 19

425

(RM

) 19

84

Bras

sic

Des

cura

inia

pin

nata

va

r. ha

licto

rum

[D. p

inna

ta v

ar.

osm

iaru

m]

Wes

tern

tans

y-m

usta

rd

G5T

? N

X

O

Fe

rtig

1943

1 (R

M)

1983

Bras

sic

Des

cura

inia

pin

nata

va

r. in

term

edia

W

este

rn ta

nsy-

mus

tard

G

5T?

N

X

Fe

rtig

1945

1 (R

M)

2001

Bras

sic

Des

cura

inia

sop

hia

Her

b-so

phia

[Flix

wee

d]

G?

E X

O

C

ollin

s 55

1 (R

M)

1983

Bras

sic

Dra

ba a

lber

tina

[D

. ste

nolo

ba]

Slen

der w

hitlo

w-g

rass

G

4 N

X X

Lich

var 5

677

(RM

) 19

83

Page 30: Vascular Plant Species Checklist of Bighorn Canyon National

30

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Bras

sic

Dra

ba o

ligos

perm

a va

r. ol

igos

perm

a Fe

w-s

eed

whi

tlow

-gr

ass

G5

N

X

Li

chva

r 596

6 (R

M)

1983

Bras

sic

Dra

ba p

raea

lta

Tall

whi

tlow

-gra

ss

G5

N

X

O

Col

lins

665

(RM

) 19

83

Br

assi

c D

raba

rept

ans

Car

olin

a w

hitlo

w-

gras

s G

5 N

X

X

Lic

hvar

593

1 (R

M)

1983

Bras

sic

Erys

imum

cap

itatu

m

var.

purs

hii

[E

. as

peru

m, E

. ca

pita

tum

var

. ar

gillo

sum

]

Sand

-dun

e w

allfl

ower

G

5T?

N

X

X H

eide

l 201

9 (B

ICA)

19

83

Bras

sic

Erys

imum

ch

eira

ntho

ides

var

. al

tum

Wor

m-s

eed

wal

lflow

er

G5

N

X Li

chva

r 611

3 (R

M)

1983

Bras

sic

Erys

imum

in

cons

picu

um

Shy

wal

lflow

er

G4G

5 N

X?

O

Lich

var 6

085

(RM

) 19

83

Vouc

her a

t BIC

A m

ay b

e m

isid

entif

ied

Bras

sic

Hes

peris

mat

rona

lis

Mot

her-o

f-the

-Ev

enin

g

[D

ame'

s ro

cket

]

G4G

5 E

X Li

chva

r 571

4 (R

M)

1983

Bras

sic

Lepi

dium

den

siflo

rum

va

r. m

acro

carp

um

Min

er's

pep

perw

ort

G5T

5 N

X

X

Col

lins

691

(RM

) 19

83

Bras

sic

Lepi

dium

per

folia

tum

C

lasp

ing

pepp

erw

ort

G?

E

X

Lich

var 5

719

(RM

) 19

83

Br

assi

c Le

sque

rella

alp

ina

Al

pine

bla

dder

pod

G4

N

X X

X Li

chva

r 646

3 (R

M)

1983

In

clud

es v

ars.

al

pina

and

sp

athu

lata

Br

assi

c Le

sque

rella

lesi

cii

Pryo

r Mou

ntai

ns

blad

derp

od

G1

N

X

H

eide

l 202

2 (R

M)

1998

Bras

sic

Lesq

uere

lla lu

dovi

cian

a Lo

uisi

ana

blad

derp

od

G5

N

X

X Li

chva

r 618

7 (R

M)

1983

Bras

sic

Mal

colm

ia a

frica

na

Afric

an a

dder

's-m

outh

G

? E

X X

D

orn

2540

(RM

) 19

76

Br

assi

c Ph

ysar

ia a

cutif

olia

Sh

arp-

leaf

twin

pod

G5

N

O

X

Lich

var 5

872

(RM

) 19

83

Br

assi

c Ph

ysar

ia d

idym

ocar

pa

Com

mon

twin

pod

G5

N

X X

X Li

chva

r 563

6 (B

ICA)

19

83

Som

e no

n-fru

iting

sp

ecim

ens

may

ac

tual

ly b

e P.

ac

utifo

lia

Bras

sic

Ror

ippa

cal

ycin

a Pe

rsis

tent

-sep

al

yello

wcr

ess

G3

N

X

Li

chva

r 601

7 (R

M)

1983

Bras

sic

Ror

ippa

cur

vipe

s va

r. cu

rvip

es

Blun

t-lea

f yel

low

cres

s G

5T?

N

X X

Li

chva

r 601

3 (R

M)

1983

Bras

sic

Ror

ippa

nas

turti

um-

aqua

ticum

[Nas

turti

um o

ffici

nale

]

Wat

ercr

ess

G5

E O

X

X Li

chva

r 588

0 (B

ICA)

19

83

Bras

sic

Ror

ippa

pal

ustri

s va

r. fe

rnal

dian

a Bo

g ye

llow

cres

s G

5T5

N

X H

eide

l 215

3 20

01

Page 31: Vascular Plant Species Checklist of Bighorn Canyon National

31

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Bras

sic

Ror

ippa

pal

ustri

s va

r. hi

spid

a Bo

g ye

llow

cres

s G

5T5

N

X

Fe

rtig

1841

4 (R

M)

1998

Bras

sic

Ror

ippa

sin

uata

Sp

read

ing

yello

wcr

ess

G5

N

X X

Li

chva

r 605

0 (R

M)

1983

Bras

sic

Scho

enoc

ram

be

linifo

lia

[Sis

ymbr

ium

lini

foliu

m]

Salm

on R

iver

pla

ins-

mus

tard

G

5 N

X

X

Lich

var 5

606

(BIC

A)

1983

Bras

sic

Sisy

mbr

ium

alti

ssim

um

Tall

hedg

e-m

usta

rd

[Tum

blem

usta

rd]

G?

E X

X X

Lich

var 5

994

(RM

) 19

82

Bras

sic

Sisy

mbr

ium

loes

elii

Fals

e Lo

ndon

rock

et

[L

oese

l's

tum

blem

usta

rd]

G?

E

X

Hei

del 1

971

(RM

) 20

01

Bras

sic

Stan

leya

pin

nata

G

olde

n pr

ince

's-

plum

e G

4G5

N

O

X

Wol

f 86

(BIC

A)

1982

Bras

sic

Stan

leya

tom

ento

sa

var.

tom

ento

sa

Woo

lly p

rince

's-p

lum

e G

4T3

N

X X

X Li

chva

r 619

6 (R

M)

1983

Bras

sic

Stre

ptan

thel

la

long

irost

ris

Long

-bea

k fid

dle-

mus

tard

G

5 N

X

Lich

var 6

010

(BIC

A)

1983

Bras

sic

Thla

spi a

rven

se

Fiel

d pe

nnyc

ress

G

? E

X

Jo

nes,

Dito

lla, a

nd

Mye

rs s

.n. R

M

1984

Cac

t O

punt

ia p

olya

cant

ha

var.

poly

acan

tha

Hai

r-spi

ne p

rickl

y-pe

ar

[

Plai

ns

pric

kly-

pear

]

G5T

5 N

X

X

Jone

s, D

itolla

, and

M

yers

s.n

. (R

M)

1983

Cac

t Pe

dioc

actu

s si

mps

onii

Snow

ball

cact

us

G4

N

X C

ollin

s 64

7 (B

ICA)

19

83

C

ampa

nul

Cam

panu

la ro

tund

ifolia

Bl

uebe

ll-of

-Sco

tland

[Har

ebel

l] G

5 N

O

X

X Li

chva

r 607

4 (B

ICA)

19

83

Can

nab

Hum

ulus

lupu

lus

var.

neom

exic

anus

C

omm

on h

op

G5

T4T5

N

X

Hei

del 2

090

(RM

) 20

01

Prev

ious

ly

repo

rted,

but

no

vouc

her

spec

imen

s lo

cate

d fo

r cor

robo

ratio

n.

Cap

par

Cle

ome

lute

a Ye

llow

spi

der-f

low

er

G5

N

X X

Li

chva

r 604

3 (R

M)

1976

Cap

par

Cle

ome

serru

lata

R

ocky

Mou

ntai

n be

epla

nt

G5

N

X M

orst

ad s

.n. (

BIC

A)

2001

Ea

rlier

repo

rt by

Li

chva

r et a

l. (1

985)

was

bas

ed

on a

mis

iden

tifie

d sp

ecim

en

Cap

par

Pola

nisi

a do

deca

ndra

va

r. tra

chys

perm

a R

ed-w

hisk

er

clam

myw

eed

G5T

? N

X

O

Li

chva

r 602

8 (R

M)

1983

Cap

rifol

i Lo

nice

ra ta

taric

a Tw

insi

ster

s

[Tat

aria

n ho

neys

uckl

e]

G?

E

X

Hei

del 2

108

(RM

) 19

99

Page 32: Vascular Plant Species Checklist of Bighorn Canyon National

32

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Cap

rifol

i Sa

mbu

cus

nigr

a ss

p.

caer

ulea

[S

. cer

ulea

]

Blac

k el

der

G5?

N

O

X Li

chva

r 613

6 (R

M)

1983

Cap

rifol

i Sy

mph

oric

arpo

s al

bus

var.

albu

s C

omm

on s

now

berry

G

5T4

N

X

X C

ollin

s 76

4 (R

M)

1983

Cap

rifol

i Sy

mph

oric

arpo

s oc

cide

ntal

is

Wes

tern

sno

wbe

rry

G5

N

X

O

Lich

var 6

415

(RM

) 19

83

Cap

rifol

i Sy

mph

oric

arpo

s or

eoph

ilus

var.

utah

ensi

s

Mou

ntai

n sn

owbe

rry

G5T

5?

N

X

C

ollin

s 72

8 (B

ICA)

19

83

Car

yoph

yll

Aren

aria

con

gest

a va

r. co

nges

ta

[E

rem

ogen

e co

nges

ta

var.

c.]

Ball-

head

san

dwor

t G

5T5?

N

X

Lich

var 5

927

(RM

) 19

83

Car

yoph

yll

Aren

aria

hoo

keri

var.

hook

eri

[E

rem

ogen

e ho

oker

i va

r. h.

]

Hoo

ker's

san

dwor

t G

5T5?

N

X

X

Lich

var 5

612

(RM

) 19

83

Car

yoph

yll

Aren

aria

ser

pylli

folia

Th

yme-

leaf

san

dwor

t G

? E

O

X

Hei

del 1

861

(MO

NTU

) 19

99

Car

yoph

yll

Cer

astiu

m a

rven

se

Fiel

d m

ouse

-ear

ch

ickw

eed

G5

N

X

Li

chva

r 555

5 (R

M)

1983

Car

yoph

yll

Cer

astiu

m n

utan

s N

oddi

ng m

ouse

-ear

ch

ickw

eed

G5

N

X Li

chva

r 617

2 (R

M)

1983

Car

yoph

yll

Min

uarti

a nu

ttallii

[A

rena

ria n

utta

llii]

Britt

le s

titch

wor

t G

5 N

X

Col

lins

650

(RM

) 19

83

Car

yoph

yll

Paro

nych

ia s

essi

liflo

ra

Low

nai

lwor

t G

5 N

X

X O

Li

chva

r 561

4 (R

M)

1983

Car

yoph

yll

Sile

ne a

ntirr

hina

Sl

eepy

cat

chfly

G

5 N

O

X H

eide

l 203

6 (R

M)

2001

Car

yoph

yll

Sile

ne m

enzi

esii

var.

visc

osa

Whi

te c

atch

fly

G5T

? N

X

Lich

var 5

870

(RM

) 19

83

Car

yoph

yll

Sper

gula

ria s

alin

a

[S

. mar

ina]

Sa

ltmar

sh s

ands

purry

G

? E

X

Fe

rtig

1483

4 (R

M)

1994

Car

yoph

yll

Stel

laria

med

ia

Com

mon

chi

ckw

eed

G?

E

X X

Lich

var 5

697

(RM

) 19

83

C

heno

podi

At

riple

x ar

gent

ea

Silv

ersc

ale

G5

N

X X

O

Jone

s, D

itolla

, and

M

yers

s.n

. (BI

CA)

19

84

Che

nopo

di

Atrip

lex

cane

scen

s va

r. ca

nesc

ens

Four

-win

g sa

ltbus

h G

5T5?

N

X

X

Col

lins

916

(RM

) 19

83

Che

nopo

di

Atrip

lex

conf

ertif

olia

Sh

adsc

ale

G5

N

O

X

Knig

ht K

-46

(RM

) 19

83

C

heno

podi

At

riple

x ga

rdne

ri va

r. ga

rdne

rii

Gar

dner

's s

altb

ush

G5T

5 N

X

Lich

var 5

604

(RM

) 19

83

Che

nopo

di

Atrip

lex

mic

rant

ha

[A

. het

eros

perm

a]

Two-

scal

e sa

ltbus

h G

? E

X

N

elso

n 50

80 (R

M)

1979

Page 33: Vascular Plant Species Checklist of Bighorn Canyon National

33

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Che

nopo

di

Atrip

lex

pow

ellii

Pow

ell's

ora

che

G4G

5 N

X

Jone

s, D

itolla

, and

M

yers

s.n

. (R

M)

1984

Che

nopo

di

Atrip

lex

rose

a Tu

mbl

ing

orac

he

G?

E O

X

C

ollin

s 91

8 (R

M)

1983

Che

nopo

di

Bass

ia s

copa

ria

[B. s

ieve

rsia

na, K

ochi

a sc

opar

ia]

Mex

ican

fire

-wee

d

[Sum

mer

-cyp

ress

] G

? E

X O

O

Kn

ight

K-5

6 (R

M)

1983

Che

nopo

di

Che

nopo

dium

at

rovi

rens

Pi

nyon

goo

sefo

ot

G5

N

X X

Jo

nes,

Dito

lla, a

nd

Mye

rs s

.n. (

BIC

A)

1984

Che

nopo

di

Che

nopo

dium

be

rland

ieri

var.

zsch

acke

i

Pit-s

eed

goos

efoo

t G

5T?

N

X

X N

elso

n 70

23 (R

M)

1980

Che

nopo

di

Che

nopo

dium

frem

ontii

Fr

emon

t's g

oose

foot

G

5 N

X

X

Jone

s, D

itolla

, and

M

yers

s.n

. (R

M)

1984

Che

nopo

di

Che

nopo

dium

le

ptop

hyllu

m

Nar

row

-leaf

go

osef

oot

G5

N

O

X

Lich

var 6

889

(RM

) 19

83

Che

nopo

di

Che

nopo

dium

pr

ater

icol

a D

eser

t goo

sefo

ot

G5

N

X X

Fe

rtig

1943

2 (R

M)

1984

Che

nopo

di

Che

nopo

dium

rubr

um

var.

rubr

um

Red

goo

sefo

ot

G5

N

X X

H

eide

l 198

6 (B

ICA)

20

01

Che

nopo

di

Che

nopo

dium

sal

inum

[C. g

lauc

um v

ar.

salin

um]

Roc

ky M

ount

ain

goos

efoo

t G

5 N

X

Nel

son

5068

(RM

) 19

79

Che

nopo

di

Cor

ispe

rmum

sp.

Bu

gsee

d sp

. G

??

N

X Li

chva

r 635

5 (R

M)

1983

O

rigin

ally

de

term

ined

as

C.

hyss

opifo

lium

or

igin

ally

; the

im

mat

ure

spec

imen

was

an

nota

ted

by

Mos

yaki

n in

ta

xono

mic

revi

sion

of

gen

us to

eith

er

C. a

mer

ican

um o

r C

. villo

sum

C

heno

podi

C

yclo

lom

a at

riplic

ifoliu

m

Win

ged-

pigw

eed

G5

N

X

O

Hei

del 2

079

(RM

) 20

01

Che

nopo

di

Gra

yia

spin

osa

Spin

y ho

p-sa

ge

G5

N

X X

X Jo

nes,

Dito

lla, a

nd

Mye

rs s

.n. (

BIC

A)

1984

Che

nopo

di

Hal

oget

on g

lom

erat

us

Saltl

over

G

? E

X X

W

olf 7

9 (B

ICA)

19

82

C

heno

podi

Kr

asch

enin

niko

via

lana

ta

[Cer

atoi

des

lana

ta]

Win

terfa

t G

5 N

X

X

Lich

var 6

056

(RM

) 19

83

Page 34: Vascular Plant Species Checklist of Bighorn Canyon National

34

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Che

nopo

di

Mon

olep

is n

utta

lliana

N

utta

ll's p

over

ty-

wee

d G

5 N

X

Col

lins

721

(RM

) 19

83

Che

nopo

di

Salic

orni

a ru

bra

Red

sal

twor

t G

4 N

X

Hei

del 2

055

(RM

) 20

01

C

heno

podi

Sa

lsol

a tra

gus

[S. a

ustra

lis, S

. kal

i] Pr

ickl

y R

ussi

an-th

istle

G

? E

X O

O

W

olf 1

51 (B

ICA)

19

82

Che

nopo

di

Suae

da m

oqui

nii

[S. n

igra

, S. f

rutic

osa,

S.

torre

yana

]

Shru

bby

seep

wee

d G

5 N

X

X X

Lich

var 6

033

(RM

) 19

83

Che

nopo

di

Suck

leya

suc

kley

ana

Pois

on s

uckl

eya

G5

N

X

Fe

rtig

1483

3 (R

M)

1994

Che

nopo

di

[Sar

coba

t] Sa

rcob

atus

ve

rmic

ulat

us

Gre

asew

ood

G5

N

X X

X C

laus

en 6

6-23

8 (R

M),

Snel

ling

18

(RM

)

1924

Com

mel

in

Trad

esca

ntia

oc

cide

ntal

is

Prai

rie s

pide

rwor

t G

5 N

X

Lich

var 6

180

(BIC

A)

1983

Con

volv

ul

Con

volv

ulus

arv

ensi

s Fi

eld

bind

wee

d G

? E

O

X

Hei

del 2

037

(RM

) 20

01

Nox

ious

wee

d in

M

T an

d W

Y.

Earli

er v

ouch

er

spec

imen

s fro

m

BIC

A w

ere

colle

cted

at L

ovel

l vi

sito

rs c

ente

r. C

onvo

lvul

Ev

olvu

lus

nutta

llianu

s Sh

aggy

dw

arf

mor

ning

-glo

ry

G5

N

X

X H

eide

l 203

2 (R

M)

2001

Con

volv

ul

Ipom

oea

lept

ophy

lla

Bush

mor

ning

-glo

ry

G3G

5 N

X

Col

lins

893

(BIC

A)

1983

Cor

n C

ornu

s se

ricea

[C. s

tolo

nife

ra]

Red

osie

r

[Red

-osi

er d

ogw

ood]

G

5 N

O

X

O

Lich

var 5

973

(RM

) 19

83

Cra

ssul

Se

dum

lanc

eola

tum

La

nce-

leaf

sto

necr

op

G5

N

X

X Jo

nes,

Dito

lla, a

nd

Mye

rs s

.n. (

BIC

A)

1984

Cuc

urbi

t Br

yoni

a al

ba

Whi

te b

ryon

y G

? E

X H

eide

l 204

5 20

01

C

upre

ss

Juni

peru

s co

mm

unis

va

r. de

pres

sa

Com

mon

juni

per

G5T

5 N

X O

C

ollin

s 56

5 (B

ICA)

19

83

Cup

ress

Ju

nipe

rus

horiz

onta

lis

Cre

epin

g ju

nipe

r G

5 N

X

Ferti

g 19

455

(RM

) 19

84

C

upre

ss

Juni

peru

s os

teos

perm

a U

tah

juni

per

G5

N

X X

C

ollin

s 55

9 (R

M)

1981

Cup

ress

Ju

nipe

rus

scop

ulor

um

Roc

ky M

ount

ain

juni

per

G5

N

X

O

Wol

f 143

(BIC

A)

1982

Cyp

er

Car

ex a

quat

ilis

Leaf

y tu

ssoc

k se

dge

[Wat

er s

edge

] G

5 N

O

O

X

Col

lins

555

(RM

) 19

83

Cyp

er

Car

ex a

urea

G

olde

n-fru

it se

dge

G5

N

O

X X

Hei

del 1

708

(MO

NTU

) 19

98

Cyp

er

Car

ex b

revi

or

Shor

t-bea

k se

dge

G5?

N

X

Hei

del 2

143

(RM

) 20

01

C

yper

C

arex

dur

iusc

ula

[C

. ste

noph

ylla

] Sp

ike-

rush

sed

ge

[N

arro

w-le

aved

se

dge]

G5

N

X

H

eide

l 185

2 (M

ON

TU)

1999

Page 35: Vascular Plant Species Checklist of Bighorn Canyon National

35

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Cyp

er

Car

ex e

lyno

ides

Bl

ack-

root

sed

ge

G5

N

X

H

eide

l 206

5 (R

M)

2001

Pr

evio

usly

re

porte

d, b

ut n

o vo

uche

r sp

ecim

ens

loca

ted

for c

orro

bora

tion.

C

yper

C

arex

filif

olia

Th

read

-leaf

sed

ge

G5

N

X X

O

Lich

var 5

566

(RM

) 19

83

C

yper

C

arex

gey

eri

Gey

er's

sed

ge

[Elk

sed

ge]

G5

N

X

H

eide

l 204

1 (R

M)

2001

Cyp

er

Car

ex g

ravi

da v

ar.

grav

ida

Hea

vy s

edge

G

5T5?

N

X

Hei

del 2

101

(RM

) 19

99

Cyp

er

Car

ex h

oodi

i H

ood'

s se

dge

G4G

5 N

X X

Col

lins

777

(RM

) 19

83

C

yper

C

arex

hys

teric

ina

Porc

upin

e se

dge

G5

N

O

X X

Hei

del 1

716

(MO

NTU

) 19

98

Cyp

er

Car

ex in

ops

ssp.

he

lioph

ila

[C

. pen

sylv

anic

a va

r. di

gyna

, C. h

elio

phila

]

Long

-sto

lon

sedg

e

[Pen

nsyl

vani

a se

dge]

G

5T5

N

X

X C

ollin

s 57

4 (R

M)

1983

Cyp

er

Car

ex in

terio

r In

land

sed

ge

G5

N

O

X X

Lich

var 6

106

(BIC

A)

1983

Cyp

er

Car

ex m

icro

pter

a va

r. m

icro

pter

a Sm

all-w

ing

sedg

e G

5?

N

X

X Li

chva

r 615

0 (R

M)

1983

Cyp

er

Car

ex n

ebra

scen

sis

Neb

rask

a se

dge

G5

N

O

X O

H

eide

l 200

5 (R

M)

2001

Ea

rlier

repo

rts

base

d on

a

mis

iden

tifie

d sp

ecim

en

Cyp

er

Car

ex p

arry

ana

var.

parry

ana

Parry

's s

edge

G

4T4

N

X

X H

eide

l 181

9 (M

ON

TU)

1983

Cyp

er

Car

ex p

ellit

a

[C. l

anug

inos

a]

Woo

lly s

edge

G

5 N

X

X O

N

elso

n 54

56 (R

M)

1980

Cyp

er

Car

ex p

etas

ata

Lidd

on s

edge

G

5 N

X

Hei

del 2

028

(RM

) 20

01

C

yper

C

arex

pra

egra

cilis

C

lust

ered

fiel

d se

dge

G5

N

O

O

X Li

chva

r 610

7 (R

M)

1983

Cyp

er

Car

ex ro

ssii

Ros

s' s

edge

G

5 N

X

Hei

del 2

023b

(RM

) 20

01

C

yper

C

arex

sci

rpoi

dea

var.

scirp

iform

is

Can

adia

n si

ngle

-sp

ike

sedg

e G

5 T4

T5

N

X H

eide

l 210

9 (R

M)

2001

Cyp

er

Car

ex s

pren

gelii

Long

-bea

k se

dge

G5?

N

X

Lich

var 5

692

(RM

) 19

83

Lich

var 5

692

orig

inal

ly

mis

iden

tifie

d as

C.

hyst

eric

ina

Cyp

er

Car

ex s

tipat

a St

alk-

grai

n se

dge

G5

N

X H

eide

l 185

8 (M

ON

TU)

1998

Cyp

er

Car

ex v

allic

ola

Valle

y se

dge

G5

N

X

Jo

nes,

Dito

lla, a

nd

Mye

rs s

.n. (

BIC

A)

1984

Cyp

er

Car

ex v

iridu

la

Littl

e gr

een

sedg

e G

5?

N

X C

ollin

s 80

4 (R

M)

1983

Page 36: Vascular Plant Species Checklist of Bighorn Canyon National

36

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Cyp

er

Car

ex v

ulpi

noid

ea

Com

mon

fox

sedg

e G

5 N

X

Lich

var 6

119

(RM

) 19

83

C

yper

El

eoch

aris

ellip

tica

[E. t

enui

s va

r. bo

real

is]

Ellip

tic s

pike

-rush

G

5 N

X

Hei

del 2

006

(RM

) 20

01

Spec

imen

im

mat

ure

Cyp

er

Eleo

char

is p

alus

tris

Com

mon

spi

ke-ru

sh

G5

N

X O

O

N

elso

n 54

57 (R

M)

1980

Cyp

er

Eleo

char

is q

uinq

ueflo

ra

[E. p

auci

flora

] Fe

w-fl

ower

spi

ke-ru

sh

G5

N

X

H

eide

l 171

7 (M

ON

TU)

1998

Cyp

er

Scho

enop

lect

us a

cutu

s va

r. oc

cide

ntal

is

[Sci

rpus

acu

tus]

Har

d-st

em c

lub-

rush

G

5 N

X

Nel

son

7032

(RM

) 19

80

Cyp

er

Scho

enop

lect

us

mar

itim

us

[Sci

rpus

mar

itim

us,

Bolb

osch

oenu

s m

ariti

mus

var

. pa

ludo

sus]

Saltm

arsh

clu

b-ru

sh

G5

N

X X

H

eide

l 197

9 (R

M)

2001

Cyp

er

Scho

enop

lect

us

pung

ens

var.

poly

phyl

lus

[Sci

rpus

pun

gens

var

. po

lyph

yllu

s]

Thre

e-sq

uare

[Thr

ee-s

quar

e bu

lrush

]

G5

N

X O

X

Nel

son

7033

(RM

) 19

80

Cyp

er

Scho

enop

lect

us

tabe

rnae

mon

tani

[S

cirp

us v

alid

us]

Soft-

stem

clu

b-ru

sh

G5

N

O

X

Col

lins

902

(RM

) 19

83

Cyp

er

Scirp

us p

allid

us

Pale

bul

rush

G

5 N

O

X Li

chva

r 642

1 (R

M)

1983

Dry

opte

rid

[A

sple

ni]

Cys

topt

eris

frag

ilis

Britt

le b

ladd

er fe

rn

G5

N

X

O

Lich

var 5

959

(RM

) 19

83

Dry

opte

rid

[A

sple

ni]

Woo

dsia

ore

gana

var

. or

egan

a O

rego

n cl

iff fe

rn

G5T

4?

N

X

X Fe

rtig

1944

7 (R

M)

1983

Ea

rlier

spe

cim

ens

mis

iden

tifie

d as

W.

scop

ulin

a El

aeag

n El

aeag

nus

angu

stifo

lia

Rus

sian

-oliv

e G

? E

X X

O

Col

lins

718

(RM

) 19

81

El

aeag

n Sh

ephe

rdia

arg

ente

a Si

lver

buf

falo

-ber

ry

G5

N

X

X H

eide

l 199

1 (B

ICA)

19

83

El

aeag

n Sh

ephe

rdia

can

aden

sis

Rus

set b

uffa

lo-b

erry

G

5 N

X O

Li

chva

r 638

8 (R

M)

1983

Equi

set

Equi

setu

m a

rven

se

Fiel

d ho

rset

ail

G5

N

O

X

Lich

var 5

867

(RM

) 19

83

Eq

uise

t Eq

uise

tum

hye

mal

e Ta

ll sc

ourin

g-ru

sh

G5T

5 N

X

Lich

var 6

353

(RM

) 19

83

Eq

uise

t Eq

uise

tum

laev

igat

um

Smoo

th s

cour

ing-

rush

G

5 N

X

Lich

var 5

865

(RM

) 19

82

Eq

uise

t Eq

uise

tum

var

iega

tum

Va

riega

ted

scou

ring-

rush

G

5 N

X

Hei

del 2

102

(RM

) 20

01

Euph

orbi

C

ham

aesy

ce

glyp

tosp

erm

a

[E

upho

rbia

gl

ypto

sper

ma]

Rib

-see

d sa

ndm

at

G5

N

X X

C

ollin

s 85

9 (R

M)

1983

Page 37: Vascular Plant Species Checklist of Bighorn Canyon National

37

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Euph

orbi

C

ham

aesy

ce m

issu

rica

var.

peta

loid

ea

[E

upho

rbia

mis

suric

a va

r. pe

talo

idea

]

Prai

rie s

andm

at

G5

N

X O

X

Jone

s, D

itolla

, and

M

yers

s.n

. (BI

CA)

19

84

Euph

orbi

C

ham

aesy

ce

serp

yllif

olia

[Eup

horb

ia s

erpy

llifol

ia]

Thym

e-le

af s

andm

at

G5

N

X

X H

eide

l 201

4 (R

M)

2001

Euph

orbi

Eu

phor

bia

brac

hyce

ra

var.

robu

sta

[E

. rob

usta

]

Hor

ned

spur

ge

G5

N

X X

X Fe

rtig

1942

7 (R

M)

1983

Euph

orbi

Eu

phor

bia

esul

a va

r. ur

alen

sis

Wol

f's-m

ilk

[Lea

fy s

purg

e]

G5T

? E

O

O

1983

N

oxio

us w

eed

in

MT

and

WY.

R

elia

bly

repo

rted

for B

CN

RA

by

Lich

var e

t al.

(198

5) a

nd K

nigh

t et

al.

(198

7), b

ut

exac

t loc

atio

ns n

ot

know

n.

Euph

orbi

Eu

phor

bia

spat

hula

ta

War

ty s

purg

e G

5 N

X

Hei

del 2

047

(RM

) 20

01

Fa

b As

traga

lus

aret

ioid

es

[A. s

erio

leuc

us v

ar.

aret

ioid

es]

Cus

hion

milk

vetc

h G

4 N

X

Hei

del 2

135

(RM

) 19

98

Firs

t obs

erve

d in

BC

NR

A by

Hei

del

in 1

998;

vou

cher

co

llect

ed in

20

01.B

ooth

sp

ecim

en a

t M

ON

T w

as

prob

ably

col

lect

ed

outs

ide

BCN

RA.

Fa

b As

traga

lus

bisu

lcat

us

var.

bisu

lcat

us

Two-

groo

ve m

ilkve

tch

G5T

5 N

X

X O

M

esse

rsch

mid

t 10

3-(B

ICA)

19

81

Fab

Astra

galu

s ca

nade

nsis

va

r. ca

nade

nsis

C

anad

ian

milk

vetc

h G

5T4?

N

X

Hei

del 2

042

(RM

) 20

01

Prev

ious

ly

repo

rted,

but

no

vouc

her

spec

imen

s lo

cate

d fo

r cor

robo

ratio

n.

Fab

Astra

galu

s ce

ram

icus

va

r. fil

ifoliu

s Pa

inte

d m

ilkve

tch

G4T

4 N

X

Lich

var 6

184

(RM

) 19

83

Fab

Astra

galu

s ch

amae

leuc

e C

icad

a m

ilkve

tch

G5T

? N

X

Lich

var 5

600

(RM

) 19

83

Fab

Astra

galu

s ci

bariu

s Br

owse

milk

vetc

h G

4 N

X

Hei

del 1

930

(BIC

A)

1999

Fab

Astra

galu

s ci

cer

Chi

ckpe

a m

ilkve

tch

G

? E

X

H

eide

l 193

6 (B

ICA)

19

99

Page 38: Vascular Plant Species Checklist of Bighorn Canyon National

38

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Fab

Astra

galu

s cr

assi

carp

us v

ar.

pays

onii

Gro

und-

plum

G

5T5

N

X

X Li

chva

r 565

8 (R

M)

1983

Fab

Astra

galu

s dr

umm

ondi

i D

rum

mon

d's

milk

vetc

h G

5 N

X

Lich

var 5

984

(RM

) 19

83

Fab

Astra

galu

s ge

yeri

Gey

er's

milk

vetc

h G

4?

N

X X

Fe

rtig

1942

1 (R

M)

1984

Fab

Astra

galu

s gi

lvifl

orus

va

r. gi

lvifl

orus

Pl

ains

milk

vetc

h G

5T?

N

X

O

Lich

var 5

574

(RM

) 19

83

Fab

Astra

galu

s gr

acilis

Sl

ende

r milk

vetc

h G

5 N

X

Jone

s , D

itolla

, and

M

yers

s.n

. (BI

CA)

19

84

Fab

Astra

galu

s hy

alin

us

Sum

mer

milk

vetc

h G

4 N

X

Col

lins

910

(RM

) 19

76

Cite

d in

Dor

n (1

978)

Fa

b As

traga

lus

laxm

anni

i va

r. ro

bust

ior

[A. a

dsur

gens

var

. ro

bust

ior]

Laxm

ann'

s m

ilkve

tch

[S

tand

ing

milk

vetc

h]

G5T

5 N

X

X

Lich

var 5

922

(RM

) 19

83

Fab

Astra

galu

s lo

tiflo

rus

Lotu

s m

ilkve

tch

G5

N

X X

Fe

rtig

1943

3 (R

M)

1983

Fab

Astra

galu

s m

iser

var

de

cum

bens

Ti

mbe

r milk

vetc

h G

5T4?

N

X O

Li

chva

r 597

9 (R

M)

1983

Fab

Astra

galu

s m

isso

urie

nsis

var

. m

isso

urie

nsis

Mis

sour

i milk

vetc

h G

5T?

N

X

O

Col

lins

547

(RM

) 19

76

Fab

Astra

galu

s or

egan

us

Ore

gon

milk

vetc

h G

4?

N

X X

Li

chva

r 559

5 (R

M)

1983

Fab

Astra

galu

s pu

rshi

i var

. pu

rshi

i Pu

rsh'

s m

ilkve

tch

G5T

5 N

X

Lich

var 5

568

(BIC

A)

1983

Fab

Astra

galu

s sh

ortia

nus

Shor

t's m

ilkve

tch

G4

N

X

Li

chva

r 600

6 (R

M)

1983

Li

chva

r 600

6 or

igin

ally

iden

tifie

d as

A. o

rega

nus;

an

nota

ted

by

Aaro

n Li

sten

199

3 Fa

b As

traga

lus

spat

ulat

us

Tufte

d m

ilkve

tch

[Spo

onle

af m

ilkve

tch]

G

5 N

X

X O

Li

chva

r 557

9 (R

M)

1983

Fab

Car

agan

a ar

bore

scen

s Si

beria

n pe

ashr

ub

G?

E

X

Wol

f 110

(BIC

A)

1982

Pl

ante

d as

an

orna

men

tal

Fab

Dal

ea c

andi

da v

ar

olig

ophy

lla

[Pet

alos

tem

on

occi

dent

ale]

Whi

te p

rairi

e-cl

over

G

5T?

N

X X

X W

olf 1

36 (B

ICA)

19

82

Fab

Dal

ea p

urpu

rea

[P

etal

oste

mon

pu

rpur

eum

]

Viol

et p

rairi

e-cl

over

G

5 N

X

Lich

var 6

923

(RM

) 19

83

Fab

Gly

cyrrh

iza

lepi

dota

va

r. gl

utin

osa

Amer

ican

lico

rice

G5

N

X X

X N

elso

n 50

75 (R

M)

1979

Page 39: Vascular Plant Species Checklist of Bighorn Canyon National

39

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Fab

Hed

ysar

um b

orea

le

var.

bore

ale

Bore

al s

wee

t-vet

ch

G5T

? N

X X

Lich

var 5

985

(RM

) 19

83

Fab

Lupi

nus

arge

nteu

s va

r. ar

gent

eus

Silv

er-s

tem

lupi

ne

G5T

5 N

X O

H

eide

l 202

6 19

83

Fab

Lupi

nus

pusi

llus

var.

inte

rmon

tanu

s R

usty

lupi

ne

G5?

T?

N

X X

Jo

nes,

Dito

lla, a

nd

Mye

rs s

.n. (

RM

) 19

84

Fab

Lupi

nus

wye

thii

var.

wye

thii

[L. p

olyp

hyllu

s va

r. hu

mic

ola]

Wye

th's

lupi

ne

G5T

4 N

X

Col

lins

766

(RM

) 19

83

Fab

Med

icag

o lu

pulin

a Bl

ack

med

ick

G?

E O

X

O

Col

lins

881

(BIC

A)

1983

Fab

Med

icag

o sa

tiva

Alfa

lfa

G?T

? E

X X

O

Col

lins

717

(BIC

A)

1982

Fab

Mel

ilotu

s al

bus

[incl

uded

in M

. of

ficin

alis

by

Karte

sz]

Whi

te s

wee

t-clo

ver

G?

E O

X C

ollin

s 83

5 (B

ICA)

19

83

Fab

Mel

ilotu

s of

ficin

alis

Ye

llow

sw

eet-c

love

r G

? E

X X

O

Lich

var 5

991

(RM

) 19

82

Fa

b O

xytro

pis

bess

eyi v

ar.

falla

x Be

ssey

's lo

cow

eed

G5T

3 N

X

X

Des

pain

16

(RM

) 19

64

Fab

Oxy

tropi

s be

ssey

i var

. ve

ntos

a Be

ssey

's lo

cow

eed

G5T

3?

N

X

H

eide

l 171

8 (R

M)

1998

Fab

Oxy

tropi

s ca

mpe

stris

va

r. sp

icat

a

[Oxy

tropi

s ca

mpe

stris

va

r. gr

acilis

]

Nor

ther

n ye

llow

lo

cow

eed

G5T

5 N

X

X

Hei

del 2

024

(RM

) 19

83

BIC

A sp

ecim

en

may

be

O. s

eric

ea

var.

spic

ata

Fab

Oxy

tropi

s la

gopu

s va

r. at

ropu

rpur

ea

Har

e-fo

ot lo

cow

eed

G4G

5 T4

T5

N

X

Li

chva

r 566

1 (R

M)

1983

Fab

Oxy

tropi

s se

ricea

var

. sp

ecio

sa

[Oxy

tropi

s se

ricea

var

. sp

icat

a]

Roc

ky M

ount

ain

loco

wee

d G

5T?

N

X

H

eide

l 199

8 (B

ICA)

20

01

Earli

er re

ports

ba

sed

on a

m

isid

entif

ied

spec

imen

Fa

b Pe

diom

elum

ar

goph

yllu

m

Silv

er-le

af In

dian

-br

eadr

oot

G5

N

O

19

98

Obs

erve

d at

T6S

R

30E

S18

by

Hei

del

Fab

Pedi

omel

um

escu

lent

um [

Psor

alea

es

cule

nta]

Larg

e In

dian

-br

eadr

oot

G5

N

X

C

ollin

s 74

1 (R

M)

1983

Fab

Psor

alid

ium

la

nceo

latu

m

[P

sora

lea

lanc

eola

ta]

Wild

lem

onw

eed

[Lem

on s

curfp

ea]

G5

N

X

X Li

chva

r 618

5 (R

M)

1983

Fab

Psor

alid

ium

tenu

iflor

um

[Pso

rale

a te

nuifl

ora]

Sl

ende

r-flo

wer

le

mon

wee

d

[S

lend

er-fl

ower

sc

urfp

ea]

G5

N

X

Jo

nes,

Dito

lla,

Mye

rs s

.n. (

BIC

A)

1984

Page 40: Vascular Plant Species Checklist of Bighorn Canyon National

40

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Fab

Spha

erop

hysa

sal

sula

R

ed b

ladd

er-v

etch

G

? E

X X

X N

elso

n 54

58 (R

M)

1980

Fab

Ther

mop

sis

rhom

bifo

lia

var.

rhom

bifo

lia

Prai

rie g

olde

n-ba

nner

[R

ound

-leav

ed

gold

en-p

ea]

G5T

? N

O

X

Li

chva

r 566

6 (B

ICA)

19

83

Fab

Trifo

lium

frag

iferu

m

Stra

wbe

rry-h

ead

clov

er

G?

E

X X

Hei

del 2

112

(RM

) 20

01

Fab

Trifo

lium

hyb

ridum

Al

sike

clo

ver

G?

E

X X

Col

lins

757

(RM

) 19

83

Fa

b Vi

cia

amer

ican

a va

r. m

inor

Am

eric

an p

urpl

e ve

tch

G5T

5 N

X

Lich

var 5

918

(RM

) 19

83

Fum

ari

Cor

ydal

is a

urea

var

. au

rea

Scra

mbl

edeg

gs

[Gol

dens

mok

e]

G5T

5 N

X

Lich

var 6

073

(RM

) 19

83

Gen

tian

Gen

tiana

affi

nis

var.

affin

is

Plea

ted

gent

ian

G5T

5 N

X

Jone

s, D

itolla

, and

M

yers

s.n

. (BI

CA)

19

84

Ger

ani

Erod

ium

cic

utar

ium

R

ed-s

tem

sto

rk's

-bill

G?

E

X

Lich

var 5

717

(RM

) 19

83

G

eran

i G

eran

ium

bic

knel

lii

Nor

ther

n cr

ane'

s-bi

ll G

5 N

X

Hei

del 2

099

(RM

) 20

01

Prev

ious

ly

repo

rted,

but

no

vouc

her

spec

imen

s lo

cate

d fo

r cor

robo

ratio

n.

Ger

ani

Ger

aniu

m v

isco

ssim

um

var.

visc

ossi

mum

St

icky

pur

ple

cran

e's-

bill

G5T

5?

N

O

X

Lich

var 5

700

(RM

) 19

83

Gro

ssul

ari

Rib

es a

mer

ican

um

Wild

bla

ck c

urra

nt

G5

N

X

O

Col

lins

591(

RM

) 19

83

G

ross

ular

i R

ibes

aur

eum

var

. au

reum

G

olde

n cu

rrant

G

5T?

N

O

X O

Li

chva

r 564

9 (R

M)

1983

Gro

ssul

ari

Rib

es c

ereu

m v

ar.

pedi

cella

re

Whi

te s

quaw

cur

rant

G

5T5

N

X X

Li

chva

r 563

2 (R

M)

1983

Gro

ssul

ari

Rib

es o

xyac

anth

oide

s va

r. se

tosu

m

Can

adia

n go

oseb

erry

G

5 T4

T5

N

X

C

ollin

s 66

2 (R

M)

1983

Hyd

roch

arit

Eger

ia d

ensa

[E

lode

a de

nsa]

Br

azilia

n w

ater

-wee

d G

5 E

X

H

eide

l 193

7 (M

ON

TU)

1999

O

rigin

ally

de

term

ined

as

Hip

puris

vul

garis

(H

eide

l and

Fer

tig

2000

) H

ydro

phyl

l El

lisia

nyc

tele

a Au

nt L

ucy

[W

ater

pod]

G

5 N

X

Lich

var 6

181

(RM

) 19

83

Hyd

roph

yll

Nam

a de

nsum

var

. pa

rvifl

orum

Le

afy

fiddl

elea

f G

5T5

N

X

Jo

nes,

Dito

lla, a

nd

Mye

r s.n

. (BI

CA)

19

84

Hyd

roph

yll

Phac

elia

gla

ndul

osa

Gla

ndul

ar s

corp

ion-

wee

d

[G

land

ular

pha

celia

]

G4

N

X

H

eide

l 205

3 (R

M)

2001

Pr

evio

usly

re

porte

d, b

ut n

o vo

uche

r sp

ecim

ens

loca

ted

for c

orro

bora

tion.

Page 41: Vascular Plant Species Checklist of Bighorn Canyon National

41

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Hyd

roph

yll

Phac

elia

has

tata

var

. ha

stat

a Si

lver

-leaf

sco

rpio

n-w

eed

G5T

5 N

X

Col

lins

783

(RM

) 19

83

Hyd

roph

yll

Phac

elia

ives

iana

Iv

es' s

corp

ion-

wee

d G

5 N

X

X X

Lich

var 5

613

(RM

) 19

76

H

ydro

phyl

l Ph

acel

ia li

near

is

Thre

ad-le

af s

corp

ion-

wee

d G

5 N

O

X

X Li

chva

r 569

0 (R

M)

1983

Irid

Iris

mis

sour

iens

is

Roc

ky M

ount

ain

iris

G5

N

X

X Li

chva

r 593

5 (R

M)

1983

Irid

Sisy

rinch

ium

id

ahoe

nse

var.

occi

dent

ale

Idah

o bl

ue-e

yed

gras

s G

5T?

N

X H

eide

l 181

7 (M

ON

TU)

1998

Irid

Sisy

rinch

ium

m

onta

num

St

rict b

lue-

eyed

gra

ss

G5

N

X Li

chva

r 606

8 (R

M)

1983

Jugl

and

Jugl

ans

cine

rea

Whi

te w

alnu

t G

5 E

X

Li

chva

r 606

7 (R

M)

1983

Pl

ante

d at

an

old

hom

este

ad

Junc

Ju

ncus

arti

cula

tus

Join

t-lea

f rus

h G

5 N

X

Hei

del 2

115

(RM

) 20

01

Ju

nc

Junc

us b

altic

us v

ar.

mon

tanu

s Ba

ltic

rush

G

5T5

N

O

X

Lich

var 5

879

(BIC

A)

1983

Junc

Ju

ncus

buf

oniu

s To

ad ru

sh

G5

N

X X

N

elso

n 50

70 (R

M)

1979

Junc

Ju

ncus

dud

leyi

[J. t

enui

s va

r. du

dley

i] D

udle

y's ru

sh

G5

N

X Li

chva

r 637

1 (R

M)

1983

Junc

Ju

ncus

ens

ifoliu

s va

r. m

onta

nus

Dag

ger-l

eaf r

ush

G5T

5 N

X

X

Nel

son

5072

(RM

) 19

79

Junc

Ju

ncus

long

isty

lis

Long

-sty

le ru

sh

G5

N

X

X Li

chva

r 612

7 (R

M)

1983

Junc

Ju

ncus

nod

osus

Tu

bero

us ru

sh

G5

N

X H

eide

l 208

7b (R

M)

2001

Junc

Ju

ncus

torre

yi

Torre

y's ru

sh

G5

N

X O

X

Nel

son

5073

(RM

) 19

79

Ju

ncag

in

Trig

loch

in m

ariti

ma

var.

elat

a Se

asid

e ar

row

-gra

ss

G5

N

O

X X

Hei

del 1

859

(BIC

A)

1998

Junc

agin

Tr

iglo

chin

pal

ustri

s M

arsh

arro

w-g

rass

G

5 N

X

Hei

del 1

723

(MO

NTU

) 19

98

Lam

i H

edeo

ma

drum

mon

dii

Dru

mm

ond'

s fa

lse

penn

yroy

al

G5

N

X X

X Jo

nes,

Dito

lla, a

nd

Mye

rs s

.n. (

BIC

A)

1983

Lam

i H

edeo

ma

hisp

ida

Rou

gh fa

lse

penn

yroy

al

G5

N

O

19

98

Obs

erve

d on

slo

pe

north

of O

k-a-

beh

Mar

ina

(T6S

R31

E S1

8 SW

1/4

) by

Hei

del

La

mi

Lyco

pus

aspe

r R

ough

wat

er-

hore

houn

d G

5 N

X

X

Nel

son

5062

(RM

) 19

79

Lam

i M

enth

a ar

vens

is

Amer

ican

wild

min

t

G5

N

X O

X

Nel

son

5063

(RM

) 19

79

Lam

i M

onar

da fi

stul

osa

var.

men

thifo

lia

Osw

ego-

tea

G5T

5 N

O

X C

ollin

s 80

1 (R

M)

1983

Page 42: Vascular Plant Species Checklist of Bighorn Canyon National

42

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Lam

i N

epet

a ca

taria

C

atni

p G

? E

O

X

Hei

del 2

093

(RM

) 20

01

Prev

ious

ly

repo

rted,

but

no

vouc

her

spec

imen

s lo

cate

d fo

r cor

robo

ratio

n.

Lam

i Sa

lvia

refle

xa

Lanc

e-le

af s

age

G?

N

O

20

01

Obs

erve

d by

Su

zann

e M

orst

ad

at h

ighw

ay tu

rn to

H

orse

shoe

Ben

d;

too

late

in s

easo

n to

col

lect

vou

cher

La

mi

Scut

ella

ria g

aler

icul

ata

Hoo

ded

skul

lcap

G

5 N

X

Col

lins

811

(RM

) 19

83

La

mi

Teuc

rium

can

aden

se

var.

occi

dent

ale

Amer

ican

ger

man

der

G5T

4

N

X

N

elso

n 70

19 (R

M)

1980

Lem

n Le

mm

a tu

rioni

fera

[L. m

inor

] Tu

rion

duck

wee

d G

5 N

X

X

Hei

del 2

125

(RM

) 20

01

Lili

Friti

llaria

atro

purp

urea

Sp

otte

d m

issi

onbe

lls

[C

heck

er li

ly]

G5

N

X Li

chva

r 567

3 (R

M)

1983

Lili

Friti

llaria

pud

ica

Yello

w m

issi

onbe

lls

[Yel

low

bells

] G

5 N

O

2001

O

bser

ved

in fr

uit

by H

eide

l Li

li Li

lium

phi

lade

lphi

cum

va

r. an

dinu

m

Woo

d lily

G

5 T4

T5

N

X H

eide

l 214

8 (B

ICA)

20

01

Prev

ious

ly

repo

rted,

but

no

vouc

her

spec

imen

s lo

cate

d fo

r cor

robo

ratio

n.

Lili

[A

lli]

Alliu

m c

ernu

um

Nod

ding

oni

on

G5

N

X

O

Jone

s, D

itolla

, and

M

yers

s.n

. (BI

CA)

19

84

Lili

[A

lli]

Alliu

m g

eyer

i var

. ge

yeri

Gey

er's

oni

on

G4G

5 T4

N

X

Col

lins

785

(RM

) 19

83

Col

lins

785

may

be

A. t

extil

e Li

li

[Alli]

Al

lium

text

ile

Whi

te w

ild o

nion

[Tex

tile

onio

n]

G5?

N

X

X X

Jone

s s.

n. (R

M)

1982

Lili

[A

spar

ag]

Aspa

ragu

s of

ficin

alis

As

para

gus

G5?

E

X X

Fe

rtig

1840

7 (R

M)

1982

Lili

[Cal

ocho

rt]

Cal

ocho

rtus

gunn

ison

ii G

unni

son'

s m

arip

osa-

lily

G5

N

X

H

eide

l 206

7 (R

M)

2001

Lili

[Cal

ocho

rt]

Cal

ocho

rtus

nutta

llii

Sego

-lily

G

5 N

X X

Lich

var 6

137

(RM

) 19

81

Lili

[Cal

ocho

rt]

Pros

arte

s tra

chyc

arpa

[Dis

poru

m

trach

ycar

pum

]

Rou

gh-fr

uit f

airy

bells

G

5 N

X

Col

lins

564

(RM

) 19

83

Page 43: Vascular Plant Species Checklist of Bighorn Canyon National

43

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Lili

[Con

valla

ri]

Mai

anth

emum

ra

cem

osum

var

. am

plex

icau

le

[Sm

ilaci

na ra

cem

osa]

Feat

hery

fals

e So

lom

on's

-sea

l [F

alse

spi

kena

rd]

G5T

? N

X X

Lich

var 5

940

(RM

) 19

83

Lili

[Con

valla

ri]

Mai

anth

emum

st

ella

tum

[Sm

ilaci

na

stel

lata

]

Star

ry fa

lse

Solo

mon

's-s

eal

[Spi

kena

rd]

G5

N

X

O

Wol

f 113

(BIC

A)

1982

Lili

[M

elan

thi]

Ziga

denu

s el

egan

s M

ount

ain

deat

hcam

as

G5

N

X

X Fe

rtig

1945

4 (R

M)

2001

Li

chva

r 616

5 w

as

colle

cted

out

side

of

the

BCN

RA

boun

dary

Li

li

[Mel

anth

i] Zi

gade

nus

vene

nosu

s va

r. gr

amin

eus

Mea

dow

dea

thca

mas

G

5T5

N

O

X

Col

lins

562

(RM

) 19

83

Lin

Linu

m le

wis

ii Pr

airie

flax

[B

lue

flax]

G

4G5

N

X X

O

T. P

eter

s, s

.n.

(BIC

A)

1981

Lin

Linu

m ri

gidu

m

Larg

e-flo

wer

yel

low

fla

x G

4G5

N

X

Jo

nes,

Dito

lla, a

nd

Mye

rs s

.n. (

BIC

A)

1984

Loas

M

entz

elia

alb

icau

lis

Whi

te-s

tem

bl

azin

gsta

r G

5 N

X

X

Col

lins

715

(RM

) 19

76

Loas

M

entz

elia

dec

apet

ala

Gum

bo-li

ly

[Eve

ning

-sta

r] G

5 N

X

O

C

ollin

s 85

1 (R

M)

1983

Loas

M

entz

elia

mon

tana

Va

riega

ted-

brac

t bl

azin

gsta

r G

4 N

X

Jone

s, D

itolla

, and

M

yers

s.n

. (BI

CA)

19

84

Loas

M

entz

elia

pum

ila

Gol

den

blaz

ings

tar

[Wyo

min

g st

ickl

eaf]

G4

N

X X

W

elp

7969

(BIC

A)

1998

Mal

v M

alva

par

viflo

ra

Smal

l-who

rl m

allo

w

G?

E

X

Hei

del 2

158

(RM

) 20

01

Prev

ious

ly

repo

rted,

but

no

vouc

her

spec

imen

s lo

cate

d fo

r cor

robo

ratio

n.

Mal

v Sp

haer

alce

a co

ccin

ea

Scar

let g

lobe

-mal

low

G

5?

N

X O

O

W

olf 9

2 (B

ICA)

19

82

M

or

Mor

us a

lba

Whi

te m

ulbe

rry

G?

E X

X

Hei

del 1

990

(RM

) 20

01

Plan

ted

as a

n or

nam

enta

l N

ycta

gin

Abro

nia

ellip

tica

[A

. fra

gran

s va

r. el

liptic

a]

Frag

rant

whi

te s

and-

verb

ena

G5

N

X O

Lich

var 6

025

(RM

) 19

83

Nyc

tagi

n M

irabi

lis li

near

is

Nar

row

-leaf

four

-o'

cloc

k G

5 N

X

X X

Lich

var 6

063

(RM

) 19

83

Nyc

tagi

n Tr

ipte

roca

lyx

mic

rant

hus

[Abr

onia

mic

rant

ha]

Smal

l-flo

wer

sa

ndpu

ffs

G5

N

X X

O

Jone

s, D

itolla

, and

M

yers

s.n

. (BI

CA)

19

84

Ole

Fr

axin

us p

enns

ylva

nica

G

reen

ash

G

5 N

X

Hei

del 2

010

(RM

) 20

01

Plan

ted

as a

n or

nam

enta

l

Page 44: Vascular Plant Species Checklist of Bighorn Canyon National

44

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Ole

Li

gust

rum

vul

gare

Eu

rope

an p

rivet

G

? E

X

M

esse

rsch

mid

t 104

(B

ICA)

19

81

Plan

ted

as a

n or

nam

enta

l O

nagr

C

alyl

ophu

s se

rrula

tus

Yello

w s

undr

ops

G5

N

X Li

chva

r 636

7 (R

M)

1983

Ona

gr

Cam

isso

nia

min

or

Gre

en R

iver

sun

cup

G4

N

X X

Jo

nes,

Dito

lla, a

nd

Mye

rs s

.n. (

BIC

A)

1984

Ona

gr

Cam

isso

nia

scap

oide

a Pa

iute

sun

cup

G5T

? N

X

Lich

var 6

061

(RM

) 19

83

O

nagr

C

ham

erio

n an

gust

ifoliu

m

[Epi

lobi

um

angu

stifo

lium

]

Nar

row

leaf

fire

wee

d G

5 N

X

Col

lins

820

(RM

) 19

83

varie

ty n

ot

dete

rmin

ed

Ona

gr

Circ

aea

alpi

na v

ar.

alpi

na

Smal

l enc

hant

er's

-ni

ghts

hade

G

5T5

N

X Li

chva

r 689

9 (R

M)

1983

Ona

gr

Epilo

bium

br

achy

carp

um

[E

. pan

icul

atum

]

Tall

annu

al

willo

whe

rb

G5

N

X H

eide

l 215

0 (R

M)

2001

Pr

evio

usly

re

porte

d, b

ut n

o vo

uche

r sp

ecim

ens

loca

ted

for c

orro

bora

tion.

O

nagr

Ep

ilobi

um c

iliatu

m v

ar.

cilia

tum

Fr

inge

d w

illow

herb

G

5T5

N

O

X

Lich

var 6

920

(RM

) 19

83

Ona

gr

Epilo

bium

cilia

tum

var

. gl

andu

losu

m

Frin

ged

willo

whe

rb

G5T

5 N

X

Lich

var 6

374

(RM

) 19

83

Ona

gr

Gau

ra c

occi

nea

Scar

let b

eebl

osso

m

[S

carle

t gau

ra]

G5

N

X X

O

Col

lins

644

(RM

) 19

83

Ona

gr

Gau

ra m

ollis

[G

. par

viflo

ra]

Velv

etw

eed

[Sm

all-f

low

er g

aura

] G

4G5

N

X C

ollin

s 83

1 (R

M)

1983

Ona

gr

Oen

othe

ra a

lbic

aulis

W

hite

-ste

m e

veni

ng-

prim

rose

G

5 N

X

Dor

n 26

35

(MO

NT?

) 19

76

Rep

orte

d by

Dor

n (1

978)

O

nagr

O

enot

hera

cae

spito

sa

var.

caes

pito

sa

[O

. ces

pito

sa v

ar.

cesp

itosa

]

Tufte

d ev

enin

g-pr

imro

se

G5T

5 N

X

X

Mes

sers

chm

idt 5

1 (B

ICA)

19

81

Ona

gr

Oen

othe

ra p

allid

a va

r. tri

choc

alyx

W

hite

-pol

e ev

enin

g-pr

imro

se

G5T

? N

X

O

X Li

chva

r 601

4 (R

M)

1983

Ona

gr

Oen

othe

ra v

illosa

var

. st

rigos

a

[O. d

epre

ssa]

Hai

ry e

veni

ng-

prim

rose

G

5T?

N

X Li

chva

r 639

6 (R

M)

1983

Orc

hid

Cor

allo

rhiz

a m

acul

ata

Sum

mer

cor

alro

ot

[Spo

tted

cora

lroot

] G

5 T3

T5

N

X

C

ollin

s 65

6 (R

M)

1983

va

riety

not

de

term

ined

O

rchi

d C

oral

lorh

iza

stria

ta v

ar.

vree

land

ii H

oode

d co

ralro

ot

[S

tripe

d co

ralro

ot]

G5T

? N

X

Col

lins

781

(RM

) 19

83

Orc

hid

Goo

dyer

a ob

long

ifolia

G

reen

-leaf

ra

ttles

nake

-pla

ntai

n

G5?

N

X

Lich

var 5

965

(RM

) 19

83

Page 45: Vascular Plant Species Checklist of Bighorn Canyon National

45

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Orc

hid

Pipe

ria u

nala

scen

sis

[H

aben

aria

un

alas

cens

is]

Alas

ka re

in-o

rchi

d G

5 N

X

Lich

var 6

395

(RM

) 19

83

Orc

hid

Plat

anth

era

dila

tata

[H

aben

aria

dilit

ata]

Sc

entb

ottle

[W

hite

bog

orc

hid]

G

5 N

X O

Li

chva

r 587

5 (R

M)

1983

va

riety

not

de

term

ined

O

rchi

d Pl

atan

ther

a hu

rone

nsis

La

ke H

uron

gre

en

orch

id

G5

N

X

Jo

nes,

Dito

lla, a

nd

Mye

rs s

.n. (

BIC

A)

1983

Fo

rmer

ly in

clud

ed

in P

lata

nthe

ra

[Hab

enar

ia]

hype

rbor

ea

Oro

banc

h O

roba

nche

fasc

icul

ata

Clu

ster

ed b

room

-rape

G

4 N

X

X

Jone

s, D

itolla

, and

M

yers

s.n

. (BI

CA)

19

84

Oro

banc

h O

roba

nche

ludo

vici

ana

var.

ludo

vici

ana

Loui

sian

a br

oom

-rape

G

5 N

O

1998

O

bser

ved

belo

w

new

pic

nic

area

at

Hor

sesh

oe B

end

(T57

N R

95W

S1)

by

Hei

del

Oro

banc

h O

roba

nche

uni

flora

var

. oc

cide

ntal

is

Nak

ed b

room

-rape

G

5 N

X

Col

lins

584

(RM

) 19

83

Pin

Abie

s la

sioc

arpa

Su

balp

ine

fir

G5

N

O

1984

R

epor

ted

in p

lots

by

Kni

ght e

t al.

(198

7)

Pin

Pice

a en

gelm

anni

i En

gelm

ann

spru

ce

G5

N

X

Li

chva

r 595

5 (N

Y)

1983

Pin

Pinu

s fle

xilis

Li

mbe

r pin

e G

5 N

X

Lich

var 5

645

(RM

) 19

83

Pi

n Pi

nus

pond

eros

a Po

nder

osa

pine

G

5 N

O

X H

eide

l 203

9 (R

M)

2001

Pr

evio

usly

re

porte

d, b

ut n

o vo

uche

r sp

ecim

ens

loca

ted

for c

orro

bora

tion.

Pi

n Ps

eudo

tsug

a m

enzi

esii

var.

glau

ca

Dou

glas

-fir

G5T

5 N

X O

C

ollin

s 56

9 (R

M)

1983

Plan

tagi

n Pl

anta

go m

ajor

G

reat

pla

ntai

n G

5 E

O

O

X H

eide

l 211

3 (R

M)

2001

Pr

evio

usly

re

porte

d, b

ut n

o vo

uche

r sp

ecim

ens

loca

ted

for c

orro

bora

tion.

Plan

tagi

n Pl

anta

go p

atag

onic

a va

r. pa

tago

nica

W

oolly

pla

ntai

n G

5T5

N

X O

X

Lich

var 6

058

(RM

) 19

83

Po

Achn

athe

rum

hy

men

oide

s

[O

ryzo

psis

hy

men

oide

s]

Indi

an ri

ce g

rass

G

5 N

X

X O

Pe

ters

99

(BIC

A)

1981

Page 46: Vascular Plant Species Checklist of Bighorn Canyon National

46

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Po

Achn

athe

rum

le

tterm

anii

[Stip

a le

tterm

anii]

Lette

rman

's ric

e gr

ass

G5

N

X

C

ollin

s 73

2 (R

M)

1983

D

uplic

ate

spec

imen

at B

ICA

is m

isid

entif

ied

Po

Achn

athe

rum

nel

soni

i va

r. do

rei

[Stip

a ne

lson

ii va

r. do

rei,

S. c

olum

bian

a,

S. w

illia

msi

i]

Nel

son'

s ric

e gr

ass

G5T

5?

N

X

O

Lich

var 5

976

(RM

) 19

83

Po

Achn

athe

rum

nel

soni

i va

r. ne

lson

ii

[Stip

a ne

lson

ii va

r. ne

lson

ii]

Nel

son'

s ric

e gr

ass

G5T

5 N

X

Hei

del 2

007

(RM

) 20

01

Po

Aegi

lops

cyl

indr

ica

Join

ted

goat

gra

ss

G?

E

X

Hei

del 1

849

(MO

NTU

) 19

99

Po

Agro

pyro

n cr

ista

tum

va

r. cr

ista

tum

C

rest

ed w

heat

gra

ss

G5T

5 E

X X

W

olf 1

24 (B

ICA)

19

81

Po

Agro

pyro

n de

serto

rum

[A. c

rista

tum

var

. de

serto

rum

]

Clu

ster

ed w

heat

gr

ass

G5T

? E

X

Li

chva

r 588

2 (R

M)

1983

Po

Agro

stis

exa

rata

Sp

iked

ben

t G

5 N

X

Lich

var 6

378

(RM

) 19

83

Po

Ag

rost

is s

cabr

a R

ough

ben

t G

5 N

O

X

O

Knig

ht K

-72

(RM

) 19

83

Po

Ag

rost

is s

tolo

nife

ra

[A

. alb

a]

Spre

adin

g be

nt

G5

E O

X Li

chva

r 689

6 (R

M)

1983

Po

Alop

ecur

us

arun

dina

ceus

C

reep

ing

foxt

ail

G?

E X

X X

Hei

del 1

984

(RM

) 19

99

Orig

inal

ly

dete

rmin

ed a

s A.

pr

aten

sis

Po

Andr

opog

on g

erar

dii

Big

blue

stem

G

5 N

X

Lich

var 6

930

(RM

) 19

83

Po

Ar

istid

a pu

rpur

ea v

ar.

fend

leria

na

[A

. fen

dler

iana

]

Purp

le th

ree-

awn

G5T

5 N

X

X

Col

lins

543

(RM

) 19

76

Po

Aris

tida

purp

urea

var

. lo

ngis

eta

[A

. lon

gise

ta]

Purp

le th

ree-

awn

G5T

5?

N

X

O

Col

lins

542

(RM

) 19

83

Po

Beck

man

nia

syzi

gach

ne

Amer

ican

slo

ugh

gras

s G

5 N

X

Hei

del 2

012

(RM

) 20

01

Po

Bout

elou

a cu

rtipe

ndul

a Si

de-o

ats

gram

a G

5 N

X

Col

lins

888

(RM

) 19

83

Po

Bo

utel

oua

grac

ilis

Blue

gra

ma

G5

N

X X

X W

olf 2

1 (B

ICA)

19

81

Po

Br

omus

car

inat

us

Cal

iforn

ia b

rom

e G

5 N

X

Lich

var 6

169

(RM

) 19

83

Po

Br

omus

cilia

tus

Frin

ged

brom

e G

5 N

X

Lich

var 6

909

(RM

) 19

83

Po

Br

omus

com

mut

atus

M

eado

w b

rom

e G

? E

O

X

Lich

var 6

408

(RM

) 19

83

Po

Br

omus

iner

mis

var

. in

erm

is

Smoo

th b

rom

e G

5T?

E O

X

O

Mes

sers

chm

idt 1

9 (B

ICA)

1981

Page 47: Vascular Plant Species Checklist of Bighorn Canyon National

47

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Po

Brom

us ja

poni

cus

Japa

nese

bro

me

G?

E

O

20

01

Prev

ious

ly

repo

rted,

but

no

vouc

her

spec

imen

s lo

cate

d fo

r cor

robo

ratio

n;

obse

rved

in h

ills

abov

e Ft

. Sm

ith.

Po

Brom

us te

ctor

um

Che

at g

rass

G

? E

X X

X Li

chva

r 562

5 (R

M)

1983

Po

Cal

amag

rost

is s

trict

a ss

p. in

expa

nsa

[C. i

nexp

ansa

]

Slim

-ste

m re

ed g

rass

G

5T5

N

X

H

eide

l 200

2 (R

M)

2001

Po

Cal

amov

ilfa

long

ifolia

Pr

airie

san

d-re

ed

G5

N

O

O

X C

ollin

s 79

5 (R

M)

1983

Po

Cat

abro

sa a

quat

ica

Wat

er w

horl

gras

s G

5 N

X

Hei

del

1857

(MO

NTU

) 19

99

Po

Cin

na la

tifol

ia

Slen

der w

ood-

reed

G

5 N

X

Hei

del 1

921

(MO

NTU

) 19

99

Po

Dac

tylis

glo

mer

ata

Orc

hard

gra

ss

G?

E O

X C

ollin

s 58

8 (R

M)

1983

Po

Dan

thon

ia c

alifo

rnic

a C

alifo

rnia

wild

oat

gr

ass

G5

N

X H

eide

l 186

0 (M

ON

TU)

1999

Po

Dan

thon

ia s

pica

ta

Pove

rty w

ild o

at g

rass

G

5 N

X

Hei

del 2

097

(RM

) 20

01

Po

D

anth

onia

uni

spic

ata

Few

-flow

er w

ild o

at

gras

s G

5 N

X

Hei

del 2

046

(RM

) 20

01

Prev

ious

ly

repo

rted,

but

no

vouc

her

spec

imen

s lo

cate

d fo

r cor

robo

ratio

n.

Po

Des

cham

psia

ce

spito

sa

Tufte

d ha

ir gr

ass

G5

N

X

O

Col

lins

736

(RM

) 19

83

Po

Dic

hant

heliu

m

acum

inat

um v

ar.

fasc

icul

atum

[D

. acu

min

atum

var

. la

nugi

nosu

m]

Tape

red

rose

tte g

rass

G

5T5

N

X

X H

eide

l 172

0 (M

ON

TU)

1998

Po

Dis

tichl

is s

pica

ta

[D. s

trict

a]

Coa

stal

sal

t gra

ss

G5

N

X O

O

N

elso

n 54

55 (R

M)

1980

Po

Echi

noch

loa

mur

icat

a va

r. m

icro

stac

hya

Rou

gh b

arny

ard

gras

s G

5T5

N

X

O

Nel

son

5069

(RM

) 19

79

Po

Elym

us c

anad

ensi

s va

r. ca

nade

nsis

N

oddi

ng w

ild ry

e

[C

anad

a w

ild ry

e]

G5T

5 N

O

O

X

Lich

var 6

363

(RM

) 19

83

Po

Elym

us e

lym

oide

s va

r. el

ymoi

des

[Sita

nion

hys

trix]

Wes

tern

bot

tle-b

rush

gr

ass

[B

ottle

brus

h sq

uirre

ltail]

G5T

5 N

X

X O

C

ollin

s 68

4 (R

M)

1983

Po

Elym

us g

lauc

us

Blue

wild

rye

G5

N

X

O

Col

lins

883

(RM

)

1983

Page 48: Vascular Plant Species Checklist of Bighorn Canyon National

48

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Po

Elym

us re

pens

[Agr

opyr

on re

pens

] C

reep

ing

wild

rye

[Q

uack

gras

s]

G5

N

X

X H

eide

l 199

4 (R

M)

2001

N

oxio

us w

eed

in

Wyo

min

g.

Prev

ious

ly

repo

rted,

but

no

vouc

her

spec

imen

s lo

cate

d fo

r cor

robo

ratio

n.

Po

Elym

us tr

achy

caul

us

var.

andi

nus

Slen

der w

ild ry

e G

5T5

N

X

H

eide

l 206

1 (R

M)

2001

Th

is ta

xon

is

incl

uded

in E

. tra

chyc

aulu

s va

r. tra

chyc

aulu

s by

Ka

rtesz

and

M

each

am (1

999)

. Po

El

ymus

trac

hyca

ulus

va

r. tra

chyc

aulu

s

[A

grop

yron

tra

chyc

aulu

m]

Slen

der w

ild ry

e G

5T5

N

X

Li

chva

r 598

9 (R

M)

1983

Po

Elym

us v

illosu

s H

airy

wild

rye

G5

N

X

H

eide

l 214

0 (R

M)

2001

Po

Elym

us v

irgin

icus

Vi

rgin

ia w

ild ry

e G

5 N

X

Lich

var 6

386

(RM

) 19

83

Po

Er

agro

stis

cilia

nens

is

Stin

k gr

ass

G?

E X

O

H

eide

l 212

8 (R

M)

2001

Po

Fest

uca

arun

dina

cea

Tall

fesc

ue

G5

E X

X X

Hei

del 1

993

(RM

) 19

99

Plan

ted

and

esca

ped

at B

ICA

Po

Fest

uca

idah

oens

is

Blue

bunc

h fe

scue

[Id

aho

fesc

ue]

G5

N

X

X Li

chva

r 598

6 (R

M)

1983

Po

Fest

uca

ovin

a Sh

eep

fesc

ue

G?

E

X

Hei

del 2

111

(RM

) 20

01

Plan

ted

and

esca

ped

at

BCN

RA

Po

Gly

ceria

stri

ata

Fow

l man

na g

rass

G

5 N

X X

Col

lins

808

(RM

) 19

83

Po

H

espe

rost

ipa

com

ata

var.

com

ata

[S

tipa

com

ata

var.

com

ata]

Nee

dle-

and-

thre

ad

G5T

5 N

X

X X

Wol

f 76

(BIC

A)

1982

Po

Hor

deum

juba

tum

Fo

x-ta

il ba

rley

G5

N

X X

M

esse

rsch

mid

t 19

(BIC

A)

1981

Po

Hor

deum

x

caes

pito

sum

H

ybrid

(bet

wee

n Fo

x-ta

il an

d M

eado

w

barle

y)

G5

N

X H

eide

l 209

4 (R

M)

2001

Po

Koel

eria

mac

rant

ha

Prai

rie K

oele

r's g

rass

[Pra

irie

june

gras

s]

G5

N

O

X O

Li

chva

r 589

6 (R

M)

1983

Po

Leer

sia

oryz

oide

s R

ice

cut g

rass

G

5 N

X

Hei

del 2

151

(Rm

) 20

01

Po

Le

ptoc

hloa

fusc

a va

r. fa

scic

ular

is

[L

. fas

cicu

laris

]

Bear

ded

spra

ngle

top

G5

N

X O

O

H

eide

l 212

3 (R

M)

2001

Page 49: Vascular Plant Species Checklist of Bighorn Canyon National

49

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Po

Leuc

opoa

kin

gii

Fals

e fe

scue

[Kin

g sp

ike-

fesc

ue]

G5

N

O

O

X C

ollin

s 57

9 (R

M)

1983

Po

Leym

us c

iner

eus

[Ely

mus

cin

ereu

s]

Gre

at B

asin

lym

e gr

ass

[G

reat

Bas

in

wild

rye]

G5

N

X X

X Li

chva

r 635

8 (R

M)

1981

Po

Mel

ica

subu

lata

Al

aska

mel

ic g

rass

G

5 N

X

Col

lins

773

(RM

) 19

83

Po

M

onro

a sq

uarro

sa

Fals

e bu

ffalo

gra

ss

G5

N

X O

Hei

del 2

077

(RM

) 20

01

Po

M

uhle

nber

gia

aspe

rifol

ia

Alka

li m

uhly

G

5 N

O

X

X H

eide

l 211

7 (R

M)

1999

Po

Muh

lenb

ergi

a cu

spid

ata

Ston

y-hi

lls m

uhly

G

4 N

X

Hei

del 1

925

(MO

NTU

) 19

99

Po

Muh

lenb

ergi

a fil

iform

is

Pullu

p m

uhly

G

5 N

X

Hei

del 1

732

(MO

NTU

) 19

98

Po

Muh

lenb

ergi

a gl

omer

ata

Sp

iked

muh

ly

[M

arsh

muh

ly]

G5

N

X Li

chva

r 691

7 (B

ICA)

19

83

Orig

inal

ly

iden

tifie

d as

M.

race

mos

a Po

M

uhle

nber

gia

mex

ican

a W

irest

em m

uhly

G

5 N

X

Hei

del 2

104

(RM

) 20

01

Po

Muh

lenb

ergi

a ra

cem

osa

Gre

en m

uhly

G

5 N

X

Hei

del 2

118

(RM

) 20

01

Orig

inal

repo

rts

base

d on

a

mis

iden

tifie

d sp

ecim

en

Po

Muh

lenb

ergi

a ric

hard

soni

s M

atte

d m

uhly

G

5 N

X X

Hei

del 1

727

(MO

NTU

) 19

98

Po

Nas

sella

viri

dula

[S

tipa

virid

ula]

G

reen

tuss

ock

gras

s

[G

reen

nee

dleg

rass

] G

5 N

X X

Col

lins

676

(RM

) 19

83

A na

tive

spec

ies,

bu

t als

o cu

ltiva

ted

at B

CN

RA

Po

Pani

cum

cap

illare

C

omm

on p

anic

gra

ss

G5

N

X H

eide

l 211

0 (R

M)

2001

Po

Pani

cum

mili

aceu

m

Mille

t G

? E

O

20

01

Obs

erve

d by

H

eide

l at f

irst

pullo

ut o

n w

est

side

of h

ighw

ay in

BC

NR

A. P

lant

s pr

obab

ly n

ot

esta

blis

hed.

Po

Pa

scop

yrum

sm

ithii

[E

lym

us s

mith

ii,

Agro

pyro

n sm

ithii]

Wes

tern

whe

at g

rass

G

5 N

X O

W

olf 1

31 (B

ICA)

19

81

Po

Phal

aris

aru

ndin

acea

R

eed

cana

ry g

rass

G

5 N

X

O

X C

ollin

s 70

6 (R

M)

1983

D

uplic

ate

spec

imen

s at

BI

CA

wer

e m

isid

entif

ied

Po

Phle

um p

rate

nse

Com

mon

tim

othy

G

? E

X

O

Col

lins

810

(RM

) 19

83

Page 50: Vascular Plant Species Checklist of Bighorn Canyon National

50

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Po

Phra

gmite

s au

stra

lis

Com

mon

reed

G

5 N

X

X

Nel

son

5083

(RM

) 19

79

Po

Pi

ptat

heru

m

mic

rant

hum

[O

ryzo

psis

mic

rant

ha]

Littl

e-se

ed m

ount

ain

rice

gras

s G

5 N

X X

Col

lins

731

(RM

) 19

83

Po

Poa

annu

a An

nual

blu

e gr

ass

G5

E

X

Hei

del 1

850

(MO

NTU

) 19

99

Po

Poa

arid

a Pr

airie

blu

e gr

ass

G5

N

X Li

chva

r 691

4 (R

M)

1983

Po

Poa

bulb

osa

Bulb

ous

blue

gra

ss

G?

E

X O

Li

chva

r 557

6 (R

M)

1983

Po

Poa

com

pres

sa

Flat

-ste

m b

lue

gras

s G

? E

X

X C

ollin

s 88

4 (R

M)

1983

Po

Poa

cusi

ckii

var.

pallid

a

[P. c

usic

kii v

ar. c

usic

kii]

Cus

ick'

s bl

ue g

rass

G

5T?

N

X

X Li

chva

r 557

5 (R

M)

1983

Po

Poa

fend

leria

na

Mut

ton

gras

s G

5 N

X X

Col

lins

678

(RM

) 19

83

Po

Po

a ju

ncifo

lia v

ar.

ampl

a C

urly

blu

e gr

ass

[Big

blu

egra

ss]

G5

N

X

H

eide

l 201

6 20

01

Incl

uded

in P

oa

secu

nda

by

Karte

sz a

nd

Mea

cham

199

9)

Po

Poa

nem

oral

is s

sp.

Inte

rior

[P

oa in

terio

r]

Fore

st b

lue

gras

s

[Inte

rior b

lueg

rass

] G

5T?

N

X

X Li

chva

r 617

3 (R

M)

1983

Po

Poa

palu

stris

Fo

wl b

lue

gras

s G

5 N

X

Lich

var 5

712

(RM

) 19

83

Po

Po

a pr

aten

sis

Kent

ucky

blu

e gr

ass

G?

E O

X

X M

esse

rsch

mid

t 2

(BIC

A)

1981

Po

Poa

secu

nda

var.

elon

gata

[P

oa s

cabr

ella

]

Cur

ly b

lue

gras

s

[C

anby

blu

egra

ss]

G5

N

O

X O

Jo

nes,

Dito

lla, a

nd

Mye

rs s

.n. (

BIC

A)

1984

In

clud

ed in

Poa

se

cund

a by

Ka

rtesz

and

M

each

am 1

999)

Po

Po

a se

cund

a va

r. se

cund

a

[Poa

san

dber

gii]

Cur

ly b

lue

gras

s

[S

andb

erg

blue

gras

s]

G5

N

O

Jone

s, D

itolla

, and

M

yers

s.n

. (BI

CA)

19

84

Incl

uded

in P

oa

secu

nda

by

Karte

sz a

nd

Mea

cham

199

9)

Po

Poly

pogo

n m

onsp

elie

nsis

An

nual

rabb

it's-

foot

gr

ass

G?

E X

Nel

son

5071

(RM

) 19

79

Po

Pseu

doro

egne

ria

spic

ata

[Ely

mus

spi

catu

s,

Agro

pyro

n sp

icat

um]

Blue

bunc

h w

heat

gras

s G

5 N

X

X X

Pete

rs 1

01 B

ICA

1981

Po

Pucc

inel

lia n

utta

lliana

N

utta

ll's a

lkal

i gra

ss

G5

N

X O

Hei

del 1

982

(RM

) 20

01

Po

Sc

hedo

nnar

dus

pani

cula

tus

Tum

ble

gras

s G

5 N

X

Hei

del 2

062

(RM

) 19

98

Po

Schi

zach

yriu

m

scop

ariu

m

[And

ropo

gon

scop

ariu

s]

Littl

e fa

lse

blue

stem

G

5 N

X

Lich

var 6

894

(RM

) 19

83

Page 51: Vascular Plant Species Checklist of Bighorn Canyon National

51

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Po

Seta

ria v

iridi

s G

reen

bris

tle g

rass

G

? E

X O

O

H

eide

l 208

0 (R

M)

2001

Pr

evio

usly

re

porte

d, b

ut n

o vo

uche

r sp

ecim

ens

loca

ted

for c

orro

bora

tion.

Po

Sp

artin

a gr

acilis

Al

kali

cord

gra

ss

G5

N

X

H

eide

l 172

6 (M

ON

TU)

1998

Po

Spar

tina

pect

inat

a Fr

eshw

ater

cor

d gr

ass

G5

N

X H

eide

l 215

7 (R

M)

2001

Pr

evio

usly

re

porte

d, b

ut n

o vo

uche

r sp

ecim

ens

loca

ted

for c

orro

bora

tion.

Po

Sp

heno

phol

is

inte

rmed

ia

[S

. obt

usat

a va

r. m

ajor

]

Slen

der w

edge

scal

e G

5 N

X

Lich

var 6

378

(RM

) 19

83

Po

Sphe

noph

olis

obt

usat

a va

r. ob

tusa

ta

Slen

der w

edge

scal

e G

5 N

X

Hei

del 2

156

(RM

) 20

01

Po

Spor

obol

us a

iroid

es

Alka

li-sa

cato

n G

5 N

X

X X

Nel

son

5454

(RM

) 19

80

Po

Sp

orob

olus

cr

ypta

ndru

s Sa

nd d

rops

eed

G5

N

X

C

ollin

s 69

9 (R

M)

1983

Po

Thin

opyr

um p

ontic

um

[E

lym

us e

long

atus

, Ag

ropy

ron

elon

gatu

m]

Eura

sian

qua

ckgr

ass

G?

E X

X

Hei

del 2

056

(RM

) 20

01

Prev

ious

ly

repo

rted,

but

no

vouc

her

spec

imen

s lo

cate

d fo

r cor

robo

ratio

n.

Po

Vulp

ia o

ctof

lora

[Fes

tuca

oct

oflo

ra]

Eigh

t-flo

wer

six

-w

eeks

gra

ss

[S

ix-

wee

ks fe

scue

]

G5

N

X

X C

ollin

s 57

9 (R

M)

1983

Pole

mon

i C

ollo

mia

line

aris

N

arro

w-le

af

mou

ntai

n-tru

mpe

t

[Nar

row

leaf

col

lom

ia]

G5

N

X H

eide

l 204

9 (R

M)

2001

Pole

mon

i G

ilia

lept

omer

ia

[A

licie

lla le

ptom

eria

] Sa

nd g

ily-fl

ower

[Gre

at B

asin

gilia

] G

5 N

X

X

Dor

n 26

31 (M

ON

T)

1976

Pole

mon

i Ip

omop

sis

pum

ila

Low

sky

rock

et

[Dw

arf g

ilia]

G4

N

X X

O

Jone

s, D

itolla

, and

M

yers

s.n

. (BI

CA)

19

84

Pole

mon

i Ip

omop

sis

spic

ata

var.

orch

idac

ea

Spik

ed s

kyro

cket

[Spi

cate

gilia

] G

4?T?

N

O

X H

eide

l 202

9 (R

M)

2001

Pole

mon

i Ip

omop

sis

spic

ata

var.

spic

ata

Spik

ed s

kyro

cket

[Spi

cate

gilia

] G

4?

T4?

N

X X

Li

chva

r 604

0 (R

M)

1976

Pole

mon

i Le

ptod

acty

lon

caes

pito

sum

[L

inan

thus

ca

espi

tosu

m]

Mat

ted

pric

kly-

phlo

x G

3G4

N

X X

Li

chva

r 600

1 (R

M)

1983

Page 52: Vascular Plant Species Checklist of Bighorn Canyon National

52

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Pole

mon

i Le

ptod

acty

lon

pung

ens

[L

inan

thus

pun

gens

] G

rani

te p

rickl

y-ph

lox

G5

N

X

X Jo

nes,

Dito

lla, a

nd

Mye

rs s

.n. (

BIC

A)

1983

Pole

mon

i Li

nant

hus

sept

entri

onal

is

[L

epto

siph

on

sept

entri

onal

is]

Nor

ther

n de

sert-

trum

pet

[Nor

ther

n lin

anth

us]

G5

N

X

H

eide

l 202

0 (B

ICA)

20

01

Earli

er re

ports

ba

sed

on a

m

isid

entif

ied

spec

imen

Po

lem

oni

Phlo

x ho

odii

var.

hood

ii C

arpe

t phl

ox

[H

ood'

s ph

lox]

G

5T5

N

X X

X Li

chva

r 557

1 (R

M)

1983

Pole

mon

i Ph

lox

hood

ii va

r. m

usco

ides

[P. m

usco

ides

, P.

bryo

ides

]

Car

pet p

hlox

[Mos

s ph

lox]

G

5T4

N

X X

Li

chva

r 559

0 (R

M)

1983

Poly

gon

Erio

gonu

m a

nnuu

m

Annu

al w

ild

buck

whe

at

G5

N

X Li

chva

r 635

9 (R

M)

1983

Poly

gon

Erio

gonu

m c

ernu

um

Nod

ding

wild

bu

ckw

heat

G

5 N

X

X O

Jo

nes,

Dito

lla, a

nd

Mye

rs s

.n. (

BIC

A)

1984

Poly

gon

Erio

gonu

m fl

avum

var

. fla

vum

Al

pine

gol

den

wild

bu

ckw

heat

G

5 T4

T5

N

X X

X Jo

nes,

Dito

lla, a

nd

Mye

rs s

.n. (

BIC

A)

1983

Poly

gon

Erio

gonu

m la

gopu

s

[E. b

revi

caul

e va

r. ca

num

]

Para

sol w

ild

buck

whe

at

[R

abbi

t buc

kwhe

at]

G3

N

X X

D

orn

2772

(RM

) 19

76

Poly

gon

Erio

gonu

m o

valif

oliu

m

var.

ochr

oleu

cum

C

ushi

on w

ild

buck

whe

at

G5T

? N

X

Hei

del 1

851

(MO

NTU

) 19

99

Poly

gon

Erio

gonu

m o

valif

oliu

m

var.

purp

ureu

m

Cus

hion

wild

bu

ckw

heat

G

5T5

N

X

X Fe

rtig

1944

5 (R

M)

1983

Poly

gon

Erio

gonu

m p

auci

floru

m

var.

pauc

iflor

um

Few

-flow

er w

ild

buck

whe

at

G5T

4

N

X X

Li

chva

r 600

4 (R

M)

1983

Poly

gon

Erio

gonu

m u

mbe

llatu

m

var.

maj

us

Sulp

hur-f

low

er w

ild

buck

whe

at

G5T

5 N

X

Jone

s, D

itolla

, and

M

yers

s.n

. (BI

CA)

19

84

Poly

gon

Poly

gonu

m a

mph

ibiu

m

var.

emer

sum

W

ater

sm

artw

eed

G5T

5 N

X

X

Nel

son

7024

(RM

) 19

80

BIC

A sp

ecim

en is

ve

geta

tive

Poly

gon

Poly

gonu

m a

vicu

lare

Ya

rd k

notw

eed

G?

E X

Col

lins

689

(RM

) 19

83

Po

lygo

n Po

lygo

num

la

path

ifoliu

m

Doc

k-le

af s

mar

twee

d G

5 E

X

O

Nel

son

5058

(RM

) 19

79

Poly

gon

Rum

ex c

rispu

s C

urly

doc

k G

? E

X X

X W

olf s

.n. (

BIC

A)

1982

Poly

gon

Rum

ex m

ariti

mus

var

. fu

egin

us

Gol

den

dock

G

5 N

X

Nel

son

5057

(RM

) 19

79

Poly

gon

Rum

ex s

alic

ifoliu

s va

r. m

exic

anus

[R. s

alic

ifoliu

s va

r. tri

angu

lival

vis]

Willo

w d

ock

G5T

5 N

X

X

Hei

del 1

989

(RM

) 20

01

Poly

gon

Rum

ex v

enos

us

Vein

y do

ck

G5?

N

O

O

X

Lich

var 6

356

(RM

)

1983

Page 53: Vascular Plant Species Checklist of Bighorn Canyon National

53

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Portu

lac

Cla

yton

ia la

nceo

lata

va

r. la

nceo

lata

La

nce-

leaf

sp

ringb

eaut

y G

5T5

N

X Li

chva

r 569

5 (R

M)

1983

Portu

lac

Cla

yton

ia p

erfo

liata

var

. in

term

onta

na

[Mon

tia p

erfo

liata

]

Min

er's

lettu

ce

G5T

? N

X

Lich

var 5

690

(RM

) 19

83

Portu

lac

Lew

isia

redi

viva

O

rego

n bi

tter-r

oot

G5

N

X

Li

chva

r 592

8 (R

M)

1983

Portu

lac

Portu

laca

ole

race

a Li

ttle-

hogw

eed

[C

omm

on p

ursl

ane]

G

? E

X

H

eide

l 213

4 (B

ICA)

20

01

Earli

er re

ports

w

ere

not

vouc

here

d fo

r co

nfirm

atio

n.

Spec

ies

is n

ativ

e to

Nor

th A

mer

ica

but i

ntro

duce

d in

M

onta

na.

Pota

mog

eton

Po

tam

oget

on fo

liosu

s Le

afy

pond

wee

d G

5 N

X

Hei

del 2

126

(RM

) 20

01

Po

tam

oget

on

Stuc

keni

a fil

iform

is v

ar.

alpi

na

[P

otam

oget

on fi

lifor

mis

va

r. al

pinu

s]

Slen

der-l

eaf f

alse

po

ndw

eed

G5T

5 N

X

Col

lins

792

(RM

) 19

83

Pota

mog

eton

St

ucke

nia

pect

inat

a

[Pot

amog

eton

pe

ctin

atus

]

Sago

fals

e po

ndw

eed

G5

N

X

H

eide

l 198

1 (R

M)

2001

Prim

ul

Dod

ecat

heon

co

njug

ens

Bonn

eville

sho

otin

g-st

ar

G5

N

X

O

Hei

del 1

825

(BIC

A)

1999

Prim

ul

Dod

ecat

heon

pu

lche

llum

D

ark-

thro

at s

hoot

ing-

star

G

5 N

X

Lich

var 5

567

(RM

) 19

83

Prim

ul

[Myr

sin]

Ly

sim

achi

a ci

liata

Fr

inge

d ye

llow

-lo

oses

trife

G

5 N

X

Lich

var 6

405

(RM

) 19

83

Pter

id

[Adi

ant]

Aspi

dotis

den

sa

Indi

an's

-dre

am

G5

N

X

H

eide

l 171

4 (M

ON

TU)

1998

Pter

id

[Adi

ant]

Che

ilant

hes

feei

Sl

ende

r lip

fern

G

5 N

X

X X

Lich

var 5

616

(RM

) 19

83

Pter

id

[Adi

ant]

Pella

ea g

labe

lla s

sp.

occi

dent

alis

[P. o

ccid

enta

lis]

Smoo

th c

liffb

rake

G

5T4

N

X

Li

chva

r 594

5 (R

M)

1983

Ran

uncu

l Ac

taea

rubr

a R

ed b

aneb

erry

G

5 N

X O

C

ollin

s 66

3 (R

M)

1983

Ran

uncu

l An

emon

e cy

lindr

ica

Long

-hea

d th

imbl

ewee

d G

5 N

X

Hei

del 2

043

(RM

) 19

83

Ran

uncu

l An

emon

e m

ultif

ida

Red

win

dflo

wer

G

5 N

X X

Lich

var 5

939

(RM

) 19

83

R

anun

cul

Cer

atoc

epha

la

test

icul

ata

[Ran

uncu

lus

test

icul

atus

]

Cur

ve-s

eed

butte

rwor

t

[H

orns

eed

butte

rcup

]

G?

E

X X

Lich

var 5

716

(RM

) 19

83

Page 54: Vascular Plant Species Checklist of Bighorn Canyon National

54

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Ran

uncu

l C

lem

atis

col

umbi

ana

var.

tenu

iloba

C

olum

bian

virg

in's-

bow

er

G5?

T4

? N

X

Hei

del 2

136

(RM

) 20

01

Earli

er re

cord

(L

ichv

ar 6

457)

co

llect

ed o

utsi

de

of B

CN

RA

Ran

uncu

l C

lem

atis

ligu

stic

ifolia

D

ecid

uous

trav

eler

's-

joy

[W

este

rn

virg

in's

-bow

er]

G5

N

X X

X N

elso

n 50

74 (R

M)

1979

Ran

uncu

l D

elph

iniu

m b

icol

or v

ar.

bico

lor

Flat

head

lark

spur

G

4G5

T?

N

X

Li

chva

r 555

6 (R

M)

1983

Ran

uncu

l Pu

lsat

illa p

aten

s ss

p.

mul

tifid

a

[Ane

mon

e pa

tens

var

. m

ultif

ida]

Amer

ican

pa

sque

flow

er

G5T

? N

O

O

Col

lins

581

(RM

) 20

01

Obs

erve

d ad

jace

nt

to O

k-a-

beh

boat

ra

mp

AND

IN

Car

bon

Cou

nty

by

Hei

del i

n 20

01.

Ran

uncu

l R

anun

culu

s cy

mba

laria

Al

kali

butte

rcup

G

5 N

X

X X

Nel

son

5061

(RM

) 19

79

If va

rietie

s ar

e re

cogn

ized

, BC

NR

A m

ater

ial i

s va

r. cy

mba

laria

. R

anun

cul

Ran

uncu

lus

gmel

inii

Less

er y

ello

w w

ater

bu

tterc

up

G5

N

X

H

eide

l 171

5 (M

ON

TU)

1998

Ea

rlier

repo

rts

wer

e fro

m o

utsi

de

BCN

RA.

R

anun

cul

Ran

uncu

lus

pens

ylva

nicu

s Pe

nnsy

lvan

ia

butte

rcup

G

5 N

X

Lich

var 6

162a

(RM

) 19

83

Ran

uncu

l R

anun

culu

s sc

eler

atus

va

r. m

ultif

idus

C

urse

d bu

tterc

up

G5T

5 N

X

X

Nel

son

5060

(RM

) 19

79

Ran

uncu

l R

anun

culu

s un

cina

tus

var.

unci

natu

s W

oodl

and

butte

rcup

G

5T?

N

O

X

Lich

var 6

101

(RM

) 19

83

Ran

uncu

l Th

alic

trum

das

ycar

pum

Pu

rple

mea

dow

-rue

G5

N

X C

ollin

s 87

3 (R

M)

1983

Ran

uncu

l Th

alic

trum

fend

leri

Fend

ler's

mea

dow

-ru

e

G5

N

X C

ollin

s 77

9 (R

M)

1983

Ros

Ag

rimon

ia g

rypo

sepa

la

Tall

hairy

gro

oveb

urr

G5

N

X Li

chva

r 639

5 (R

M)

1983

Ros

Ag

rimon

ia s

triat

a W

oodl

and

groo

vebu

rr G

5 N

X

Hei

del 1

924

(MO

NTU

) 19

98

Ros

Am

elan

chie

r aln

ifolia

va

r. al

nifo

lia

Sask

atoo

n se

rvic

e-be

rry

G5T

5 N

O

O

X

Lich

var 5

667

(RM

) 19

83

Ros

C

erco

carp

us le

difo

lius

var.

ledi

foliu

s

Cur

l-lea

f mou

ntai

n-m

ahog

any

G5T

5 N

X

X X

Col

lins

541

(RM

) 19

83

Ros

C

rata

egus

chr

ysoc

arpa

[C

. col

umbi

ana

var.

chry

soca

rpa]

Fire

-ber

ry h

awth

orn

G5T

5 N

X

Hei

del 2

142

(RM

) 20

01

Page 55: Vascular Plant Species Checklist of Bighorn Canyon National

55

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Ros

D

asip

hora

frut

icos

a va

r. flo

ribun

da

[P

oten

tilla

frut

icos

a,

Pent

aphy

lloid

es

florib

unda

]

Gol

den-

hard

hack

[Shr

ubby

cin

quef

oil]

G5

N

X H

eide

l 208

6 (R

M)

2001

Ea

rlier

reco

rds

wer

e co

llect

ed

from

the

plan

tings

at

the

Love

ll vi

sito

r ce

nter

. R

os

Frag

aria

ves

ca

Woo

dlan

d st

raw

berry

G

5 N

X

Lich

var 6

191

(RM

) 19

83

R

os

Frag

aria

virg

inia

na

Virg

inia

stra

wbe

rry

G5

N

X

C

ollin

s 66

7 (R

M)

1983

Ros

G

eum

ale

ppic

um

Yello

w a

vens

G

5 N

X

Lich

var 6

168

(RM

) 19

83

R

os

Geu

m m

acro

phyl

lum

va

r. pe

rinci

sum

La

rge-

leaf

ave

ns

G5T

? N

X X

Lich

var 5

944

(RM

) 19

83

Ros

G

eum

trifl

orum

var

. tri

floru

m

Old

-man

's-w

hisk

ers

G5T

5?

N

X

X C

ollin

s 57

3 (R

M)

1983

Ros

Ke

lsey

a un

iflor

a O

ne-fl

ower

kel

seya

G

5 N

X

X

Dor

n 34

66 (R

M)

1976

Ros

Pe

troph

yton

ca

espi

tosu

m

Roc

ky M

ount

ain

rock

mat

G

4G5

N

X X

C

ollin

s 86

5 (R

M)

1983

Ros

Ph

ysoc

arpu

s m

onog

ynus

M

ount

ain

nine

bark

G

4G5

N

O

X X

Col

lins

659

(RM

) 19

83

Ros

Po

tent

illa a

rgut

a Ta

ll ci

nque

foil

G5

N

X H

eide

l 214

5 (R

M)

2001

Ros

Po

tent

illa g

raci

lis v

ar.

fast

igia

ta

[P. g

raci

lis v

ar. n

utta

llii,

P. g

raci

lis v

ar. g

labr

ata]

Gra

cefu

l cin

quef

oil

G5T

5 N

X

Col

lins

748

(RM

) 19

83

Ros

Po

tent

illa n

orve

gica

N

orw

egia

n ci

nque

foil

G5

E X

X

Ferti

g 19

435

a (R

M)

1983

Ros

Po

tent

illa o

vina

var

. ov

ina

Shee

p ci

nque

foil

G5?

T5

? N

X

Lich

var 5

657

(RM

) 19

83

Ros

Po

tent

illa p

arad

oxa

Bush

y ci

nque

foil

G5

N

X X

N

elso

n 50

66 (R

M)

1979

Ros

Po

tent

illa p

ensy

lvan

ica

Penn

sylv

ania

ci

nque

foil

G5

N

X

Li

chva

r 587

4 (R

M)

1983

Ros

Po

tent

illa ri

valis

Br

ook

cinq

uefo

il G

5 N

X

Ferti

g 19

435

b (R

M)

1998

Ros

Pr

unus

am

eric

ana

Amer

ican

plu

m

G5

N

O

X

Col

lins

746

(RM

) 19

83

R

os

Prun

us v

irgin

iana

var

. m

elan

ocar

pa

Cho

ke c

herry

G

5 N

O

X

X Li

chva

r 568

6 (R

M)

1983

Ros

R

osa

acic

ular

is v

ar.

sayi

[R

osa

sayi

]

Pric

kly

rose

G

5T5

N

X

X Li

chva

r 591

5 (R

M)

1983

Ros

R

osa

arka

nsan

a Pr

airie

rose

G

5 N

X

Hei

del 2

038

(RM

) 20

01

R

os

Ros

a w

oods

ii W

oods

' ros

e G

5 N

X O

W

olf 1

09 (B

ICA)

19

82

R

os

Rub

us id

aeus

var

. st

rigos

us

[R

. ida

eus

var.

acul

eatis

sim

us]

Com

mon

red

rasp

berry

G

5T5

N

X

X Li

chva

r 609

7 (R

M)

1983

Page 56: Vascular Plant Species Checklist of Bighorn Canyon National

56

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Ros

R

ubus

par

viflo

rus

Wes

tern

thim

ble-

berry

G

5 N

X

Hei

del 2

147

(RM

) 20

01

Ros

Sp

iraea

bet

ulifo

lia v

ar.

luci

da

Shin

y-le

af

mea

dow

swee

t G

4G5

T?

N

X

C

ollin

s 76

3 (R

M)

1983

Rub

i G

aliu

m a

parin

e va

r. ec

hino

sper

mon

St

icky

-willy

G

5 N

X X

Ferti

g 19

446

(RM

) 20

01

Rub

i G

aliu

m b

orea

le

Nor

ther

n be

dstra

w

G5

N

O

X

Lich

var 6

147

(RM

) 19

83

R

ubi

Gal

ium

trifl

orum

Fr

agra

nt b

edst

raw

G

5 N

X

Lich

var 6

098

(RM

) 19

83

Sa

lic

Popu

lus

alba

W

hite

pop

lar

G5

E X

X

Lich

var 6

046

(RM

) 19

83

Sa

lic

Popu

lus

angu

stifo

lia

Nar

row

leaf

co

ttonw

ood

G5

N

X

Li

chva

r 587

6 (R

M)

1983

Salic

Po

pulu

s ba

lsam

ifera

ss

p. tr

icho

carp

a

[Pop

ulus

bal

sam

ifera

va

r. ca

lifor

nica

]

Bals

am p

opla

r G

5T5

N

X

X Fe

rtig

1944

4 (R

M)

2001

Salic

Po

pulu

s de

ltoid

es s

sp.

mon

olife

ra

[P

opul

us d

elto

ides

var

. de

ltoid

es v

ar.

occi

dent

alis

]

East

ern

cotto

nwoo

d G

5 T4

T5

N

X X

X N

elso

n 70

25 (R

M)

1980

Salic

Po

pulu

s tre

mul

oide

s Q

uaki

ng a

spen

G

5 N

X

Lich

var 5

961

(RM

) 19

83

Sa

lic

Popu

lus

x ac

umin

ata

Lanc

elea

f cot

tonw

ood

HYB

N

X

X X

Lich

var 5

995

(RM

) 19

82

Sa

lic

Salix

am

ygda

loid

es

Peac

h-le

af w

illow

G

5 N

X

X X

Nel

son

7026

(RM

) 19

80

Sa

lic

Salix

beb

bian

a G

ray

willo

w

G5

N

X H

eide

l 170

9 (M

ON

TU)

1983

Salic

Sa

lix d

rum

mon

dian

a D

rum

mon

d's

willo

w

G4G

5 N

X X

Lich

var 5

702

(RM

) 19

83

Sa

lic

Salix

exi

gua

var.

exig

ua

Nar

row

leaf

willo

w

G5T

5 N

X

X O

N

elso

n 70

27 (R

M)

1980

Salic

Sa

lix fr

agilis

C

rack

willo

w

G?

E X

Col

lins

709

(RM

) 19

83

Sa

lic

Salix

lute

a

[S

. erio

ceph

ala

var

wat

soni

i]

Yello

w w

illow

G

4G5

N

X

H

eide

l 215

2 (R

M)

2001

Ea

rlier

repo

rts h

ad

been

bas

ed o

n m

isid

entif

ied

spec

imen

s Sa

ntal

C

oman

dra

umbe

llata

va

r. pa

llida

Bast

ard

toad

flax

G5T

5 N

X

X O

Li

chva

r 556

1 (R

M)

1983

Saxi

frag

Con

imite

lla w

illiam

sii

Willi

ams'

mite

rwor

t G

3?

N

X

C

ollin

s 66

1 (R

M)

1983

D

uplic

ate

of

Col

lins

661

at

BIC

A w

as

mis

iden

tifie

d as

M

itella

st

auro

peta

la

Saxi

frag

Heu

cher

a pa

rvifo

lia

Li

ttlel

eaf a

lum

root

G

5 N

X O

Li

chva

r 588

3 (R

M)

1983

Page 57: Vascular Plant Species Checklist of Bighorn Canyon National

57

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Saxi

frag

Lith

ophr

agm

a pa

rvifl

orum

Pr

airie

woo

dlan

dsta

r G

5 N

X

Lich

var 5

560

(RM

) 19

83

Saxi

frag

Mite

lla s

taur

opet

ala

Side

-flow

er B

isho

p's-

cap

G5

N

X

O

Col

lins

661

(BIC

A)

1983

D

uplic

ate

of

Col

lins

661

at R

M

was

iden

tifie

d as

C

onim

itella

w

illia

msi

i Sa

xifra

g Su

llivan

tia h

apem

anii

var.

hape

man

ii H

apem

an's

coo

lwor

t G

3T3

N

X

X Li

chva

r 587

1 (R

M)

1976

Saxi

frag

Tele

soni

x he

uche

rifor

mis

[Boy

kini

a he

uche

rifor

mis

]

Fals

e sa

xifra

ge

G4

N

X

Li

chva

r 593

0 (R

M)

1983

Saxi

frag

[Par

nass

i] Pa

rnas

sia

palu

stris

var

. m

onta

nens

is

Mar

sh G

rass

-of-

Parn

assu

s G

5T?

N

X H

eide

l 208

7 (R

M)

1983

Scro

phul

ari

Bess

eya

wyo

min

gens

is

Wyo

min

g co

rald

rops

G

5 N

X O

Li

chva

r 566

0 (R

M)

1983

Scro

phul

ari

Cas

tille

ja a

ngus

tifol

ia

var.

angu

stifo

lia

Nor

thw

este

rn In

dian

-pa

intb

rush

G

5T?

N

X

Li

chva

r 555

7 (R

M)

1982

Scro

phul

ari

Cas

tille

ja li

narii

folia

W

yom

ing

Indi

an-

pain

tbru

sh

G5

N

X

O

19

83

Obs

erve

d by

H

eide

l in

Big

Hor

n C

o., M

T in

200

1.

Scro

phul

ari

Cas

tille

ja m

inor

ssp

. m

inor

[C

. exi

lis]

Alka

li In

dian

-pa

intb

rush

G

5 N

X

Hei

del 2

130

(RM

) 20

01

Scro

phul

ari

Cas

tille

ja s

essi

liflo

ra

Gre

at P

lain

s In

dian

-pa

intb

rush

G

5 N

X O

Jo

nes,

Dito

lla, a

nd

Mye

rs s

.n. (

BIC

A)

1984

Scro

phul

ari

Col

linsi

a pa

rvifl

ora

Smal

l-flo

wer

blu

e-ey

ed m

ary

G5

N

X

X C

ollin

s 58

3 (R

M)

1983

Scro

phul

ari

Mim

ulus

gut

tatu

s Se

ep m

onke

y-flo

wer

G

5 N

X

Lich

var 6

170

(RM

) 19

83

Sc

roph

ular

i O

rthoc

arpu

s lu

teus

G

olde

n-to

ngue

ow

l-cl

over

G

5 N

X

Jone

s, D

itolla

, and

M

yers

s.n

. (BI

CA)

19

84

Scro

phul

ari

Pens

tem

on a

ridus

St

iff-le

af b

eard

tong

ue

G5?

N

X

Lich

var 5

951

(RM

) 19

83

Sc

roph

ular

i Pe

nste

mon

eria

nthe

rus

var.

eria

nthe

rus

Fuzz

y-to

ngue

be

ardt

ongu

e G

4T4

N

X X

D

orn

2554

(M

ON

T?)

1976

R

epor

ted

by D

orn

(197

8)

Scro

phul

ari

Pens

tem

on la

ricifo

lius

var.

laric

ifoliu

s La

rch-

leaf

be

ardt

ongu

e G

4 T3

T4

N

O

X

Lich

var 6

197

(RM

) 19

83

Scro

phul

ari

Pens

tem

on n

itidu

s va

r. ni

tidus

W

axy-

leaf

be

ardt

ongu

e G

5T?

N

X

X M

esse

rsch

mid

t s.n

. (B

ICA)

19

81

Scro

phul

ari

Pens

tem

on ra

dico

sus

Mat

-root

bea

rdto

ngue

G

5 N

X X

Lich

var 5

987

(RM

) 19

83

Sc

roph

ular

i Ve

rbas

cum

thap

sus

Gre

at m

ulle

in

G?

E

X

Hei

del 2

107

(RM

) 19

83

Sc

roph

ular

i Ve

roni

ca a

mer

ican

a Am

eric

an b

rook

lime

G5

N

O

X

Col

lins

745

(RM

) 19

83

Sc

roph

ular

i Ve

roni

ca a

naga

llis-

aqua

tica

Blue

wat

er s

peed

wel

l G

5 E

X

N

elso

n 50

59 (R

M)

1979

Page 58: Vascular Plant Species Checklist of Bighorn Canyon National

58

Fam

ily

(__a

ceae

) Sp

ecie

s N

ame

[s

ynon

yms]

C

omm

on N

ame

Glo

bal

rank

N

ativ

e/

exot

ic

WY

- B

ig

Hor

n C

o M

T-

Car

bon

Co

MT-

B

ig

Hor

n C

o Sa

mpl

e Vo

uche

r Ye

ar fi

rst

colle

cted

C

omm

ents

Scro

phul

ari

Vero

nica

per

egrin

a va

r. xa

lape

nsis

N

eckw

eed

G5T

5 N

X O

Li

chva

r 615

7 (R

M)

1983

Sela

gine

ll Se

lagi

nella

den

sa

Den

se s

pike

-mos

s G

5 N

O

X Li

chva

r 565

6 (R

M)

1983

Smila

c

[Lili]

Sm

ilax

lasi

oneu

ra

[S

. her

bace

a va

r. la

sion

euro

n]

Blue

Rid

ge c

arrio

n-flo

wer

G

5 N

O

X Li

chva

r 614

6 (R

M)

1983

Sola

n Ly

cium

bar

baru

m

Mat

rimon

y-vi

ne

G?

E X

Hei

del 2

009

(RM

) 20

01

So

lan

Phys

alis

het

erop

hylla

C

lam

my

grou

nd-

cher

ry

G5

N

X

C

ollin

s 83

0 (R

M)

1983

Sola

n Ph

ysal

is lo

ngifo

lia

Long

-leaf

gro

und-

cher

ry

G5

N

X X

H

eide

l 207

5 (R

M)

2001

Sola

n So

lanu

m d

ulca

mar

a C

limbi

ng n

ight

shad

e G

? E

O

X X

Lich

var 6

130

(RM

) 19

83

So

lan

Sola

num

trifl

orum

Cut

-leav

ed

nigh

tsha

de

G5

N

X

H

eide

l 208

3 (R

M)

2001

Tam

aric

Ta

mar

ix c

hine

nsis

Fi

ve-s

tam

en ta

mar

isk

[S

alt-c

edar

] G

? E

X X

O

M. L

. Rob

erts

463

1 (R

M)

1976

N

oxio

us w

eed

in

WY.

Ty

ph

Typh

a an

gust

ifolia

N

arro

w-le

af c

at-ta

il G

5 N

X

Dor

n 70

86 (R

M)

1996

So

me

early

re

ports

wer

e fro

m

Love

ll vi

sito

rs

cent

er

Typh

Ty

pha

latif

olia

Br

oad-

leaf

cat

-tail

G5

N

X

O

Nel

son

7035

(RM

) 19

80

U

lm

Ulm

us p

umila

Si

beria

n el

m

G?

E X

X O

Kn

ight

K-1

2 (R

M)

1983

Pl

ante

d as

an

orna

men

tal

Ulm

[C

eltid

] C

eltis

occ

iden

talis

C

omm

on h

ackb

erry

G

5 N

X

X

Lich

var 6

066

(RM

) 19

83

Plan

ted

as a

n or

nam

enta

l U

rtic

Parie

taria

pen

sylv

anic

a Pe

nnsy

lvan

ia p

ellit

ory

G5

N

O

X X

Col

lins

634

(RM

) 19

83

U

rtic

Urti

ca d

ioic

a ss

p.

grac

ilis

[U

. d. v

ar. p

roce

ra]

Stin

ging

net

tle

G5T

5 N

O

X

X H

eide

l 209

1 (B

ICA)

19

83

Vale

rian

Vale

riana

dio

ica

var.

sylv

atic

a M

arsh

val

eria

n G

5 T4

T5

N

O

X

Col

lins

670

(RM

) 19

83

Verb

en

Verb

ena

brac

teat

a C

arpe

t ver

vain

G

4G5

N

X O

O

Li

chva

r 602

6 (R

M)

1982

Verb

en

Verb

ena

hast

ata

Sim

pler

's-jo

y G

5 N

O

X W

olf 1

22 (B

ICA)

19

82

Vi

ol

Viol

a ad

unca

H

ook-

spur

vio

let

G5

N

X Li

chva

r 590

1 (R

M)

1983

Viol

Vi

ola

cana

dens

is

Can

adia

n w

hite

vio

let

G5

N

X

X C

ollin

s 76

2 (R

M)

1983

Viol

Vi

ola

nutta

llii

Yello

w p

rairi

e vi

olet

G

5 N

X

Lich

var 6

186

(RM

) 19

83

Vi

ol

Viol

a pa

lust

ris

Alpi

ne m

arsh

vio

let

G5

N

X

H

eide

l 182

6 (B

ICA)

19

99

Vi

ol

Viol

a va

llicol

a Va

lley

viol

et

G4Q

N

X

Lich

var 5

558

(RM

) 19

83

Vi

sc

Arce

utho

bium

cy

anoc

arpu

m

Lim

ber p

ine

dwar

f-m

istle

toe

G4

N

X

H

eide

l 206

8 (R

M)

2001

Vit

Vitis

ripa

ria

Riv

er-b

ank

grap

e G

5 N

X

Col

lins

742

(RM

) 19

83

Za

nnic

helli

Zann

iche

llia p

alus

tris

Hor

ned

pond

wee

d G

5 N

X

X

Hei

del 2

078

(RM

) 20

01

Zy

goph

yll

Trib

ulus

terre

stris

Pu

nctu

revi

ne

G?

E

X

Hei

del 2

073

(RM

) 20

01

Page 59: Vascular Plant Species Checklist of Bighorn Canyon National

Appendix B.

Rejected Taxa

The following vascular plant species have been reported from Bighorn Canyon National Recreation Area based on collections at the RM and BICA herbaria and reports from Lichvar et al. (1985), Knight et al. (1987), Akashi (1988), and Heidel and Fertig (2000), but are now known to be either misidentified or collected outside the NRA. The list does not include species that are known by a different synonym (synonyms are listed in Appendix A). Family (__aceae)

Species Name Common Name Comments

Api Ligusticum filicinum Fern-leaf wild lovage Cited in Lichvar et al. (1985), but specimen misidentified.

Api Ligusticum porteri Porter's wild lovage Historical collection by Booth cited in Lichvar et al. (1985), but specimen misidentified.

Api Lomatium cous Cous-root desert-parsley Cited in Lichvar et al. (1985), but specimen from outside BCNRA.

Ast Antennaria media [A. alpina var. media]

Rocky Mountain pussytoes [Alpine pussytoes]

Cited in Lichvar et al. (1985), but specimen from outside BCNRA.

Ast Antennaria parvifolia Little-leaf pussytoes Cited in Lichvar et al. (1985), but specimen misidentified (A. rosea)

Ast Arnica rydbergii Subalpine leopardbane Cited in Lichvar et al. (1985), but specimen misidentified (A. fulgens).

Ast Artemisia annua Annual wormwood Report in Heidel and Fertig (2000) based on a misidentified specimen.

Ast Chrysothamnus viscidiflorus var. latifolius

Green rabbitbrush Cited in Lichvar et al. (1985), but this variety does not occur in Montana or Wyoming. This name may have been misapplied to specimens of C. linifolius.

Ast Erigeron compositus var. discoideus

Dwarf mountain fleabane [Cutleaf fleabane]

Cited in Lichvar et al. (1985), but all labeled specimens are E. allocotus. An additional unlabled Lichvar collection cannot be reliably placed within BCNRA.

Ast Erigeron linearis Desert yellow fleabane Reported from a plot by Knight et al. (1987), but voucher is misidentified.

Ast Erigeron speciosus Aspen fleabane Cited in Lichvar et al. (1985), but specimen misidentified.

Ast Helianthella quinquenervis Nodding dwarf-sunflower Cited in Lichvar et al. (1985), but specimen from outside BCNRA.

Ast Hieracium cynoglossoides Hound-tongue hawkweed Cited in Lichvar et al. (1985), but specimen misidentified.

Ast Malacothrix torreyi Torrey's desert-dandelion Cited in Lichvar et al. (1985), but specimen misidentified.

Ast Packera plattensis [Senecio plattensis]

Prairie groundsel Cited in Lichvar et al. (1985), but specimen misidentified.

Ast Packera subnuda [Senecio cymbalarioides, included in Packera streptanthifolia by Kartesz and Meacham 1999)

Rocky Mountain groundsel Report in Heidel and Fertig (2000) based on misidentified specimen.

Ast Solidago simplex var. nana [S. spathulata var. nana]

Mt. Albert goldenrod Cited in Lichvar et al. (1985), but specimen from outside BCNRA.

Ast Symphyotrichum falcatum [Aster falcatus, variety not indicated]

Rough white prairie American aster

Cited in Lichvar et al. (1985), but specimen probably misidentified (Aster ericoides)

Ast Symphyotrichum foliaceum var. parryi [Aster foliaceus var. parryi]

Alpine leafy-head American aster

Cited in Lichvar et al. (1985), but no voucher specimen has been

Ast Tetradymia spinosa Short-spine horsebrush Report in Heidel and Fertig (2000) based on an observation by Heidel just outside BCNRA boundaries in 1999.

Aster Cirsium eatonii [C. tweedyi]

Eaton's thistle Earlier reports were based on specimens collected outside of BCNRA.

Page 60: Vascular Plant Species Checklist of Bighorn Canyon National

60

Family (__aceae)

Species Name Common Name Comments

Boragin Cryptantha caespitosa Tufted cat's-eye Report from Knight et al. (1987) is based on misidentified specimens of C. cana.

Boragin Mertensia viridis [included in M. oblongifolia by Kartesz and Meacham 1999]

Green bluebells Cited in Lichvar et al. (1985), but specimen probably from outside BCNRA (single voucher is unlabeled and precise location cannot be determined).

Boragin Onosmodium molle var molle

Soft-hair marbleseed Reported by Lichvar et al. (1985), but probably represents var. occidentale. Var. molle is endemic to the southeastern United States.

Brassic Arabis hirsuta Hairy rockcress Cited in Lichvar et al. (1985), but specimen from outside BCNRA.

Brassic Arabis williamsii var. saximontana [A. pendulocarpa var. saximontana, A. microphylla var. s., Boechera williamsii var. s.]

Wind River rockcress Cited in Lichvar et al. (1985), but specimen misidentified (A. holboellii var. pendulocarpa)

Brassic Draba crassifolia Snowbed whitlow-grass Cited in Lichvar et al. (1985), but specimen misidentified (D. praealta and D. albertina).

Brassic Smelowskia calycina var. americana

Fern-leaf false candytuft Reported by Lichvar et al. (1985), but no voucher found for corroboration; locality probably outside of BCNRA.

Caprifoli Lonicera utahensis Rocky Mountain honeysuckle

Cited in Lichvar et al. (1985), but specimen misidentified.

Caprifoli [Adox]

Sambucus racemosa var. melanocarpa

Red elder Cited in Lichvar et al. (1985), but specimen misidentified (Rhus glabra)

Caryophyll Minuartia obtusiloba [Arenaria obtusiloba]

Alpine stitchwort Cited in Lichvar et al. (1985), but specimen from outside BCNRA.

Chenopodi Chenopodium album Lamb's-quarters Specimen from BCNRA (Wolf 146) is immature and probably represents C. berlandieri.

Chenopodi Kochia americana [Bassia americana]

Greenmolly Cited in Lichvar et al. (1985), but specimen misidentified.

Commelin Tradescantia bracteata Long-bract spiderwort Cited in Lichvar et al. (1985), but probably misidentified (T. occidentalis).

Cyp Carex canescens Hoary sedge Reported by Heidel and Fertig (2000) based on a misidentification.

Cyp Carex raynoldsii Raynold's sedge Cited in Lichvar et al. (1985), but specimen from outside BCNRA.

Cyp Carex scopulorum Holm's Rocky Mountain sedge

Mistakenly reported in Heidel and Fertig (2000).

Dryopterid [Aspleni]

Polystichum lonchitis Northern holly-fern Cited in Lichvar et al. (1985), but specimen from outside BCNRA.

Fab Astragaluls bourgovii Bourgov's milkvetch Report in Heidel and Fertig (2000) based on misidentified specimen.

Fab Cytisus scoparius Scotch broom Report from Heidel and Fertig (2000) is based on a misidentified specimen at BICA.

Fab Hedysarum sulphurescens Yellow sweet-vetch Cited in Lichvar et al. (1985), but specimen from outside BCNRA.

Fab Lupinus sericeus Pursh's silky lupine Cited in Lichvar et al. (1985), but specimen misidentified.

Fab Oxytropis besseyi var. besseyi

Bessey's locoweed Cited in Lichvar et al. (1985), but specimen misidentified (O. lagopus).

Fab Oxytropis deflexa (variety not indicated)

Pendant-pod locoweed Cited in Lichvar et al. (1985), but specimen misidentified.

Hippurid Hippuris vulgaris Common mare's-tail Report from Heidel and Fertig (2000) based on a misidentified specimen.

Irid Sisyrinchium angustifolium Narrow-leaf blue-eyed grass

Cited in Lichvar et al. (1985), but specimen misidentified (S. montanum).

Junc Juncus parryi Parry's rush Cited in Lichvar et al. (1985), but specimen from outside BCNRA.

Junc Juncus tenuis var. tenuis Poverty rush Report from Lichvar et al. (1985) is based on material of J. dudleyi.

Lili [Alli]

Allium brevistylum Short-stem onion Cited in Lichvar et al. (1985), but specimen from outside BCNRA.

Page 61: Vascular Plant Species Checklist of Bighorn Canyon National

61

Family (__aceae)

Species Name Common Name Comments

Onagr Gayophytum ramossimum Pinyon groundsmoke Report from Heidel and Fertig (2000) based on a misidentified specimen.

Pa Alopecurus pratensis Field meadow-foxtail Report from Heidel and Fertig (2000) based on a misidentified specimen.

Po Calamagrostis purpurascens

Purple reed grass Cited in Lichvar et al. (1985), but specimen from outside BCNRA.

Po Echinochloa crus-galli Large barnyard grass Cited in Lichvar et al. (1985), but specimen probably misidentified (E. muricata).

Po Elymus scribneri [Agopyron scribneri]

Scribner's wildrye Report from Heidel and Fertig (2000) based on a misidentified specimen.

Po Poa alpina Alpine bluegrass Cited in Lichvar et al. (1985), but specimen from outside BCNRA.

Po Poa glaucifolia [included in Poa arida by Kartesz and Meacham, 1999)

Prairie bluegrass Cited in Lichvar et al. (1985), but specimen misidentified.

Po Trisetum spicatum Narrow false oat [Spike trisetum]

Cited in Lichvar et al. (1985), but specimens either misidentified or from outside BCNRA.

Polemoni Phlox multiflora Rocky Mountain phlox Cited in Lichvar et al. (1985), but specimens from outside BCNRA.

Polemoni Polemonium viscosum Sticky Jacob's-ladder Cited in Lichvar et al. (1985), but specimens from outside BCNRA.

Polygon Eriogonum mancum Imperfect wild buckwheat [Mancos wild buckwheat]

Report from Heidel and Fertig (2000) based on a misidentified specimen.

Polygon Eriogonum ovalifolium var ovalifolium

Cushion wild bukwheat Cited in Lichvar et al. (1985), but specimen misidentified.

Polygon Polygonum achoreum Leathery knotweed Cited in Lichvar et al. (1985), but specimen misidentified.

Polygon Polygonum bistortoides American bistort Cited in Lichvar et al. (1985), but specimens from outside BCNRA.

Ranuncul Ranunculus macounii Macoun's buttercup Cited in Lichvar et al. (1985), but specimens from outside BCNRA.

Ranuncul Thalictrum occidentale Western meadow-rue Cited in Lichvar et al. (1985), but specimen misidentified.

Ros Ivesia gordonii Alpine mousetail Cited in Lichvar et al. (1985), but specimens from outside BCNRA.

Ros Physocarpus malvaceus Mallow-leaf ninebark Cited in Lichvar et al. (1985), but specimen misidentified.

Ros Potentilla biennis Biennial cinquefoil Cited in Lichvar et al. (1985), but specimens from outside BCNRA.

Ros Potentilla diversifolia var. diversifolia

Mountain meadow cinquefoil

Cited in Lichvar et al. (1985), but specimens from outside BCNRA.

Ros Potentilla fissa Big-flower cinquefoil Cited in Lichvar et al. (1985), but specimen misidentified (P. paradoxa)

Salic Salix boothii Booth's willow Cited in Lichvar et al. (1985), but specimen misidentified (S. bebbiana)

Salic Salix monticola Park willow Cited in Lichvar et al. (1985), but specimen misidentified (S. bebbiana)

Saxifrag Saxifraga rhomboidea Diamond-leaf saxifrage Cited in Lichvar et al. (1985), but specimens from outside BCNRA.

Scrophulari Castilleja angustifolia var. dubia [C. chromosa. C. applegatei var. martinii in Kartesz and Meacham, 1999]

Wavy-leaf Indian-paintbrush

Cited in Lichvar et al. (1985), but specimen misidentified.

Scrophulari Castilleja miniata Great red Indian-paintbrush Cited in Lichvar et al. (1985), but specimens from outside BCNRA.

Scrophulari Castilleja pulchella Showy Indian-paintbrush Cited in Lichvar et al. (1985), but specimens from outside BCNRA.

Scrophulari Pedicularis cystopteridifolia Fern-leaf lousewort Cited in Lichvar et al. (1985), but specimens from outside BCNRA.

Scrophulari Penstemon caryi Cary's beardtongue Cited in Lichvar et al. (1985), but specimens from outside BCNRA.

Scrophulari Penstemon humilis Low beardtongue Cited in Lichvar et al. (1985), but specimen misidentified.

Page 62: Vascular Plant Species Checklist of Bighorn Canyon National

62

Family (__aceae)

Species Name Common Name Comments

Scrophulari Penstemon procerus [P. confertus var. procerus]

Pincushion beardtongue Cited in Lichvar et al. (1985), but specimens from outside BCNRA.

Valerian Valeriana edulis Tobacco-root Cited in Lichvar et al. (1985), but specimens from outside BCNRA.

Page 63: Vascular Plant Species Checklist of Bighorn Canyon National

63

Appendix C.

Unconfirmed Taxa

The following vascular plant species have been reported from Bighorn Canyon National Recreation Area based on reports from Lichvar et al. (1985), Knight et al. (1987), Akashi (1988), and Heidel and Fertig (2000), but these reports cannot be confirmed due to a lack of authenticated voucher material at the RM or BICA herbaria. None of these species could be relocated during field surveys of BCNRA in 2001. Family (__aceae)

Species Name Common Name Comments

Api Lomatium simplex var. simplex [L. triternatum ssp. platycarpum]

Great Basin desert-parsley Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey

Aster Antennaria racemosa Hooker's pussytoes Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey

Aster Oreostemma alpigena var. haydenii [Aster alpigenus]

Tundra mountaincrown [Hayden's alpine aster]

Reported by Heidel in Heidel and Fertig (2000), but population could not be relocated in 2001 survey for verification.

Aster Picradeniopsis oppositifolia Opposite-leaf false-bahia Cited in Lichvar et al. (1985) and reported from Barry's Landing in Lichvar's 1983 field notes. No voucher has been found and this population was not relocated in 2001 survey.

Aster Senecio serra var. serra Tall ragwort Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey

Aster Symphyotrichum ascendens [Aster ascendens, A. chilensis var. ascendens]

Western American aster Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey

Aster Symphyotrichum foliaceum var. parryi [Aster foliaceus var. parryi]

Alpine leafy-head American aster

Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Aster Tetraneuris acaulis var. caespitosa [Hymenoxys acaulis var. caespitosa]

Stemless four-nerve daisy Lichvar (5907) is an unlabeled specimen of this taxon that may be within BCNRA (exact location cannot be determined and Lichvar's original field book has been lost).

Brassic Arabis lignifera [Boechera lignifera]

Owens Valley rockcress Reported by Dorn (1978), but specimen and presence within BCNRA cannot be verified. Other reports are based on misidentifications.

Brassic Cardaria chalepensis Lens-pod hoary-cress Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey. Considered a noxious weed in Montana.

Brassic Draba nemorosa Woodland whitlow-grass Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Brassic Sisymbrium loeselii False London rocket Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Campanul Triodanis leptocarpa Slim-pod Venus'-looking-glass

Reported by Lichvar et al. (1985) from the north end of BCNRA, but no voucher has been located for corroboration.

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Family (__aceae)

Species Name Common Name Comments

Campanul Triodanis perfoliata Clasping-leaf Venus'-looking-glass

Reported by Lichvar et al. (1985) from the north end of BCNRA, but no voucher has been located for corroboration.

Caryophyll Silene latifolia ssp. alba [Lychnis alba]

Bladder-campion Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Cyp Carex utriculata [C. rostrata]

Northwest Territories sedge [Beaked sedge]

Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Euphorbi Euphorbia cyparissias Cypress spurge Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Fab Amorpha canescens Leadplant Mentioned as characteristic of BCNRA Great Plains flora in Lichvar et al. (1985) but not cited in checklist and not located in 2001 survey.

Fab Astragalus kentrophyta (variety not indicated)

Spiny milkvetch Reports by Akashi (1988) cannot be confirmed with voucher specimens.

Fab Trifolium pratense Red clover Cited in Lichvar et al. (1985) and Akashi (1988) but no voucher specimens found for corroboration and not recorded in 2001 survey.

Gentian Frasera speciosa [Swertia radiata]

Monument-plant [Green gentian]

Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Junc Juncus compressus Round-fruit rush Reported by Knight et al. (1987), but no voucher specimen found for confirmation and not recorded in 2001 survey.

Junc Juncus regelii Regel's rush Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Lami Stachys palustris Woundwort [Swamp hedge-nettle]

Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Lentibulari Utricularia macrorhiza [U. vulgaris, U. major]

Greater bladderwort Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Lili [Antheric]

Leucocrinum montanum Star-lily Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey. Lichvar's 1983 field notes indicate an observation along one BCNRA's access roads, but exact location is unclear.

Onagr Camissonia andina Blackfoot River suncup Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Orchid Calypso bulbosa Fairy-slipper orchid Mentioned as characteristic of BHCNRA Rocky Mt flora in Lichvar et al. (1985), but not cited in checklist and not recorded in 2001 surveys.

Po Elymus lanceolatus var. riparius [Agropyron riparium; included in var. lanceolatus by Kartesz and Meacham, 1999; ]

Streamside wild rye Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

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Family (__aceae)

Species Name Common Name Comments

Po Hordeum pusillum Little barley Reported by Heidel in Heidel and Fertig (2000) based on an unvouchered observation in 1999. Population could not be relocated for confirmation in 2001.

Po x Elyhordeum macounii [Elymus x macounii]

Possibly observed in BCNRA by Heidel, but confirmation needed.

Polemoni Gilia pinnatifida [Aliciella pinnatifida]

Sticky gily-flower Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Polemoni Gilia tweedyi Tweedy's gily-flower Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Polemoni Polemonium occidentale Western Jacob's-ladder Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Polemoni Polemonium pulcherrimum Beautiful Jacob's-ladder Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Polygon Eriogonum caespitosum Matted wild buckwheat Reported in Knight et al. (1987), but no voucher specimen available for confirmation.

Pyrol [Eric]

Orthilia secunda [Pyrola secunda]

Sidebells Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Pyrol [Eric]

Pyrola asarifolia Pink wintergreen Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Ranuncul Delphinium geyeri Geyer's larkspur Reported in Heidel and Fertig (2000) based on a Booth specimen at MONTU that remains to be verified. Exact location of collection needs to be determined.

Ranuncul Myosurus apetalus [M. aristatus]

Bristly mousetail Unlabeled specimen from Lichvar may be from vicinity of BCNRA.

Ros Argentina anserina [Potentilla anserina]

Common silverweed Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Ros Crataegus douglasii Black hawthorn Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Rubi Galium trifidum Three-petal bedstraw Cited by Lichvar et al. (1985), but no vocuher specimens found for corroboration. Heidel made a tentative observation at T8S R29E S27 in 2001.

Scrophulari Mimulus suksdorfii Miniature monkeyflower Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

Scrophulari Penstemon glaber var. glaber

Western smooth beardtongue

Cited in Lichvar et al. (1985), but no voucher specimens found for corroboration and not recorded in 2001 survey.

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Appendix D.

Potential Vascular Plant Flora of Bighorn Canyon National Recreation Area

The following list contains species that are known from the vicinity of Bighorn Canyon National Recreation Area in Montana and Wyoming, but which have not yet been documented in the NRA. Species are considered of “High” probability of occurrence in the study area if they are globally common and occur in habitat types that are well-represented within the NRA. Species of “Medium”probability are less abundant or occur in habitats that are uncommon within the area. “Low” probability taxa are rare species or occur in habitats that are rare or found just outside the park’s boundaries. See Appendix A for other codes and definitions. Family (__aceae)

Species Name Common Name Native/ exotic

Probability Comments

Amaranth Amaranthus powellii Green amaranth N High Found in adjacent areas of Big Horn County, WY

Api Lomatium cous Cous-root desert-parsley

N Low Most likely at higher elevations in Carbon County.

Api Lomatium simplex var. simplex [L. triternatum ssp. platycarpum]

Great Basin desert-parsley

N Low Potential habitat in foothills of Pryor Mountains in Carbon County.

Aster Antennaria dimorpha Cushion pussytoes N Medium Potential habitat in foothills of Pryor Mountains and in north unit.

Aster Antennaria media [A. alpina var. media]

Rocky Mountain pussytoes [Alpine pussytoes]

N Low Known from vicinity of BCNRA. Potential habitat most likely at high elevation sites in Carbon County.

Aster Antennaria parvifolia Little-leaf pussytoes N High Potential habitat in foothlls of Pryor Mountains and in north unit.

Aster Antennaria racemosa Hooker's pussytoes N Medium Potential habitat in coniferous forests in Carbon County.

Aster Arnica mollis Cordilleran leopardbane

N Low Potential habitat in coniferous forests in Carbon County.

Aster Crepis occidentalis var. costata

Large-flower hawk's-beard

N Low Most likely at moderate elevations in Big Horn County WY & Big Horn County, MT.

Aster Erigeron compositus var. discoideus

Dwarf mountain fleabane [Cutleaf fleabane]

N Medium Known from vicinity of BCNRA. Potential habitat most likely at medium to high elevation sites.

Aster Erigeron divergens Spreading fleabane N Medium Most likely at moderate elevations in Big Horn County WY & Big Horn County, MT.

Aster Erigeron engelmannii Engelmann's fleabane

N Medium Most likely at moderate elevations in Big Horn County WY & Big Horn County, MT.

Aster Erigeron peregrinus var. callianthemus

Subalpine fleabane N Low Most likely at higher elevations in Carbon County.

Aster Erigeron speciosus Aspen fleabane N High Previous reports were misidentified, but potential habitat is present at higher elevations in Carbon County.

Aster Eurybia integrifolia [Aster integrifolius]

Thick-stem wood-aster

N Low Most likely at higher elevations in Carbon County.

Aster Eurybia merita [Aster sibiricus var. meritus, A. meritus]

Behring wood-aster N Low Most likely at higher elevations in Carbon County.

Aster Helianthella quinquenervis

Nodding dwarf-sunflower

N Medium Previous reports were misidentified, but potential habitat is present at higher elevations in Carbon County.

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Family (__aceae)

Species Name Common Name Native/ exotic

Probability Comments

Aster Madia glomerata Mountain tarplant N High Weedy native could occur along roads and trails over much of the higher elevations of BCNRA.

Aster Malacothrix torreyi Torrey's desert-dandelion

N Medium Found in adjacent areas of Big Horn County, WY

Aster Oreostemma alpigena var. haydenii [Aster alpigenus]

Tundra mountaincrown [Hayden's alpine aster]

N Medium Observed in vicinity of BCNRA in Montana by Heidel in 1999, but population could not be relocated in 2001.

Aster Picradeniopsis oppositifolia

Opposite-leaf false-bahia

N High Reported from Barry's Landing by Lichvar, but population could not be confirmed. Boat ramp development may have eliminated suitable habitat.

Aster Senecio serra var. serra

Tall ragwort N High Most likely at higher elevations in Carbon County.

Aster Senecio triangularis Arrow-leaf ragwort N Medium Most likely at higher elevations in Carbon County.

Aster Solidago simplex var. nana [S. spathulata var. nana]

Mt. Albert goldenrod N Low Most likely at higher elevations in Carbon County.

Aster Stenotus acaulis [Haplopappus acaulis]

Stemless mock goldenweed

N High Most likely at moderate elevations in Big Horn County WY & Big Horn County, MT.

Aster Symphyotrichum ascendens [Aster ascendens, A. chilensis var. ascendens]

Western American aster

N High Potential habitat throughout lower elevation areas of BCNRA.

Aster Symphyotrichum ericoides var. stricticaule [Aster ericoides var. stricticaulis]

White heath American-aster

N Medium Potential habitat in northern unit (Big Horn County, MT).

Aster Symphyotrichum foliaceum var. parryi [Aster foliaceus var. parryi]

Alpine leafy-head American aster

N Medium Most likely at higher elevations in Carbon County.

Aster Symphyotrichum spathulatum [Aster occidentalis]

Mountain American aster

N Low Most likely at higher elevations in Carbon County.

Aster Tetradymia spinosa Short-spine horsebrush

N High Observed just outside BNCRA boundaries by Heidel in 1999.

Aster Tetraneuris acaulis var. caespitosa [Hymenoxys acaulis var. caespitosa]

Stemless four-nerve daisy

N High An unlabeled collection by Lichvar is probably from the general vicinity of BCNRA.

Boragin Cryptantha caespitosa Tufted cat's-eye N Medium Reported from Big Horn County in Dorn (2001), but these may represent misidentified C. cana.

Boragin Lappula cenchrusoides Great Plains sheepburr

N Medium Most likely at moderate elevations in Big Horn County WY & Big Horn County, MT.

Boragin Mertensia viridis [included in M. oblongifolia by Kartesz and Meacham 1999]

Green bluebells N Medium Known from vicinity of BCNRA. Potential habitat most likely at medium to high elevation sites.

Boragin Plagiobothrys scouleri var. hispidulus

Meadow popcorn-flower

N High Potential habitat along Yellowtail Reservoir in Bighorn County, WY and Big Horn County, MT.

Brassic Arabis drummondii [Boechera angustifolia]

Canadian rockcress N High Most likely at higher elevations in Carbon County.

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Family (__aceae)

Species Name Common Name Native/ exotic

Probability Comments

Brassic Arabis hirsuta Hairy rockcress N Medium Most likely at higher elevations in Carbon County.

Brassic Arabis lignifera [Boechera lignifera]

Owens Valley rockcress

N Low Reported by Dorn (1978) from vicinity of BCNRA, but specimen cannot be relocated and may be misidentified. Potential habitat at lower elevations in Bighorn County, WY and Carbon County.

Brassic Cardaria chalepensis Lens-pod hoary-cress

E Medium Potential habitat along roads and trails at lower elevations.

Brassic Lepidium densiflorum var. densiflorum

Miner's pepperwort N Medium Most likely at moderate elevations in Big Horn County WY & Big Horn County, MT.

Callitrich Callitriche palustris Vernal water-starwort

N Medium Potential habitat along margins of Yellowtail Reservoir.

Campanul Triodanis leptocarpa Slim-pod Venus'-looking-glass

N High Reported by Lichvar et al. (1985) from northern end of BCNRA.

Campanul Triodanis perfoliata Clasping-leaf Venus'-looking-glass

N High Reported by Lichvar et al. (1985) from northern end of BCNRA.

Caryophyll Arenaria congesta var. lithophila [Eremogene congesta var. lithophila]

Ball-head sandwort N High Observed in 2001 on east flanks of Pryor Mountain, just outside the NRA.

Caryophyll Moehringia lateriflora [Arenaria lateriflora]

Blunt-leaf groove-sandwort

N High Potential habitat in Big Horn County, MT on slopes above dam.

Caryophyll Silene drummondii Drummond's catchfly

N Medium Most likely at higher elevations in Carbon County.

Caryophyll Silene latifolia ssp. alba [Lychnis alba]

Bladder-campion E Medium No voucher - high prob.

Caryophyll Spergularia rubra Ruby sand-spurry E High Potential habitat along margins of Yellowtail Reservoir.

Chenopodi Chenopodium foliosum [C. capitatum var. parvicapitatum, C. overi]

Leafy goosefoot N High Most likely at higher elevations in Carbon County.

Chenopodi Chenopodium simplex Giant-seed goosefoot

N Low Most likely at moderate elevations in Big Horn County WY & Big Horn County, MT.

Chenopodi Endolepis suckleyi [Atriplex suckleyi]

Suckley's saltbush N High Potential habitat in shale barren habitats in Big Horn County, WY.

Chenopodi Suaeda calceoliformis Paiuteweed N Medium Potential habitat in shale-clay wetland habitats in Big Horn County, WY.

Cyper Carex phaeocephala Mountain hare sedge

N Medium Most likely at higher elevations in Carbon County.

Cyper Carex raynoldsii Raynold's sedge N Low Most likely at higher elevations in Carbon County.

Cyper Carex utriculata [C. rostrata]

Northwest Territories sedge [Beaked sedge]

N High Potential habitat along Yellowtail Reservoir in Bighorn County, WY and Big Horn County, MT.

Dryopterid [Aspleni]

Polystichum lonchitis Mountain holly fern N Low Known from vicinity of BCNRA. Potential habitat most likely at high elevation sites in Carbon County.

Dryopterid [Aspleni]

Woodsia scopulina Rocky Mountain cliff fern

N Medium Potential habitat in northern unit (Big Horn County, MT).

Eric Vaccinium scoparium Grouseberry [Grouse whortleberry]

N Low Most likely at higher elevations in Carbon County.

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Family (__aceae)

Species Name Common Name Native/ exotic

Probability Comments

Fab Astragalus alpinus Alpine milkvetch N Medium Most likely at higher elevations in Carbon County.

Fab Astragalus eucosmus Elegant milkvetch N Low Most likely at higher elevations in Carbon County.

Fab Astragalus kentrophyta var. tegetarius

Spiny milkvetch N Medium Potential habitat in foothlls of Pryor Mountains and in north unit.

Fab Astragalus tenellus Loose-flower milkvetch

N High Potential habitat in northern unit (Big Horn County, MT).

Fab Astragalus vexilliflexus Bent-flower milkvetch

N Medium Potential habitat in northern unit (Big Horn County, MT).

Fab Hedysarum occidentale Western sweet-vetch

N Low Potential habitat in northern unit (Big Horn County, MT).

Fab Hedysarum sulphurescens

Yellow sweet-vetch N Low Known from vicinity of BCNRA. Potential habitat most likely at medium to high elevation sites.

Fab Lupinus sericeus Pursh's silky lupine N Medium Most likely at moderate elevations in Big Horn County WY & Big Horn County, MT.

Fab Oxytropis sericea var. sericea

Rocky Mountain locoweed

N Medium Most likely at moderate elevations in Big Horn County WY & Big Horn County, MT.

Fab Trifolium pratense Red clover E High Most likely at low elevation disturbed sites in Big Horn County, WY and Big Horn County, MT.

Fab Trifolium repens White clover E High Most likely at low elevation disturbed sites in Big Horn County, WY and Big Horn County, MT.

Gentian Frasera speciosa [Swertia radiata]

Monument-plant [Green gentian]

N Medium Most likely at medium to high elevations in Montana counties.

Gentian Gentianella amarella var. amarella

Autumn dwarf-gentian

N Medium Potential habitat in northern unit (Big Horn County, MT).

Gerani Geranium richardsonii White crane's-bill N High Potential habitat in northern unit (Big Horn County, MT).

Halorag Myriophyllum sibiricum Siberian water-milfoil N High Potential habitat along Yellowtail Reservoir in Bighorn County, WY and Big Horn County, MT.

Hydrophyll Nemophila breviflora Great Basin baby-blue-eyes

N Medium Most likely at higher elevations in Carbon County.

Isoet Isoetes bolanderi Bolander's quillwort N Low Potential habitat in northern unit (Big Horn County, MT).

Junc Juncus compressus Round-fruit rush E High Potential habitat along Yellowtail Reservoir in Bighorn County, WY and Big Horn County, MT.

Junc Juncus confusus Colorado rush E Medium Known from vicinity of BCNRA. Potential habitat most likely at medium elevation sites.

Junc Juncus parryi Parry's rush N Low Known from vicinity of BCNRA. Potential habitat most likely at high elevation sites in Carbon County.

Junc Luzula parviflora Small-flower wood-rush

N Medium Most likely at higher elevations of Montana counties.

Lami Prunella vulgaris Common self-heal N Medium Most likely at higher elevations of Montana counties.

Lami Stachys palustris Woundwort [Swamp hedge-nettle]

N High Most likely at higher elevations of Montana counties.

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Family (__aceae)

Species Name Common Name Native/ exotic

Probability Comments

Lentibulari Utricularia macrorhiza [U. vulgaris, U. major]

Greater bladderwort N Medium Potential habitat along Yellowtail Reservoir in Bighorn County, WY and Big Horn County, MT.

Lili [Antheric]

Leucocrinum montanum

Star-lily N Medium Potential habitat occurs at medium to high elevation areas, mostly in Montana counties.

Lili [Alli]

Allium brevistylum Short-stem onion N Medium Most likely at higher elevations of Montana counties.

Lili [Calochort]

Streptopus amplexicaulis

Clasping twisted-stalk

N Medium Most likely at higher elevations in Carbon County.

Loas Mentzelia dispersa Nevada blazing-star N High Potential habitat in northern unit (Big Horn County, MT).

Onagr Camissonia andina Blackfoot River suncup

N High Most likely at moderate elevations in Big Horn County WY & Big Horn County, MT.

Onagr Gayophytum diffusum ssp. parviflorum [G. diffusum var. strictipes]

Spreading groundsmoke

N High Most likely at moderate elevations in Big Horn County WY & Big Horn County, MT.

Po Alopecurus aequalis Short-awn meadow-foxtail

N High Potential habitat along Yellowtail Reservoir in Bighorn County, WY and Big Horn County, MT.

Po Calamagrostis canadensis

Bluejoint [Bluejoint reedgrass]

N Medium Potential habitat along Yellowtail Reservoir in Bighorn County, WY and Big Horn County, MT.

Po Calamagrostis purpurascens

Purple reed grass N Medium Known from vicinity of BCNRA. Potential habitat most likely at high elevation sites in Carbon County.

Po Danthonia intermedia Timber wild oatgrass N Medium Most likely at higher elevations in Carbon County.

Po Echinochloa crus-galli Large barnyard grass

E Low Most likely at low elevation disturbed sites in Big Horn County, WY and Big Horn County, MT.

Po Elymus lanceolatus var. lanceolatus

Streamside wild rye N High Potential habitat throughout BCNRA.

Po Eremopyrum triticeum [Agropyron triticeum]

Annual false wheatgrass

E Medium Most likely at moderate elevations in Big Horn County WY & Big Horn County, MT.

Po Hordeum pusillum Little barley N Medium Most likely at moderate elevations in Big Horn County WY & Big Horn County, MT.

Po Phleum alpinum Mountain timothy N Low Most likely at higher elevations in Carbon County.

Po Poa alpina Alpine bluegrass N Low Known from vicinity of BCNRA. Potential habitat most likely at high elevation sites in Carbon County.

Po Poa wheeleri [P. nervosa var. wheeleri]

Wheeler's bluegrass N Medium Most likely at higher elevations in Carbon County.

Po Trisetum spicatum Narrow false oat [Spike trisetum]

N Low Known from vicinity of BCNRA. Potential habitat most likely at high elevation sites in Carbon County.

Po x Elyhordeum macounii [Elymus x macounii]

N High Possibly observed in BCNRA by Heidel, but confirmation needed.

Polemoni Gilia pinnatifida [Aliciella pinnatifida]

Sticky gily-flower N High Most likely at moderate elevations in Big Horn County WY & Big Horn County, MT.

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Family (__aceae)

Species Name Common Name Native/ exotic

Probability Comments

Polemoni Gilia tweedyi Tweedy's gily-flower N High Most likely at moderate elevations in Big Horn County WY & Big Horn County, MT.

Polemoni Ipomopsis congesta var. congesta

Ballhead skyrocket N Medium Most likely at moderate elevations in Big Horn County WY & Big Horn County, MT.

Polemoni Phlox gracilis ssp. humilis [Microsteris gracilis var. humilior]

Slender phlox N Medium Most likely at moderate elevations in Montana counties.

Polemoni Phlox multiflora Rocky Mountain phlox

N Medium Most likely at higher elevations of Montana counties.

Polemoni Polemonium occidentale

Western Jacob's-ladder

N Low Most likely at moderate elevations in Big Horn County, MT.

Polemoni Polemonium pulcherrimum

Beautiful Jacob's-ladder

N Low Most likely at moderate elevations in Big Horn County, MT.

Polemoni Polemonium viscosum Sticky Jacob's-ladder

N Low Known from vicinity of BCNRA. Potential habitat most likely at high elevation sites in Carbon County.

Polygon Eriogonum caespitosum

Matted wild buckwheat

N Low Most likely at moderate elevations in Big Horn County, MT.

Polygon Polygonum achoreum Leathery knotweed N High Most likely at low elevation disturbed sites in Big Horn County, WY and Big Horn County, MT.

Polygon Polygonum douglasii var. douglasii

Douglas' knotweed N High Potential habitat in northern unit (Big Horn County, MT).

Polygon Polygonum polygaloides var. confertiflorum [P. kelloggii var. confertiflorum]

White-margin knotweed

N Medium Potential habitat in northern unit (Big Horn County, MT).

Polygon Rumex stenophyllus Narrow-leaf dock E Medium Most likely at low elevation disturbed sites in Big Horn County, WY and Big Horn County, MT.

Portulac Lewisia pygmaea Alpine bitterroot N Low Most likely at higher elevations of Montana counties.

Primul Androsace septentrionalis var. subulifera

Pygmy-flower rock-jasmine

N High Most likely at higher elevations of Montana counties.

Primul [Myrsin]

Glaux maritima Sea-milkwort N High Potential habitat in saline wetlands along Yellowtail Reservoir.

Pyrol [Eric]

Orthilia secunda [Pyrola secunda]

Sidebells N Medium Most likely at higher elevations in Carbon County.

Pyrol [Eric]

Pyrola asarifolia Pink wintergreen N Low Most likely at higher elevations in Carbon County.

Ranuncul Clematis hirsutissima Sugarbowls N Medium Most likely at higher elevations of Montana counties.

Ranuncul Delphinium geyeri Geyer's larkspur N Medium Known from vicinity of BCNRA in Montana from an historical collection by Booth. Potential habitat also occurs in Big Horn County, WY.

Ranuncul Myosurus apetalus [M. aristatus]

Bristly mousetail N Medium Potential habitat along Yellowtail Reservoir in Bighorn County, WY and Big Horn County, MT.

Ranuncul Ranunculus flammula var. filiformis

Greater creeping spearwort

N Medium Potential habitat along Yellowtail Reservoir in Bighorn County, WY and Big Horn County, MT.

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Family (__aceae)

Species Name Common Name Native/ exotic

Probability Comments

Ranuncul Ranunculus glaberrimus var. ellipticus

Sagebrush buttercup N Medium Most likely at higher elevations of Montana counties.

Ranuncul Ranunculus macounii Macoun's buttercup N Medium Most likely at higher elevations of Montana counties.

Rhamn Ceanothus velutinus Tobacco-brush [Buckbrush]

N Low Most likely at higher elevations of Montana counties.

Ros Argentina anserina [Potentilla anserina]

Common silverweed N High Potential habitat along Yellowtail Reservoir in Bighorn County, WY and Big Horn County, MT.

Ros Crateagus douglasii Black hawthorn N High Potential habitat in northern unit (Big Horn County, MT).

Ros Ivesia gordonii Alpine mousetail N Medium Most likely at higher elevations in Carbon County.

Ros Potentilla biennis Biennial cinquefoil N High Potential habitat along Yellowtail Reservoir in Bighorn County, WY and Big Horn County, MT.

Ros Potentilla concinna var. concinna

Red cinquefoil N Medium Most likely at higher elevations of Montana counties.

Ros Potentilla diversifolia var. diversifolia

Mountain meadow cinquefoil

N Low Known from vicinity of BCNRA. Potential habitat most likely at high elevation sites in Carbon County.

Ros Potentilla glandulosa var. pseudorupestris

Sticky cinquefoil N Medium Most likely at higher elevations of Montana counties.

Ros Potentilla pulcherrima [P. gracilis var. pulcherrima]

Soft cinquefoil N Medium Most likely at higher elevations of Montana counties.

Rubi Galium bifolium Twin-leaf bedstraw N High Potential habitat in semi-disturbed areas throughout BCNRA

Rubi Galium trifidum Three-petal bedstraw

N High Potential habitat along Yellowtail Reservoir in Bighorn County, WY and Big Horn County, MT.

Ruppi Ruppia cirrhosa [R. maritima]

Spiral ditch-grass N Low Potential habitat along Yellowtail Reservoir in Bighorn County, WY and Big Horn County, MT.

Saxifrag Saxifraga rhomboidea Diamond-leaf saxifrage

N Low Known from vicinity of BCNRA. Potential habitat most likely at high elevation sites in Carbon County.

Scrophulari Castilleja cusickii Cusick's Indian-paintbrush

N Low Most likely at higher elevations of Montana counties.

Scrophulari Castilleja miniata Great red Indian-paintbrush

N Low Known from vicinity of BCNRA. Potential habitat most likely at high elevation sites in Carbon County.

Scrophulari Penstemon caryi Cary's beardtongue N Medium Voucher from outside BCNRA Scrophulari Penstemon eriantherus

var. cleburnei Fuxxy-tongue beardtongue

N Low Potential habitat in low elevation areas of Big Horn County, WY.

Scrophulari Penstemon glaber var. glaber

Western smooth beardtongue

N Medium Most likely at higher elevations of Montana counties.

Scrophulari Penstemon procerus [P. confertus var. procerus]

Pincushion beardtongue

N Medium Known from vicinity of BCNRA. Potential habitat most likely at high elevation sites in Carbon County.

Scrophulari Veronica serpyllifolia var. humifusa

Thyme-leaf speedwell

N Medium Most likely at higher elevations of Montana counties.

Solan Hyoscyamus niger Black henbane E Medium Potential habitat in semi-disturbed areas throughout BCNRA

Page 73: Vascular Plant Species Checklist of Bighorn Canyon National

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Family (__aceae)

Species Name Common Name Native/ exotic

Probability Comments

Valerian Valeriana edulis Tobacco-root N Low Known from vicinity of BCNRA. Potential habitat most likely at high elevation sites in Carbon County.

Viol Viola praemorsa ssp. liguifolia [V. praemorsa var. altior]

Upland yellow violet N Medium Most likely at higher elevations of Montana counties.