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Aestivation
The mode of arrangement of sepals and petals in a floral bud is known as aestivation.
There are five types of aestivation:
i) Valvate: the unit of a whorl i.e. sepal or petal contacts with each other by their
margins but does not overlap each other.
ii) Twisted: the sepals or petals are overlapped each other at the margin
iii) Imbricate: Among the five sepals or petals one is inside, one is outside
and remaining is overlapped at the margin.
iv) Quincuncial: Among the five petals or sepals, two are inside, two are outside and
remaining is overlapped at the margin.
v) Vexillary: among the five petals, the posterior or largest one covers the two lateral
and the latter two covers the interior two. This type of aestivation is commonly found
in the member of Fabaceae family.
Placentation
Placenta: A parenchymatous outgrowth-in the inner wall of the ovary to which the ovule or
ovules are attached.
Placentation: The manner in which the placentae are distributed in the cavity of the ovary is
known as placentation
Types of placentation
i) Marginal- The ovary is one chambered and the placenta develops along the
junction of the two margins of the carpel called the ventral suture e.g.
Fabaceae.
ii) Axile-The ovary is two or many chambered- usually as many as the number of
carpels –and the placentae bearing the ovules develop from central axis e.g.
lemon, orange, china rose, lady’s finger, tomato, potato etc.
iii) Parietal- the ovary is one chambered and the placentae bearing the ovules
develop on the inner wall of the ovary e.g. papaya
iv) Central- The septa or portion walls in the
young ovary soon break down so that ovary becomes one chambred and
placenta bearing the ovules develops from the central axis e.g. Dianthus
v) Free central- The placenta arises from the base of the ovary, projects far into
its cavity as a swollen central axis and bears the ovule all over its surface e.g.
Primula
vi) Basal- In basal placentation, the ovary is one chambered and the placenta
develops directly on the thalamus, bearing the single ovule at the base of the
ovary e.g. rice.
vii) Superficial- the ovary is multichamber, the carpels being numerous, as in
axile placentation. But the placentae developed all round the inner surface of
the partition walls as in water lily.
Androecium
The androcium composed of male parts of flower which is essential whorl for sexual
reproduction. It is composed of stamens and the male flower is termed as staminate flower. Each
stamen is composed of three parts viz. anther connective and filament.
Didynamous
When the stamens are unequal i.e. two are longer and two are shorter.
Tetradynamous
When four stamens are longer and two stamens are shorter e.g. mustard
Monadelphous
The filaments of all the stamens are united to form a tube like structure but the anthers are remaining free e.g. cotton, okra, China rose etc.
Diadelphous
The filaments of all the stamens are united into two bundles e.g. pea, soybean etc.
Polyadelphous
The filaments of all the stamens are united into many bundles and anthers are remaining free e.g. citrus.
Synandrous
The stamens of the androecium are united throughout the whole body i.e. anther and filaments are united e.g. member of Cucurbitaceae family.
Syngenesious
The anthers of stamens are fused and united but the filaments remain free e.g. sunflower, pumkin etc.
Adnate
When the filament is attached throughout the whole length of the anther i.e. from the base to the top e.g. Nicotiana, Magnolia etc.
Basifexed
When the filament is fixed to the base of the anther e.g. Brassica
Dorsifixed
When the filament of stamen is fixed with the dorsal site of the stamen e.g. Sesbania, Annona etc.
Versatile
When the filament is attached with the anther in a single point forming the anther as pendulous e.g. rice.
Gynoecium
It is the female reproductive part of flower and essential whorl of flower for reproductive process. It is composed of pistil or carpel.
Pistil
It is the unit of gynoecium having three parts viz. stigma, style and ovary. It is also termed as megasporophyll.
Hypogynous
Ovary is situated above the thallamus but other whorl remains at the base of thalamus. It is called as superior ovary.
Floral formula
Floral formula means symbol, number or alphabet for expressing the structure and information of flower. It can represent or express the particular structure of a species or generalize the structure of taxa.
Floral diagram
It is a graphic representation of flower structure. It shows the number of floral organs, their arrangement and fusion. Floral diagram is useful for identification of plant and help in understanding the evolution of plant.
Symbols Used in Floral Formulae
Parameter Symbol
Bracteate Br
Ebracteate Ebr
Bracteolate Brl
Ebracteolate Ebrl
Actinomorphic
Zygomorphic %
Bisexual
Male flower
Female flower ♀Epicalyx EpK
Calyx K
Corolla C
Androecium A
Gynoecium G
Ovary superior G
Ovary inferior G
Many ∞