a new species of draba in the kananaskis range of southwestern alberta

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NOTES Validation of the moss name Atuichrcm abecuistatu~n (Ren. & Card.) Irel. ROBERT R. IRELAND Natior~nl Mrtseutns of Cannrla, Mrwerrtn of Nnturnl Scietrces, Ottui~.n, Crrtmda Received October 3, 1969 IRELAND, R. R. 1970. Validation of the nioss namc Atriclrltm allecristatrrrrl (Ren. & Card.) Irel. Can. J. Bot. 48: 1897. Dr. Gary L. Smith, New York Botanical Garden, recently informed me that my new com- bination, Atrickrrrn altecristatum (Ren. & Card.) Irel. (Can. J. Bot. 47(3): 353. 1969), is illegitimate according to Article 11 of the International Rules of Botanical Nomenclature. A species described in 1904, Cntharinen tnollis Holz., was listed in the syilonomy of A. altecristatwn; therefore, 1 failed to use the oldest name within rank. Dr. Smith further stated that he did not believe that C. n7ollis should be considered a synonym of A. altecristnturn. After a reexamination of an isotype of C. nzoNis in the New York Botanical Garden, I now agree with this opinion. I would like to make the name A. altecristatunz legitimate by citing the synonymy as I now understand it. Atrichurn altecristatrrm (Ren. & Card.) Irel. comb. nov., Atriclrurn urzdzrlatun? var. ctltecrista- turn Ren. & Card. (Bot. Gaz. 15: 58. 1890), Catharinen urrdulnta var. alleghe~riensis Jenn. (Man. Mosses W. Pennsylv. 188. 1913), Atriclzurn undulatun7 var. crllegher7ier7se (Jenn.) Jenn. (Man. Mosses W. Pennsylv. Ed. 2, 162. 1951). A new species of Draba in the Kananaskis Range of southwestern Alberta' GERALD A. MULLIGAN Plnnt Research Itrstitrtte, Carradn Department of Agricrtltrlre, Ottatva, Cai~ndr~ Received May 22, 1970 MULLIGAN, G. A. 1970. A new species of Draba in the Kananaskis Range of southwestern Alberta. Can. J. Bot. 48: 1897-1898. A new yellow-flowered species of Draba, D. kananaskis G. A. Mulligan, 11 = 32, is described from the Kananaskis Range in southwestern Alberta. It is suggested that it may be an endemic in an area unglaciated during the Wisconsin. Draba kananaskis G. A. Mulligan, sp. nov. Plantae perennes laxe caespitosae. Folia basilaria integra vel denticulata obovata, 5-20 mm longa, 2-9 mm lata, gracile petiolata; paginae pilis stellatis plerumque 4-radiis subsessilibus sparse praeditae; margines pilis simplicibus furcatis et stellatis instructae. Caules floriferi 3-20 cm alti, pilis simplicibus furcatis et stellatis vestiti, 1-2 foliis ovatis vel anguste ovatis praediti. Pedicelli siliculis breviores. Sepala ca. lPlant Research Institute Contribution No. 777. 2 mm longa, glabra vel pilis paucis simplicibus instructa. Petala flava, spathulato-oblanceolata, 3.54.0 mm longa; siliculae anguste ovatae, 6-10 mm longa, 2-3 mm lata, pilis paucis parvis simplicibus instructa; styli 0.3-0.5 mm longi. Semina 20-24, ca. 1.3 mm longa. Plant perennial, loosely tufted; basal leaves entire to denticulate, obovate, 5-20 mm long, 2-9 mm broad with slender petioles; leaf sur- faces with scattered, nearly sessile, mostly 4-rayed hairs; leaf margins with simple, forked and stellate hairs; flowering stems 3-20 cm tall Can. J. Bot. Downloaded from www.nrcresearchpress.com by Depository Services Program on 11/16/14 For personal use only.

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Page 1: A new species of               Draba               in the Kananaskis Range of southwestern Alberta

NOTES

Validation of the moss name Atuichrcm abecuistatu~n (Ren. & Card.) Irel.

ROBERT R. IRELAND Natior~nl Mrtseutns of Cannrla, Mrwerrtn of Nnturnl Scietrces, Ottui~.n, Crrtmda

Received October 3, 1969

IRELAND, R. R. 1970. Validation of the nioss namc Atriclrltm allecristatrrrrl (Ren. & Card.) Irel. Can. J. Bot. 48: 1897.

Dr. Gary L. Smith, New York Botanical Garden, recently informed me that my new com- bination, Atrickrrrn altecristatum (Ren. & Card.) Irel. (Can. J. Bot. 47(3): 353. 1969), is illegitimate according to Article 11 of the International Rules of Botanical Nomenclature. A species described in 1904, Cntharinen tnollis Holz., was listed in the syilonomy of A. altecristatwn; therefore, 1 failed to use the oldest name within rank. Dr. Smith further stated that he did not believe that C. n7ollis should be considered a synonym of A. altecristnturn. After a reexamination of an isotype

of C. nzoNis in the New York Botanical Garden, I now agree with this opinion. I would like to make the name A. altecristatunz legitimate by citing the synonymy as I now understand it.

Atrichurn altecristatrrm (Ren. & Card.) Irel. comb. nov., Atriclrurn urzdzrlatun? var. ctltecrista- turn Ren. & Card. (Bot. Gaz. 15: 58. 1890), Catharinen urrdulnta var. alleghe~riensis Jenn. (Man. Mosses W. Pennsylv. 188. 1913), Atriclzurn undulatun7 var. crllegher7ier7se (Jenn.) Jenn. (Man. Mosses W. Pennsylv. Ed. 2, 162. 1951).

A new species of Draba in the Kananaskis Range of southwestern Alberta'

GERALD A. MULLIGAN Plnnt Research Itrstitrtte, Carradn Department of Agricrtltrlre, Ottatva, Cai~ndr~

Received May 22, 1970

MULLIGAN, G. A. 1970. A new species of Draba in the Kananaskis Range of southwestern Alberta. Can. J. Bot. 48: 1897-1898.

A new yellow-flowered species of Draba, D. kananaskis G. A. Mulligan, 11 = 32, is described from the Kananaskis Range in southwestern Alberta. It is suggested that it may be an endemic in an area unglaciated during the Wisconsin.

Draba kananaskis G. A. Mulligan, sp. nov. Plantae perennes laxe caespitosae. Folia

basilaria integra vel denticulata obovata, 5-20 mm longa, 2-9 mm lata, gracile petiolata; paginae pilis stellatis plerumque 4-radiis subsessilibus sparse praeditae; margines pilis simplicibus furcatis et stellatis instructae. Caules floriferi 3-20 cm alti, pilis simplicibus furcatis et stellatis vestiti, 1-2 foliis ovatis vel anguste ovatis praediti. Pedicelli siliculis breviores. Sepala ca.

lPlant Research Institute Contribution No. 777.

2 mm longa, glabra vel pilis paucis simplicibus instructa. Petala flava, spathulato-oblanceolata, 3 .54.0 mm longa; siliculae anguste ovatae, 6-10 mm longa, 2-3 mm lata, pilis paucis parvis simplicibus instructa; styli 0.3-0.5 mm longi. Semina 20-24, ca. 1.3 mm longa.

Plant perennial, loosely tufted; basal leaves entire to denticulate, obovate, 5-20 mm long, 2-9 mm broad with slender petioles; leaf sur- faces with scattered, nearly sessile, mostly 4-rayed hairs; leaf margins with simple, forked and stellate hairs; flowering stems 3-20 cm tall

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Page 2: A new species of               Draba               in the Kananaskis Range of southwestern Alberta

1898 CANADIAN JOURNAL O F BOTANY. VOL. 48, 1970

with few simple to forked and stellate hairs and with one or two ovate to narrowly ovate leaves; pedicels shorter than silicles; sepals ca. 2 inm long, glabrous or with few simple hairs; petals yellow, spathulate-oblanceolate, 3.5-4.0 mm long; silicles narrowly ovate, 6-10 inn1 long, 2-3 mm broad, with few simple hairs, 20-24 seeded, seeds ca. 1.3 mm long; styles 0.3-0.5 mm long.

TYPE: Near Snow Ridge Ski Resort, 25 mi south of highway 1 on Kananaskis-Coleman Road, eastern slope of Kananaskis Range, about 50'48' N, 1 15'12' W, altitude 7250 ft, n = 32, G. A. & D. G. Mulligan 3477, Aug. 8, 1969 (holotype, DAO).

Draba lcananaslcis is normally self-fertilized in nature and is octoploid with the common base number of the genus, x = 8. Some strictly North American species of Draba have higher base numbers (Mulligan 1966; Bocher 1966). This species is characterized by its yellow flowers, one or two stem leaves, loosely tufted rosette, and nearly sessile cruciform hairs (Fig. 1) on the surface of the basal leaves. It is not particularly closely related to any other species of Draba.

FIG. 1. Nearly sessile cruciform hairs of D. lcnt~n~lnskis, X 85.

Draba kanar~aslcis is known to occur only on the eastern slope of the Kananaskis Range a t the location where the type specimen was col- lected. It is not included in A Revision of the Drabas of Western Nort11 America (Hitchcock 1941) or the Vasculal- Plants of the Pac~fic Northrvest (Hitchcock et a/. 1964), both of which cover the area in which this taxon occurs. Since specimens of this species were not found in the herbaria of the Canada Department of Agricul- ture, Ottawa (DAO), National Museum of Canada, Ottawa (CAN), University of Washing- ton, Seattle (WTU), or Oregon State College, Corvallis (OSC), it can be concluded that it is not a common species. It does not seem to have been collected before. Relatively easy access to the mountain slope where the type of D. lcar~anaskis was collected was only made possible recently when a road was built into a new ski area. Since collecting of alpine communi- ties in this area is much limited because of the absence of roads, it may be some time before the total range of D. kat~anaskis is determined.

Draba lcnnanaslcis is found in the rain shadow of the Rocky Mountains at a location just north and west of an area that is shown as unglaciated during the Wisconsin by V. K. Prest on his 1969 map of the Retreat of Wisconsin and recent ice in North America (map 1257A, Geological Survey of Canada). It is quite possible that D. kananaslcis may occur in an arca which was uilglaciated during the Wisconsin and that it may be an endemic with a very rcstricted distribution.

BOCHER, T. W. 1966. Experimental and cytological s t~~dies on plant species. IX. Some arctic and montane crucifers. K. Dan. V~densk. Sclsk. Biol. Skr. pp. 1-74.

HITCHCOCK, C. L. 1941. A revision of the drabas of west- ern North America. Univ. Wash. Publ. Biol. 11: 1-132.

HITCHCOCK, C. L., A. CRONQUIST, M. OWN BEY?^^^ J. W. THOMPSON. 1964. Vasc~~lar plants of the Pac~fic North- west. Part 2. Salicaceae to Saxifragaceae. Univ. Wash. Publ. Biol. 17(2): 1-597.

MULLIGAN, G. A. 1966. Chromosome numbers of the family Cruciferae. 111. Can. J. Bot. 44: 309-319.

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