UNIVERSITY OF WISCONSIN

STEVENS POINT STEVENS POINT WISCONSIN

THE MUSEUM OF NATURAL

HISTORY ATLAS OF WISCONSIN BRYOPHYTES

Report No 16

Reports on the Fauna and Flora of Wisconsin

Patt 1

ATLAS OF WISCONSIN BRYOPHYTES

November 21 1979

THE MUSEUM OF NATURAL HISTORY Stevens Point Wisconsin

Reports on the Fauna and Flora of Wisconsin No 16

Part I

Climacium dendroides (Tree moss)

ATLAS OF WISCONSIN BRYOPHYTES

By Frank D Bowers

Curator of Non-Vascular Plants-Herbarium Museum of Natural History and

Department of Biology and

Sally Kauss Freckmann Research Associate

Museum of Natural History Universit y of Wisconsin

Stevens Point Wisconsin 54481 U S A

2

CONTENTS

ABSTRACT 4

ACKNOWLEDCEMENTS 4

INTRODUCTION 6

GEOGRAPHIC CLIMATIC AND BOTANICAL fEATURES OF WISCONSIN 7

SYSTEMATIC ARLNGEMENT OF WISCONSIN BRYOPHYTES 11

MAPS OF BRYOPH) iE TAXA REPORTED FOR WISCONSIN 29

HORNWORTS AND LIVERWORTS 30

MUSCI 54

FOSSIL AND SUBFOSSIL MOSSES OF WISCONSIN 119

PALEONTOLOGICAL Rb J3RENCES 119

SYNONYMS AND EXCLUDED TAXA 120

PIONEER BRYOLOGISTS OF WISCONSIN 126

LITERATURE CITED 127

3

ABSTRACf

117This atlas of Wisconsin bryophytes shows the distribution of)30 taxa (137 hepatics 389 mosses)

on state maps with a dot in each county for which each taxon has been reported in the literature A list of synonyms and excluded taxa is included along with an extensive bibliography Pertinent features of the natural history and history of bryology in the state are discussed

ACKNOWLEDGEMENTS

We want to thank Drs Fred Hermann and John Thomson of the University of Wisconsin-Madison herbarium and Botany department for locating and lending packets and for references to aid us in accumushylating a more accurate complete report The illustration of Climacium dendroides on the cover was drawn by Robert Freckmann Of course we are grateful to all the people who have collected specimens of Wisshyconsin bryophytes and preserved them in herbariums making this distributional study possible

4

ONEIDA

LINCOLN

TAYLOR

t---------- MARATHON C1l-RK

SAu~

t

~

~ C i-W=AS_-r7-1

l=itHLANO

)

lOCK

INGTb

lt

WALWORTH

5

INTRODUCfION

This atlas of the bryophyte flora of Wisconsin is an attempt to show the distribution of all bryoshyphyte species known to occur in the state by county occurrenoes based on published reports in the literashyture It is not an annot~ted check list (only a feW problematic taxa were examined) but is a literature search intended as a foundation for a complete flora or for otherbryological research

The literature of btYo1ogy is large and spread over numerous journals reports monographs and manuals It would be almost impossible to search all sources for every reference attributing bryophyte species to Wisconsin and to particular localities in the state Wehav checked many inlportant references and paid special attention to works dealing with Wisconsiit botany and the bryologyof the upper Midwest Several bibliographies and indices were checked thoroughly including Green an~ Curtis (1955) the Bry ologist indices (Crum 1959 and Miller 1976) andt)le Wisconsin Academy of Sciences Arts and Letters indices

Only those published repots which app~red to)e substanti2ted by herbarium specimens and which gave- ~fficient location dat to P~CetheW(On in amiddot particula(county were-used in the initial compilation Species described in manuals in generalized way implying their occurrence in Wisconsin (Without menmiddot tioning Wisconsin specifically or citing a particular location in the state) were excluded eg Gnmmia raui (Wynne 1943) and Pohlia annotinll var decipiens (Crum 1976) Ecological papers which were either too general in taxonomic determinations (eg Sphagnum sp Polytrichum sp) or in location data were also excluded Some other taxa attributed to Wisconsin which have been excluded are

Cylindrothectum compr~ (Cheney 189tmiddotDane Co) identified later by Grout as Entodon seducshytrtx

Dicranell4 humilis (Holzjne~ 1925middotBuffalo Co) probablyD vllJia since Dhumilis is not found in North Arnerl~

Didymodon triarium (Foote 1 966-SW Visconsin) identified by Fred Hennann as Barbulaacuta Funoria microstoma (anonymous 1975middotDane Co) excluded because it is outside of its geographic

range and the packe~ QaMot be located Phil(motis calcarea (Cheney and Evans 1944-Dane Co) excluded in Crum Steere and Anderson

(1973) Plagiobryum demissum and P zierl (Cheney and Evans 1944-various counties) are both Arcticshy

alpine species not likely to be found in isconsin Two of Cheneys collections of Pl1giobyrum have been referred to Bryum

Pogonatum alpinum var septentrionale (Che~ey and Evans 1944-Chippewa Co) was misidentified (Ireland 1960) and is P umigerum

Rhizomnium fAPpoJachIanum and R punctalu1fl ssp chlorophyDosum (Koponen 1973middotseveral counshy ties) excluded North American bryologiflts have not recognized these taxa in America

Sphagnum tenellum (Cheney and Evans 1944-Waukesha Co)an Atlantic species not occurring in Wisconsin (Cruro 1975)

The nomenclature and systematic arrangement of hornwort 00 hepatic species in the atlas follow Stotler and Crandall-5totler (1977) and of moss species fonow Crum Steere and Anderson (1973) except in such instances as the JreptlIfockIdus fonns whiGh follow Wijk et aL (19S9middot1973) Species reported in the literature under names which are now listed as synonyms on either of these lists were converted to the accepted name A dot is placed in each county of Wisconsin from which the taxon has been reported The numbers after the name of the taxon correspond to the numbered references in the bibliography which giye ~act location data OEat least location data -s~fficient to place it in-particular counties in the state The numbers in parentheses indicate references which attribute the taxon to Wisconsin without providing exact loltation data

Fossil and subfossil reports are summarized Slfparately in our atlas only six species (Aulacomnium turgidum lJ1achythecium turgidum Ditrichum Jlelficaule Hypnum bambergti Scorpidium 6fJisities and Tortel14 1rIgi1i$) are known exclUsively from the fossil record ~

6

GEOGRAPHIC CLIMATIC AND BOTANICAL FEATURES OF WISCONSIN

Wisconsin is a north central state bounded on the east by Lake Michigan on the north by Michigan and Lake Superior on the south by Illinois and on the west by the Mississippi and St Croix rivers It is located midway between the Prime Meridian (0deg) of Longitude and 1800 Longitude and midway between the equator and the north pole thus placing it in the center of the northern half of the western hemisphere

The topography of the state is relatively flat a few hills exceed 1900 feet with Thns Hill at 1952 feet forming the highest point in Price County Lake Michigan represents the lowest point at 580 feet The northern and eastern two-thirds of the state were heavily glaciated during the last stage of the Pleistocene and now contain many of the best examples of continental glaciation topOgraphy including considerable ground moraine large terminal recessional and intetlobate moraines kames eskers drumlins and small depressions or kettle-holes Most of the nearly 9000 lakes of 20 acres or more lie within this part of the state The southwestern third of the state is the Driftless Area It includes most of the large flat generally sandy bed of Glacial Lake Wisconsin plus the highly dissected ridge and valley section with numerous cliffs and rock-outcroppings (Figure 1)

Much of the bedrock in the state lies below glacial till but exposures include igneous and metamiddot morphic rocks primarily granitic especially in the north as well as sedimentary layers primarily sandshystones and dolomitic limestones especially along Lake Michigan and in the DriftlesS Area

The climate of Wisconsin is classified as humid-temperate with cold winters The annual precipimiddot tation averages approximately 30 inches which is sufficiently even in its seasonal distribution to support forest vegetation throughout most of the state Nevertheless according to Curtis (I 959) the pre-settlement vegetation of the state (J 830-1850) included about 27 prairie and oak-savanna plus about 64 forests

The state is presently divided into seventy-two counties Marathon County is the largest (1586 sq mi) the smallest counties include Pepin (235 sq mi) Ozaukee (236 sq mi) andMilw8ukee (237 sq mi) We included Menominee County though its future status as a county is in doubt since it regained its Indian reservation status and rights in 1973 This county was established April 30 1961 from the Menomishynee Indian reservation which consisted of parts of Shawano and Oconto counties Wisconsin ranks 26th in the United States in area with 54464 square miles and 16th in population with 4651000

More ecological research is needed on the bryophyte flora of Wisconsin especially to relate it to the vegetation and to such other environmental features as soils climate and glaCial deposits Plant communishyties in Wisconsin are many and diverse these are summarized on Figure 2 from studies conducted by Curtis (1959) The state is divided into northern and southern floristic provinces by a tension zone deshyscribed by Curtis (1959) This tension zone is an eeotonal region vatying from 10middot31 mt wide where the ranges of many vascular plants overlap or terminatej it runs in an S-curve northeast from Milwaukee to Hudson in St Croix County (Figure 2) The principal plant communities of the northern province apshyproaching their southern boundaries within the zone include the northern conifermiddothardwood forests of sugar maple white pine hemlock and yellow birch and the Sphagnum bogs with tamarack and black spruce Southern communities at or near their northern limits within this zone include prairies oak and jack pine savannas and hardwood forests of maples oaks ash ehn and cottonwood

We hope to encourage research on cryptogamic ecology with our checklist We know that some species are northern while others are southern (eg cimacium) fits the tension zone concept Freckmann (I977) while Bryoxiphium is only found in the Driftless Area of the state Steere (1937) and Love and Love (1953) The tension zone theory may prove to be acceptable for bryophytes We speculate that certain bryophytes can indicate particular plant communities since many bryophytes are restricted to preshycise environmental requirements Many papers or theses have dealt with cryptogamic bark and rock ecology in Wisconsin (Culberson 1955 Foote 1962 Hale 1952 1955 and Cole 1975)

We hope that this atlas will stimulate collecting in counties poorly sampled so that our knowledge of the bryophytes can be increased It should aid anyone interested in floras for preservation environmental impact statements and distributional studies

7

A numerical summary of the bryophyte flora of the state based on our literature search is presented in Table 1 (below) This includes 517 taxa 70 families and 176 genera This numerical summary is comshypared with the published floras of Michigan mosses (Crum 1976) and Minnesota hepatics (Schuster 1953) Thus we surmise that further collecting could lead to the same approximate numbers for Wisconsin bryoshyphytes

Table 1 Comparison of Recorded Bryophytes of Wisconsin with Michigan and Minnesota and Summary of Wisconsin Taxa

Wisconsin Michigan (Crum 1976)

Minnesota (Schuster 1955)

Hepaticsamp Hornworts

Taxa Families Genera

137 28 47

174 32 52

Mosses

Taxa Families Genera

380 42

129

503 42

149

8

-

~

~ tIt

Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map

ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area

9

Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959

tit beol Forest e Phillie

Northernmiddot hardwood forest reg Pine 5cMInna

copy Southernmiddot hordwood forest CD Oak Savanna

10

SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES

ANTHOCEROTAE (Hornworts)

1 Anthocerotaceae Anthoceros 1 emnd Prosk

macounii M A Howe punctatum 1

Phaeoceros Prosk laevis (1) Prosk subsp laevis

Notothylas Sul1 orbicularis (Schwein) Sullo

HEPATICAE (liverworts)

2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum

trlchophyllum (L) Dum subsp trlchophylluni

3 Ptilidiaceae Ptilidium Nees

ci1iare (L) Hampe pulcherrimum (G Web) Hampe

4 Trichocoleaceae Trichocolea Dum nom cons

tomentella (Ehrh) Dum

5 Lepidoziaceae Kunia von Martens

setacea (Web) GroUe

Lepidozia (Dum) Dum reptans (1) Dum

Bazzania S Gray trilobata (1) S Gray var triZobata

6 Calypogejaceae 011ypogeja Raddi

flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda

11

7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum

biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps

Noweilia Mitt cunifolilz (Dicks) Mitt

8 Adelanthaceae Odontoschimuz (Dwn) Dum

denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw

c

9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$

divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD

10 Geo~c~ larptuttllus Nees

scutatus (Web et Mohr) Spruce

Geocalyx Nees grGVeolens (Schrad) Nees

11 Lophocoleaceae Lophocolea (Dum) Dum

heteropllyll4 (Schrad) Own minorNees

Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees

12 PlagiochiJaceae Plagio(hila Dum nom cons

tlBpienioides (L) Dum var asplenioidu

13 Jungermanniaceae Btriilophoril Loeske

attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske

12

Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones

Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg

Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum

Mylill S Gray nom cons anomala (Hook) S Gray

Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis

Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook

14 Scapaniaceae Diplophyllum (Dum) Dum nom cons

apiculatum (Evans) Steph var apiculatum

Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta

13

cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata

15 Radulaceae Raduia Dum nom COllS

compianata (L) Dum obconica Sull

16 Porellaceae Parella L

pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb

17 Jubulaceae FruUanill Raddi

bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt

18 lejeuneaceae Calolejeunea (Spruce) Schlffn

biddlecomiae (Aust) Evans

Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch

19 Codoniaceae Fossombronia Raddi

foveolata Lindb

20 Pa11aviciniaceae Paklvieinlil S Gray nom cons

lyeiii (Hook) Carruth

14

Moerckia Gott hibemica (Hook) Gott

21 PeWaceae Pellfa Raddi nom cons propos

endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr

22 Metzgeriaceae Metzgerla Raddi

conjugata Undb

23 Blasiacae B L

pulilm L

24 Aneurareae AneumDum

pinguis (L) Dum

Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth

25 Aytoniaceae Reboulill Raddi nom conI

hemllphaerica (L) Radeli

ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe

26 Conocephalaceae Conocephalum Wigg nom cons

conicum (L) Lindb

27 Marchantiaceae Preiuia Corda

quadrata (Scop) Nees

Marchantia L polymorpha L

28 Ricciaceae Riccwcarpus Corda

natans (L) Corda

Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm

15

i

CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust

MUSCI (Mosses)

29 Sphagnaceae SpIutgnum L

capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm

WtlI7IItorjii Russ wuljfanum Gira

30 Andreaeaceae AndNrrerI Hedw

rupeitrlr Oedw var pest

31 Fissident~e Fluid_ lledw

trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u

16

ATLAS OF WISCONSIN BRYOPHYTES

November 21 1979

THE MUSEUM OF NATURAL HISTORY Stevens Point Wisconsin

Reports on the Fauna and Flora of Wisconsin No 16

Part I

Climacium dendroides (Tree moss)

ATLAS OF WISCONSIN BRYOPHYTES

By Frank D Bowers

Curator of Non-Vascular Plants-Herbarium Museum of Natural History and

Department of Biology and

Sally Kauss Freckmann Research Associate

Museum of Natural History Universit y of Wisconsin

Stevens Point Wisconsin 54481 U S A

2

CONTENTS

ABSTRACT 4

ACKNOWLEDCEMENTS 4

INTRODUCTION 6

GEOGRAPHIC CLIMATIC AND BOTANICAL fEATURES OF WISCONSIN 7

SYSTEMATIC ARLNGEMENT OF WISCONSIN BRYOPHYTES 11

MAPS OF BRYOPH) iE TAXA REPORTED FOR WISCONSIN 29

HORNWORTS AND LIVERWORTS 30

MUSCI 54

FOSSIL AND SUBFOSSIL MOSSES OF WISCONSIN 119

PALEONTOLOGICAL Rb J3RENCES 119

SYNONYMS AND EXCLUDED TAXA 120

PIONEER BRYOLOGISTS OF WISCONSIN 126

LITERATURE CITED 127

3

ABSTRACf

117This atlas of Wisconsin bryophytes shows the distribution of)30 taxa (137 hepatics 389 mosses)

on state maps with a dot in each county for which each taxon has been reported in the literature A list of synonyms and excluded taxa is included along with an extensive bibliography Pertinent features of the natural history and history of bryology in the state are discussed

ACKNOWLEDGEMENTS

We want to thank Drs Fred Hermann and John Thomson of the University of Wisconsin-Madison herbarium and Botany department for locating and lending packets and for references to aid us in accumushylating a more accurate complete report The illustration of Climacium dendroides on the cover was drawn by Robert Freckmann Of course we are grateful to all the people who have collected specimens of Wisshyconsin bryophytes and preserved them in herbariums making this distributional study possible

4

ONEIDA

LINCOLN

TAYLOR

t---------- MARATHON C1l-RK

SAu~

t

~

~ C i-W=AS_-r7-1

l=itHLANO

)

lOCK

INGTb

lt

WALWORTH

5

INTRODUCfION

This atlas of the bryophyte flora of Wisconsin is an attempt to show the distribution of all bryoshyphyte species known to occur in the state by county occurrenoes based on published reports in the literashyture It is not an annot~ted check list (only a feW problematic taxa were examined) but is a literature search intended as a foundation for a complete flora or for otherbryological research

The literature of btYo1ogy is large and spread over numerous journals reports monographs and manuals It would be almost impossible to search all sources for every reference attributing bryophyte species to Wisconsin and to particular localities in the state Wehav checked many inlportant references and paid special attention to works dealing with Wisconsiit botany and the bryologyof the upper Midwest Several bibliographies and indices were checked thoroughly including Green an~ Curtis (1955) the Bry ologist indices (Crum 1959 and Miller 1976) andt)le Wisconsin Academy of Sciences Arts and Letters indices

Only those published repots which app~red to)e substanti2ted by herbarium specimens and which gave- ~fficient location dat to P~CetheW(On in amiddot particula(county were-used in the initial compilation Species described in manuals in generalized way implying their occurrence in Wisconsin (Without menmiddot tioning Wisconsin specifically or citing a particular location in the state) were excluded eg Gnmmia raui (Wynne 1943) and Pohlia annotinll var decipiens (Crum 1976) Ecological papers which were either too general in taxonomic determinations (eg Sphagnum sp Polytrichum sp) or in location data were also excluded Some other taxa attributed to Wisconsin which have been excluded are

Cylindrothectum compr~ (Cheney 189tmiddotDane Co) identified later by Grout as Entodon seducshytrtx

Dicranell4 humilis (Holzjne~ 1925middotBuffalo Co) probablyD vllJia since Dhumilis is not found in North Arnerl~

Didymodon triarium (Foote 1 966-SW Visconsin) identified by Fred Hennann as Barbulaacuta Funoria microstoma (anonymous 1975middotDane Co) excluded because it is outside of its geographic

range and the packe~ QaMot be located Phil(motis calcarea (Cheney and Evans 1944-Dane Co) excluded in Crum Steere and Anderson

(1973) Plagiobryum demissum and P zierl (Cheney and Evans 1944-various counties) are both Arcticshy

alpine species not likely to be found in isconsin Two of Cheneys collections of Pl1giobyrum have been referred to Bryum

Pogonatum alpinum var septentrionale (Che~ey and Evans 1944-Chippewa Co) was misidentified (Ireland 1960) and is P umigerum

Rhizomnium fAPpoJachIanum and R punctalu1fl ssp chlorophyDosum (Koponen 1973middotseveral counshy ties) excluded North American bryologiflts have not recognized these taxa in America

Sphagnum tenellum (Cheney and Evans 1944-Waukesha Co)an Atlantic species not occurring in Wisconsin (Cruro 1975)

The nomenclature and systematic arrangement of hornwort 00 hepatic species in the atlas follow Stotler and Crandall-5totler (1977) and of moss species fonow Crum Steere and Anderson (1973) except in such instances as the JreptlIfockIdus fonns whiGh follow Wijk et aL (19S9middot1973) Species reported in the literature under names which are now listed as synonyms on either of these lists were converted to the accepted name A dot is placed in each county of Wisconsin from which the taxon has been reported The numbers after the name of the taxon correspond to the numbered references in the bibliography which giye ~act location data OEat least location data -s~fficient to place it in-particular counties in the state The numbers in parentheses indicate references which attribute the taxon to Wisconsin without providing exact loltation data

Fossil and subfossil reports are summarized Slfparately in our atlas only six species (Aulacomnium turgidum lJ1achythecium turgidum Ditrichum Jlelficaule Hypnum bambergti Scorpidium 6fJisities and Tortel14 1rIgi1i$) are known exclUsively from the fossil record ~

6

GEOGRAPHIC CLIMATIC AND BOTANICAL FEATURES OF WISCONSIN

Wisconsin is a north central state bounded on the east by Lake Michigan on the north by Michigan and Lake Superior on the south by Illinois and on the west by the Mississippi and St Croix rivers It is located midway between the Prime Meridian (0deg) of Longitude and 1800 Longitude and midway between the equator and the north pole thus placing it in the center of the northern half of the western hemisphere

The topography of the state is relatively flat a few hills exceed 1900 feet with Thns Hill at 1952 feet forming the highest point in Price County Lake Michigan represents the lowest point at 580 feet The northern and eastern two-thirds of the state were heavily glaciated during the last stage of the Pleistocene and now contain many of the best examples of continental glaciation topOgraphy including considerable ground moraine large terminal recessional and intetlobate moraines kames eskers drumlins and small depressions or kettle-holes Most of the nearly 9000 lakes of 20 acres or more lie within this part of the state The southwestern third of the state is the Driftless Area It includes most of the large flat generally sandy bed of Glacial Lake Wisconsin plus the highly dissected ridge and valley section with numerous cliffs and rock-outcroppings (Figure 1)

Much of the bedrock in the state lies below glacial till but exposures include igneous and metamiddot morphic rocks primarily granitic especially in the north as well as sedimentary layers primarily sandshystones and dolomitic limestones especially along Lake Michigan and in the DriftlesS Area

The climate of Wisconsin is classified as humid-temperate with cold winters The annual precipimiddot tation averages approximately 30 inches which is sufficiently even in its seasonal distribution to support forest vegetation throughout most of the state Nevertheless according to Curtis (I 959) the pre-settlement vegetation of the state (J 830-1850) included about 27 prairie and oak-savanna plus about 64 forests

The state is presently divided into seventy-two counties Marathon County is the largest (1586 sq mi) the smallest counties include Pepin (235 sq mi) Ozaukee (236 sq mi) andMilw8ukee (237 sq mi) We included Menominee County though its future status as a county is in doubt since it regained its Indian reservation status and rights in 1973 This county was established April 30 1961 from the Menomishynee Indian reservation which consisted of parts of Shawano and Oconto counties Wisconsin ranks 26th in the United States in area with 54464 square miles and 16th in population with 4651000

More ecological research is needed on the bryophyte flora of Wisconsin especially to relate it to the vegetation and to such other environmental features as soils climate and glaCial deposits Plant communishyties in Wisconsin are many and diverse these are summarized on Figure 2 from studies conducted by Curtis (1959) The state is divided into northern and southern floristic provinces by a tension zone deshyscribed by Curtis (1959) This tension zone is an eeotonal region vatying from 10middot31 mt wide where the ranges of many vascular plants overlap or terminatej it runs in an S-curve northeast from Milwaukee to Hudson in St Croix County (Figure 2) The principal plant communities of the northern province apshyproaching their southern boundaries within the zone include the northern conifermiddothardwood forests of sugar maple white pine hemlock and yellow birch and the Sphagnum bogs with tamarack and black spruce Southern communities at or near their northern limits within this zone include prairies oak and jack pine savannas and hardwood forests of maples oaks ash ehn and cottonwood

We hope to encourage research on cryptogamic ecology with our checklist We know that some species are northern while others are southern (eg cimacium) fits the tension zone concept Freckmann (I977) while Bryoxiphium is only found in the Driftless Area of the state Steere (1937) and Love and Love (1953) The tension zone theory may prove to be acceptable for bryophytes We speculate that certain bryophytes can indicate particular plant communities since many bryophytes are restricted to preshycise environmental requirements Many papers or theses have dealt with cryptogamic bark and rock ecology in Wisconsin (Culberson 1955 Foote 1962 Hale 1952 1955 and Cole 1975)

We hope that this atlas will stimulate collecting in counties poorly sampled so that our knowledge of the bryophytes can be increased It should aid anyone interested in floras for preservation environmental impact statements and distributional studies

7

A numerical summary of the bryophyte flora of the state based on our literature search is presented in Table 1 (below) This includes 517 taxa 70 families and 176 genera This numerical summary is comshypared with the published floras of Michigan mosses (Crum 1976) and Minnesota hepatics (Schuster 1953) Thus we surmise that further collecting could lead to the same approximate numbers for Wisconsin bryoshyphytes

Table 1 Comparison of Recorded Bryophytes of Wisconsin with Michigan and Minnesota and Summary of Wisconsin Taxa

Wisconsin Michigan (Crum 1976)

Minnesota (Schuster 1955)

Hepaticsamp Hornworts

Taxa Families Genera

137 28 47

174 32 52

Mosses

Taxa Families Genera

380 42

129

503 42

149

8

-

~

~ tIt

Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map

ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area

9

Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959

tit beol Forest e Phillie

Northernmiddot hardwood forest reg Pine 5cMInna

copy Southernmiddot hordwood forest CD Oak Savanna

10

SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES

ANTHOCEROTAE (Hornworts)

1 Anthocerotaceae Anthoceros 1 emnd Prosk

macounii M A Howe punctatum 1

Phaeoceros Prosk laevis (1) Prosk subsp laevis

Notothylas Sul1 orbicularis (Schwein) Sullo

HEPATICAE (liverworts)

2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum

trlchophyllum (L) Dum subsp trlchophylluni

3 Ptilidiaceae Ptilidium Nees

ci1iare (L) Hampe pulcherrimum (G Web) Hampe

4 Trichocoleaceae Trichocolea Dum nom cons

tomentella (Ehrh) Dum

5 Lepidoziaceae Kunia von Martens

setacea (Web) GroUe

Lepidozia (Dum) Dum reptans (1) Dum

Bazzania S Gray trilobata (1) S Gray var triZobata

6 Calypogejaceae 011ypogeja Raddi

flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda

11

7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum

biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps

Noweilia Mitt cunifolilz (Dicks) Mitt

8 Adelanthaceae Odontoschimuz (Dwn) Dum

denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw

c

9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$

divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD

10 Geo~c~ larptuttllus Nees

scutatus (Web et Mohr) Spruce

Geocalyx Nees grGVeolens (Schrad) Nees

11 Lophocoleaceae Lophocolea (Dum) Dum

heteropllyll4 (Schrad) Own minorNees

Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees

12 PlagiochiJaceae Plagio(hila Dum nom cons

tlBpienioides (L) Dum var asplenioidu

13 Jungermanniaceae Btriilophoril Loeske

attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske

12

Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones

Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg

Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum

Mylill S Gray nom cons anomala (Hook) S Gray

Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis

Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook

14 Scapaniaceae Diplophyllum (Dum) Dum nom cons

apiculatum (Evans) Steph var apiculatum

Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta

13

cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata

15 Radulaceae Raduia Dum nom COllS

compianata (L) Dum obconica Sull

16 Porellaceae Parella L

pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb

17 Jubulaceae FruUanill Raddi

bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt

18 lejeuneaceae Calolejeunea (Spruce) Schlffn

biddlecomiae (Aust) Evans

Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch

19 Codoniaceae Fossombronia Raddi

foveolata Lindb

20 Pa11aviciniaceae Paklvieinlil S Gray nom cons

lyeiii (Hook) Carruth

14

Moerckia Gott hibemica (Hook) Gott

21 PeWaceae Pellfa Raddi nom cons propos

endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr

22 Metzgeriaceae Metzgerla Raddi

conjugata Undb

23 Blasiacae B L

pulilm L

24 Aneurareae AneumDum

pinguis (L) Dum

Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth

25 Aytoniaceae Reboulill Raddi nom conI

hemllphaerica (L) Radeli

ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe

26 Conocephalaceae Conocephalum Wigg nom cons

conicum (L) Lindb

27 Marchantiaceae Preiuia Corda

quadrata (Scop) Nees

Marchantia L polymorpha L

28 Ricciaceae Riccwcarpus Corda

natans (L) Corda

Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm

15

i

CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust

MUSCI (Mosses)

29 Sphagnaceae SpIutgnum L

capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm

WtlI7IItorjii Russ wuljfanum Gira

30 Andreaeaceae AndNrrerI Hedw

rupeitrlr Oedw var pest

31 Fissident~e Fluid_ lledw

trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u

16

Climacium dendroides (Tree moss)

ATLAS OF WISCONSIN BRYOPHYTES

By Frank D Bowers

Curator of Non-Vascular Plants-Herbarium Museum of Natural History and

Department of Biology and

Sally Kauss Freckmann Research Associate

Museum of Natural History Universit y of Wisconsin

Stevens Point Wisconsin 54481 U S A

2

CONTENTS

ABSTRACT 4

ACKNOWLEDCEMENTS 4

INTRODUCTION 6

GEOGRAPHIC CLIMATIC AND BOTANICAL fEATURES OF WISCONSIN 7

SYSTEMATIC ARLNGEMENT OF WISCONSIN BRYOPHYTES 11

MAPS OF BRYOPH) iE TAXA REPORTED FOR WISCONSIN 29

HORNWORTS AND LIVERWORTS 30

MUSCI 54

FOSSIL AND SUBFOSSIL MOSSES OF WISCONSIN 119

PALEONTOLOGICAL Rb J3RENCES 119

SYNONYMS AND EXCLUDED TAXA 120

PIONEER BRYOLOGISTS OF WISCONSIN 126

LITERATURE CITED 127

3

ABSTRACf

117This atlas of Wisconsin bryophytes shows the distribution of)30 taxa (137 hepatics 389 mosses)

on state maps with a dot in each county for which each taxon has been reported in the literature A list of synonyms and excluded taxa is included along with an extensive bibliography Pertinent features of the natural history and history of bryology in the state are discussed

ACKNOWLEDGEMENTS

We want to thank Drs Fred Hermann and John Thomson of the University of Wisconsin-Madison herbarium and Botany department for locating and lending packets and for references to aid us in accumushylating a more accurate complete report The illustration of Climacium dendroides on the cover was drawn by Robert Freckmann Of course we are grateful to all the people who have collected specimens of Wisshyconsin bryophytes and preserved them in herbariums making this distributional study possible

4

ONEIDA

LINCOLN

TAYLOR

t---------- MARATHON C1l-RK

SAu~

t

~

~ C i-W=AS_-r7-1

l=itHLANO

)

lOCK

INGTb

lt

WALWORTH

5

INTRODUCfION

This atlas of the bryophyte flora of Wisconsin is an attempt to show the distribution of all bryoshyphyte species known to occur in the state by county occurrenoes based on published reports in the literashyture It is not an annot~ted check list (only a feW problematic taxa were examined) but is a literature search intended as a foundation for a complete flora or for otherbryological research

The literature of btYo1ogy is large and spread over numerous journals reports monographs and manuals It would be almost impossible to search all sources for every reference attributing bryophyte species to Wisconsin and to particular localities in the state Wehav checked many inlportant references and paid special attention to works dealing with Wisconsiit botany and the bryologyof the upper Midwest Several bibliographies and indices were checked thoroughly including Green an~ Curtis (1955) the Bry ologist indices (Crum 1959 and Miller 1976) andt)le Wisconsin Academy of Sciences Arts and Letters indices

Only those published repots which app~red to)e substanti2ted by herbarium specimens and which gave- ~fficient location dat to P~CetheW(On in amiddot particula(county were-used in the initial compilation Species described in manuals in generalized way implying their occurrence in Wisconsin (Without menmiddot tioning Wisconsin specifically or citing a particular location in the state) were excluded eg Gnmmia raui (Wynne 1943) and Pohlia annotinll var decipiens (Crum 1976) Ecological papers which were either too general in taxonomic determinations (eg Sphagnum sp Polytrichum sp) or in location data were also excluded Some other taxa attributed to Wisconsin which have been excluded are

Cylindrothectum compr~ (Cheney 189tmiddotDane Co) identified later by Grout as Entodon seducshytrtx

Dicranell4 humilis (Holzjne~ 1925middotBuffalo Co) probablyD vllJia since Dhumilis is not found in North Arnerl~

Didymodon triarium (Foote 1 966-SW Visconsin) identified by Fred Hennann as Barbulaacuta Funoria microstoma (anonymous 1975middotDane Co) excluded because it is outside of its geographic

range and the packe~ QaMot be located Phil(motis calcarea (Cheney and Evans 1944-Dane Co) excluded in Crum Steere and Anderson

(1973) Plagiobryum demissum and P zierl (Cheney and Evans 1944-various counties) are both Arcticshy

alpine species not likely to be found in isconsin Two of Cheneys collections of Pl1giobyrum have been referred to Bryum

Pogonatum alpinum var septentrionale (Che~ey and Evans 1944-Chippewa Co) was misidentified (Ireland 1960) and is P umigerum

Rhizomnium fAPpoJachIanum and R punctalu1fl ssp chlorophyDosum (Koponen 1973middotseveral counshy ties) excluded North American bryologiflts have not recognized these taxa in America

Sphagnum tenellum (Cheney and Evans 1944-Waukesha Co)an Atlantic species not occurring in Wisconsin (Cruro 1975)

The nomenclature and systematic arrangement of hornwort 00 hepatic species in the atlas follow Stotler and Crandall-5totler (1977) and of moss species fonow Crum Steere and Anderson (1973) except in such instances as the JreptlIfockIdus fonns whiGh follow Wijk et aL (19S9middot1973) Species reported in the literature under names which are now listed as synonyms on either of these lists were converted to the accepted name A dot is placed in each county of Wisconsin from which the taxon has been reported The numbers after the name of the taxon correspond to the numbered references in the bibliography which giye ~act location data OEat least location data -s~fficient to place it in-particular counties in the state The numbers in parentheses indicate references which attribute the taxon to Wisconsin without providing exact loltation data

Fossil and subfossil reports are summarized Slfparately in our atlas only six species (Aulacomnium turgidum lJ1achythecium turgidum Ditrichum Jlelficaule Hypnum bambergti Scorpidium 6fJisities and Tortel14 1rIgi1i$) are known exclUsively from the fossil record ~

6

GEOGRAPHIC CLIMATIC AND BOTANICAL FEATURES OF WISCONSIN

Wisconsin is a north central state bounded on the east by Lake Michigan on the north by Michigan and Lake Superior on the south by Illinois and on the west by the Mississippi and St Croix rivers It is located midway between the Prime Meridian (0deg) of Longitude and 1800 Longitude and midway between the equator and the north pole thus placing it in the center of the northern half of the western hemisphere

The topography of the state is relatively flat a few hills exceed 1900 feet with Thns Hill at 1952 feet forming the highest point in Price County Lake Michigan represents the lowest point at 580 feet The northern and eastern two-thirds of the state were heavily glaciated during the last stage of the Pleistocene and now contain many of the best examples of continental glaciation topOgraphy including considerable ground moraine large terminal recessional and intetlobate moraines kames eskers drumlins and small depressions or kettle-holes Most of the nearly 9000 lakes of 20 acres or more lie within this part of the state The southwestern third of the state is the Driftless Area It includes most of the large flat generally sandy bed of Glacial Lake Wisconsin plus the highly dissected ridge and valley section with numerous cliffs and rock-outcroppings (Figure 1)

Much of the bedrock in the state lies below glacial till but exposures include igneous and metamiddot morphic rocks primarily granitic especially in the north as well as sedimentary layers primarily sandshystones and dolomitic limestones especially along Lake Michigan and in the DriftlesS Area

The climate of Wisconsin is classified as humid-temperate with cold winters The annual precipimiddot tation averages approximately 30 inches which is sufficiently even in its seasonal distribution to support forest vegetation throughout most of the state Nevertheless according to Curtis (I 959) the pre-settlement vegetation of the state (J 830-1850) included about 27 prairie and oak-savanna plus about 64 forests

The state is presently divided into seventy-two counties Marathon County is the largest (1586 sq mi) the smallest counties include Pepin (235 sq mi) Ozaukee (236 sq mi) andMilw8ukee (237 sq mi) We included Menominee County though its future status as a county is in doubt since it regained its Indian reservation status and rights in 1973 This county was established April 30 1961 from the Menomishynee Indian reservation which consisted of parts of Shawano and Oconto counties Wisconsin ranks 26th in the United States in area with 54464 square miles and 16th in population with 4651000

More ecological research is needed on the bryophyte flora of Wisconsin especially to relate it to the vegetation and to such other environmental features as soils climate and glaCial deposits Plant communishyties in Wisconsin are many and diverse these are summarized on Figure 2 from studies conducted by Curtis (1959) The state is divided into northern and southern floristic provinces by a tension zone deshyscribed by Curtis (1959) This tension zone is an eeotonal region vatying from 10middot31 mt wide where the ranges of many vascular plants overlap or terminatej it runs in an S-curve northeast from Milwaukee to Hudson in St Croix County (Figure 2) The principal plant communities of the northern province apshyproaching their southern boundaries within the zone include the northern conifermiddothardwood forests of sugar maple white pine hemlock and yellow birch and the Sphagnum bogs with tamarack and black spruce Southern communities at or near their northern limits within this zone include prairies oak and jack pine savannas and hardwood forests of maples oaks ash ehn and cottonwood

We hope to encourage research on cryptogamic ecology with our checklist We know that some species are northern while others are southern (eg cimacium) fits the tension zone concept Freckmann (I977) while Bryoxiphium is only found in the Driftless Area of the state Steere (1937) and Love and Love (1953) The tension zone theory may prove to be acceptable for bryophytes We speculate that certain bryophytes can indicate particular plant communities since many bryophytes are restricted to preshycise environmental requirements Many papers or theses have dealt with cryptogamic bark and rock ecology in Wisconsin (Culberson 1955 Foote 1962 Hale 1952 1955 and Cole 1975)

We hope that this atlas will stimulate collecting in counties poorly sampled so that our knowledge of the bryophytes can be increased It should aid anyone interested in floras for preservation environmental impact statements and distributional studies

7

A numerical summary of the bryophyte flora of the state based on our literature search is presented in Table 1 (below) This includes 517 taxa 70 families and 176 genera This numerical summary is comshypared with the published floras of Michigan mosses (Crum 1976) and Minnesota hepatics (Schuster 1953) Thus we surmise that further collecting could lead to the same approximate numbers for Wisconsin bryoshyphytes

Table 1 Comparison of Recorded Bryophytes of Wisconsin with Michigan and Minnesota and Summary of Wisconsin Taxa

Wisconsin Michigan (Crum 1976)

Minnesota (Schuster 1955)

Hepaticsamp Hornworts

Taxa Families Genera

137 28 47

174 32 52

Mosses

Taxa Families Genera

380 42

129

503 42

149

8

-

~

~ tIt

Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map

ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area

9

Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959

tit beol Forest e Phillie

Northernmiddot hardwood forest reg Pine 5cMInna

copy Southernmiddot hordwood forest CD Oak Savanna

10

SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES

ANTHOCEROTAE (Hornworts)

1 Anthocerotaceae Anthoceros 1 emnd Prosk

macounii M A Howe punctatum 1

Phaeoceros Prosk laevis (1) Prosk subsp laevis

Notothylas Sul1 orbicularis (Schwein) Sullo

HEPATICAE (liverworts)

2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum

trlchophyllum (L) Dum subsp trlchophylluni

3 Ptilidiaceae Ptilidium Nees

ci1iare (L) Hampe pulcherrimum (G Web) Hampe

4 Trichocoleaceae Trichocolea Dum nom cons

tomentella (Ehrh) Dum

5 Lepidoziaceae Kunia von Martens

setacea (Web) GroUe

Lepidozia (Dum) Dum reptans (1) Dum

Bazzania S Gray trilobata (1) S Gray var triZobata

6 Calypogejaceae 011ypogeja Raddi

flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda

11

7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum

biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps

Noweilia Mitt cunifolilz (Dicks) Mitt

8 Adelanthaceae Odontoschimuz (Dwn) Dum

denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw

c

9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$

divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD

10 Geo~c~ larptuttllus Nees

scutatus (Web et Mohr) Spruce

Geocalyx Nees grGVeolens (Schrad) Nees

11 Lophocoleaceae Lophocolea (Dum) Dum

heteropllyll4 (Schrad) Own minorNees

Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees

12 PlagiochiJaceae Plagio(hila Dum nom cons

tlBpienioides (L) Dum var asplenioidu

13 Jungermanniaceae Btriilophoril Loeske

attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske

12

Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones

Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg

Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum

Mylill S Gray nom cons anomala (Hook) S Gray

Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis

Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook

14 Scapaniaceae Diplophyllum (Dum) Dum nom cons

apiculatum (Evans) Steph var apiculatum

Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta

13

cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata

15 Radulaceae Raduia Dum nom COllS

compianata (L) Dum obconica Sull

16 Porellaceae Parella L

pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb

17 Jubulaceae FruUanill Raddi

bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt

18 lejeuneaceae Calolejeunea (Spruce) Schlffn

biddlecomiae (Aust) Evans

Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch

19 Codoniaceae Fossombronia Raddi

foveolata Lindb

20 Pa11aviciniaceae Paklvieinlil S Gray nom cons

lyeiii (Hook) Carruth

14

Moerckia Gott hibemica (Hook) Gott

21 PeWaceae Pellfa Raddi nom cons propos

endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr

22 Metzgeriaceae Metzgerla Raddi

conjugata Undb

23 Blasiacae B L

pulilm L

24 Aneurareae AneumDum

pinguis (L) Dum

Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth

25 Aytoniaceae Reboulill Raddi nom conI

hemllphaerica (L) Radeli

ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe

26 Conocephalaceae Conocephalum Wigg nom cons

conicum (L) Lindb

27 Marchantiaceae Preiuia Corda

quadrata (Scop) Nees

Marchantia L polymorpha L

28 Ricciaceae Riccwcarpus Corda

natans (L) Corda

Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm

15

i

CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust

MUSCI (Mosses)

29 Sphagnaceae SpIutgnum L

capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm

WtlI7IItorjii Russ wuljfanum Gira

30 Andreaeaceae AndNrrerI Hedw

rupeitrlr Oedw var pest

31 Fissident~e Fluid_ lledw

trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u

16

CONTENTS

ABSTRACT 4

ACKNOWLEDCEMENTS 4

INTRODUCTION 6

GEOGRAPHIC CLIMATIC AND BOTANICAL fEATURES OF WISCONSIN 7

SYSTEMATIC ARLNGEMENT OF WISCONSIN BRYOPHYTES 11

MAPS OF BRYOPH) iE TAXA REPORTED FOR WISCONSIN 29

HORNWORTS AND LIVERWORTS 30

MUSCI 54

FOSSIL AND SUBFOSSIL MOSSES OF WISCONSIN 119

PALEONTOLOGICAL Rb J3RENCES 119

SYNONYMS AND EXCLUDED TAXA 120

PIONEER BRYOLOGISTS OF WISCONSIN 126

LITERATURE CITED 127

3

ABSTRACf

117This atlas of Wisconsin bryophytes shows the distribution of)30 taxa (137 hepatics 389 mosses)

on state maps with a dot in each county for which each taxon has been reported in the literature A list of synonyms and excluded taxa is included along with an extensive bibliography Pertinent features of the natural history and history of bryology in the state are discussed

ACKNOWLEDGEMENTS

We want to thank Drs Fred Hermann and John Thomson of the University of Wisconsin-Madison herbarium and Botany department for locating and lending packets and for references to aid us in accumushylating a more accurate complete report The illustration of Climacium dendroides on the cover was drawn by Robert Freckmann Of course we are grateful to all the people who have collected specimens of Wisshyconsin bryophytes and preserved them in herbariums making this distributional study possible

4

ONEIDA

LINCOLN

TAYLOR

t---------- MARATHON C1l-RK

SAu~

t

~

~ C i-W=AS_-r7-1

l=itHLANO

)

lOCK

INGTb

lt

WALWORTH

5

INTRODUCfION

This atlas of the bryophyte flora of Wisconsin is an attempt to show the distribution of all bryoshyphyte species known to occur in the state by county occurrenoes based on published reports in the literashyture It is not an annot~ted check list (only a feW problematic taxa were examined) but is a literature search intended as a foundation for a complete flora or for otherbryological research

The literature of btYo1ogy is large and spread over numerous journals reports monographs and manuals It would be almost impossible to search all sources for every reference attributing bryophyte species to Wisconsin and to particular localities in the state Wehav checked many inlportant references and paid special attention to works dealing with Wisconsiit botany and the bryologyof the upper Midwest Several bibliographies and indices were checked thoroughly including Green an~ Curtis (1955) the Bry ologist indices (Crum 1959 and Miller 1976) andt)le Wisconsin Academy of Sciences Arts and Letters indices

Only those published repots which app~red to)e substanti2ted by herbarium specimens and which gave- ~fficient location dat to P~CetheW(On in amiddot particula(county were-used in the initial compilation Species described in manuals in generalized way implying their occurrence in Wisconsin (Without menmiddot tioning Wisconsin specifically or citing a particular location in the state) were excluded eg Gnmmia raui (Wynne 1943) and Pohlia annotinll var decipiens (Crum 1976) Ecological papers which were either too general in taxonomic determinations (eg Sphagnum sp Polytrichum sp) or in location data were also excluded Some other taxa attributed to Wisconsin which have been excluded are

Cylindrothectum compr~ (Cheney 189tmiddotDane Co) identified later by Grout as Entodon seducshytrtx

Dicranell4 humilis (Holzjne~ 1925middotBuffalo Co) probablyD vllJia since Dhumilis is not found in North Arnerl~

Didymodon triarium (Foote 1 966-SW Visconsin) identified by Fred Hennann as Barbulaacuta Funoria microstoma (anonymous 1975middotDane Co) excluded because it is outside of its geographic

range and the packe~ QaMot be located Phil(motis calcarea (Cheney and Evans 1944-Dane Co) excluded in Crum Steere and Anderson

(1973) Plagiobryum demissum and P zierl (Cheney and Evans 1944-various counties) are both Arcticshy

alpine species not likely to be found in isconsin Two of Cheneys collections of Pl1giobyrum have been referred to Bryum

Pogonatum alpinum var septentrionale (Che~ey and Evans 1944-Chippewa Co) was misidentified (Ireland 1960) and is P umigerum

Rhizomnium fAPpoJachIanum and R punctalu1fl ssp chlorophyDosum (Koponen 1973middotseveral counshy ties) excluded North American bryologiflts have not recognized these taxa in America

Sphagnum tenellum (Cheney and Evans 1944-Waukesha Co)an Atlantic species not occurring in Wisconsin (Cruro 1975)

The nomenclature and systematic arrangement of hornwort 00 hepatic species in the atlas follow Stotler and Crandall-5totler (1977) and of moss species fonow Crum Steere and Anderson (1973) except in such instances as the JreptlIfockIdus fonns whiGh follow Wijk et aL (19S9middot1973) Species reported in the literature under names which are now listed as synonyms on either of these lists were converted to the accepted name A dot is placed in each county of Wisconsin from which the taxon has been reported The numbers after the name of the taxon correspond to the numbered references in the bibliography which giye ~act location data OEat least location data -s~fficient to place it in-particular counties in the state The numbers in parentheses indicate references which attribute the taxon to Wisconsin without providing exact loltation data

Fossil and subfossil reports are summarized Slfparately in our atlas only six species (Aulacomnium turgidum lJ1achythecium turgidum Ditrichum Jlelficaule Hypnum bambergti Scorpidium 6fJisities and Tortel14 1rIgi1i$) are known exclUsively from the fossil record ~

6

GEOGRAPHIC CLIMATIC AND BOTANICAL FEATURES OF WISCONSIN

Wisconsin is a north central state bounded on the east by Lake Michigan on the north by Michigan and Lake Superior on the south by Illinois and on the west by the Mississippi and St Croix rivers It is located midway between the Prime Meridian (0deg) of Longitude and 1800 Longitude and midway between the equator and the north pole thus placing it in the center of the northern half of the western hemisphere

The topography of the state is relatively flat a few hills exceed 1900 feet with Thns Hill at 1952 feet forming the highest point in Price County Lake Michigan represents the lowest point at 580 feet The northern and eastern two-thirds of the state were heavily glaciated during the last stage of the Pleistocene and now contain many of the best examples of continental glaciation topOgraphy including considerable ground moraine large terminal recessional and intetlobate moraines kames eskers drumlins and small depressions or kettle-holes Most of the nearly 9000 lakes of 20 acres or more lie within this part of the state The southwestern third of the state is the Driftless Area It includes most of the large flat generally sandy bed of Glacial Lake Wisconsin plus the highly dissected ridge and valley section with numerous cliffs and rock-outcroppings (Figure 1)

Much of the bedrock in the state lies below glacial till but exposures include igneous and metamiddot morphic rocks primarily granitic especially in the north as well as sedimentary layers primarily sandshystones and dolomitic limestones especially along Lake Michigan and in the DriftlesS Area

The climate of Wisconsin is classified as humid-temperate with cold winters The annual precipimiddot tation averages approximately 30 inches which is sufficiently even in its seasonal distribution to support forest vegetation throughout most of the state Nevertheless according to Curtis (I 959) the pre-settlement vegetation of the state (J 830-1850) included about 27 prairie and oak-savanna plus about 64 forests

The state is presently divided into seventy-two counties Marathon County is the largest (1586 sq mi) the smallest counties include Pepin (235 sq mi) Ozaukee (236 sq mi) andMilw8ukee (237 sq mi) We included Menominee County though its future status as a county is in doubt since it regained its Indian reservation status and rights in 1973 This county was established April 30 1961 from the Menomishynee Indian reservation which consisted of parts of Shawano and Oconto counties Wisconsin ranks 26th in the United States in area with 54464 square miles and 16th in population with 4651000

More ecological research is needed on the bryophyte flora of Wisconsin especially to relate it to the vegetation and to such other environmental features as soils climate and glaCial deposits Plant communishyties in Wisconsin are many and diverse these are summarized on Figure 2 from studies conducted by Curtis (1959) The state is divided into northern and southern floristic provinces by a tension zone deshyscribed by Curtis (1959) This tension zone is an eeotonal region vatying from 10middot31 mt wide where the ranges of many vascular plants overlap or terminatej it runs in an S-curve northeast from Milwaukee to Hudson in St Croix County (Figure 2) The principal plant communities of the northern province apshyproaching their southern boundaries within the zone include the northern conifermiddothardwood forests of sugar maple white pine hemlock and yellow birch and the Sphagnum bogs with tamarack and black spruce Southern communities at or near their northern limits within this zone include prairies oak and jack pine savannas and hardwood forests of maples oaks ash ehn and cottonwood

We hope to encourage research on cryptogamic ecology with our checklist We know that some species are northern while others are southern (eg cimacium) fits the tension zone concept Freckmann (I977) while Bryoxiphium is only found in the Driftless Area of the state Steere (1937) and Love and Love (1953) The tension zone theory may prove to be acceptable for bryophytes We speculate that certain bryophytes can indicate particular plant communities since many bryophytes are restricted to preshycise environmental requirements Many papers or theses have dealt with cryptogamic bark and rock ecology in Wisconsin (Culberson 1955 Foote 1962 Hale 1952 1955 and Cole 1975)

We hope that this atlas will stimulate collecting in counties poorly sampled so that our knowledge of the bryophytes can be increased It should aid anyone interested in floras for preservation environmental impact statements and distributional studies

7

A numerical summary of the bryophyte flora of the state based on our literature search is presented in Table 1 (below) This includes 517 taxa 70 families and 176 genera This numerical summary is comshypared with the published floras of Michigan mosses (Crum 1976) and Minnesota hepatics (Schuster 1953) Thus we surmise that further collecting could lead to the same approximate numbers for Wisconsin bryoshyphytes

Table 1 Comparison of Recorded Bryophytes of Wisconsin with Michigan and Minnesota and Summary of Wisconsin Taxa

Wisconsin Michigan (Crum 1976)

Minnesota (Schuster 1955)

Hepaticsamp Hornworts

Taxa Families Genera

137 28 47

174 32 52

Mosses

Taxa Families Genera

380 42

129

503 42

149

8

-

~

~ tIt

Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map

ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area

9

Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959

tit beol Forest e Phillie

Northernmiddot hardwood forest reg Pine 5cMInna

copy Southernmiddot hordwood forest CD Oak Savanna

10

SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES

ANTHOCEROTAE (Hornworts)

1 Anthocerotaceae Anthoceros 1 emnd Prosk

macounii M A Howe punctatum 1

Phaeoceros Prosk laevis (1) Prosk subsp laevis

Notothylas Sul1 orbicularis (Schwein) Sullo

HEPATICAE (liverworts)

2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum

trlchophyllum (L) Dum subsp trlchophylluni

3 Ptilidiaceae Ptilidium Nees

ci1iare (L) Hampe pulcherrimum (G Web) Hampe

4 Trichocoleaceae Trichocolea Dum nom cons

tomentella (Ehrh) Dum

5 Lepidoziaceae Kunia von Martens

setacea (Web) GroUe

Lepidozia (Dum) Dum reptans (1) Dum

Bazzania S Gray trilobata (1) S Gray var triZobata

6 Calypogejaceae 011ypogeja Raddi

flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda

11

7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum

biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps

Noweilia Mitt cunifolilz (Dicks) Mitt

8 Adelanthaceae Odontoschimuz (Dwn) Dum

denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw

c

9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$

divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD

10 Geo~c~ larptuttllus Nees

scutatus (Web et Mohr) Spruce

Geocalyx Nees grGVeolens (Schrad) Nees

11 Lophocoleaceae Lophocolea (Dum) Dum

heteropllyll4 (Schrad) Own minorNees

Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees

12 PlagiochiJaceae Plagio(hila Dum nom cons

tlBpienioides (L) Dum var asplenioidu

13 Jungermanniaceae Btriilophoril Loeske

attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske

12

Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones

Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg

Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum

Mylill S Gray nom cons anomala (Hook) S Gray

Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis

Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook

14 Scapaniaceae Diplophyllum (Dum) Dum nom cons

apiculatum (Evans) Steph var apiculatum

Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta

13

cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata

15 Radulaceae Raduia Dum nom COllS

compianata (L) Dum obconica Sull

16 Porellaceae Parella L

pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb

17 Jubulaceae FruUanill Raddi

bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt

18 lejeuneaceae Calolejeunea (Spruce) Schlffn

biddlecomiae (Aust) Evans

Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch

19 Codoniaceae Fossombronia Raddi

foveolata Lindb

20 Pa11aviciniaceae Paklvieinlil S Gray nom cons

lyeiii (Hook) Carruth

14

Moerckia Gott hibemica (Hook) Gott

21 PeWaceae Pellfa Raddi nom cons propos

endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr

22 Metzgeriaceae Metzgerla Raddi

conjugata Undb

23 Blasiacae B L

pulilm L

24 Aneurareae AneumDum

pinguis (L) Dum

Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth

25 Aytoniaceae Reboulill Raddi nom conI

hemllphaerica (L) Radeli

ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe

26 Conocephalaceae Conocephalum Wigg nom cons

conicum (L) Lindb

27 Marchantiaceae Preiuia Corda

quadrata (Scop) Nees

Marchantia L polymorpha L

28 Ricciaceae Riccwcarpus Corda

natans (L) Corda

Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm

15

i

CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust

MUSCI (Mosses)

29 Sphagnaceae SpIutgnum L

capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm

WtlI7IItorjii Russ wuljfanum Gira

30 Andreaeaceae AndNrrerI Hedw

rupeitrlr Oedw var pest

31 Fissident~e Fluid_ lledw

trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u

16

ABSTRACf

117This atlas of Wisconsin bryophytes shows the distribution of)30 taxa (137 hepatics 389 mosses)

on state maps with a dot in each county for which each taxon has been reported in the literature A list of synonyms and excluded taxa is included along with an extensive bibliography Pertinent features of the natural history and history of bryology in the state are discussed

ACKNOWLEDGEMENTS

We want to thank Drs Fred Hermann and John Thomson of the University of Wisconsin-Madison herbarium and Botany department for locating and lending packets and for references to aid us in accumushylating a more accurate complete report The illustration of Climacium dendroides on the cover was drawn by Robert Freckmann Of course we are grateful to all the people who have collected specimens of Wisshyconsin bryophytes and preserved them in herbariums making this distributional study possible

4

ONEIDA

LINCOLN

TAYLOR

t---------- MARATHON C1l-RK

SAu~

t

~

~ C i-W=AS_-r7-1

l=itHLANO

)

lOCK

INGTb

lt

WALWORTH

5

INTRODUCfION

This atlas of the bryophyte flora of Wisconsin is an attempt to show the distribution of all bryoshyphyte species known to occur in the state by county occurrenoes based on published reports in the literashyture It is not an annot~ted check list (only a feW problematic taxa were examined) but is a literature search intended as a foundation for a complete flora or for otherbryological research

The literature of btYo1ogy is large and spread over numerous journals reports monographs and manuals It would be almost impossible to search all sources for every reference attributing bryophyte species to Wisconsin and to particular localities in the state Wehav checked many inlportant references and paid special attention to works dealing with Wisconsiit botany and the bryologyof the upper Midwest Several bibliographies and indices were checked thoroughly including Green an~ Curtis (1955) the Bry ologist indices (Crum 1959 and Miller 1976) andt)le Wisconsin Academy of Sciences Arts and Letters indices

Only those published repots which app~red to)e substanti2ted by herbarium specimens and which gave- ~fficient location dat to P~CetheW(On in amiddot particula(county were-used in the initial compilation Species described in manuals in generalized way implying their occurrence in Wisconsin (Without menmiddot tioning Wisconsin specifically or citing a particular location in the state) were excluded eg Gnmmia raui (Wynne 1943) and Pohlia annotinll var decipiens (Crum 1976) Ecological papers which were either too general in taxonomic determinations (eg Sphagnum sp Polytrichum sp) or in location data were also excluded Some other taxa attributed to Wisconsin which have been excluded are

Cylindrothectum compr~ (Cheney 189tmiddotDane Co) identified later by Grout as Entodon seducshytrtx

Dicranell4 humilis (Holzjne~ 1925middotBuffalo Co) probablyD vllJia since Dhumilis is not found in North Arnerl~

Didymodon triarium (Foote 1 966-SW Visconsin) identified by Fred Hennann as Barbulaacuta Funoria microstoma (anonymous 1975middotDane Co) excluded because it is outside of its geographic

range and the packe~ QaMot be located Phil(motis calcarea (Cheney and Evans 1944-Dane Co) excluded in Crum Steere and Anderson

(1973) Plagiobryum demissum and P zierl (Cheney and Evans 1944-various counties) are both Arcticshy

alpine species not likely to be found in isconsin Two of Cheneys collections of Pl1giobyrum have been referred to Bryum

Pogonatum alpinum var septentrionale (Che~ey and Evans 1944-Chippewa Co) was misidentified (Ireland 1960) and is P umigerum

Rhizomnium fAPpoJachIanum and R punctalu1fl ssp chlorophyDosum (Koponen 1973middotseveral counshy ties) excluded North American bryologiflts have not recognized these taxa in America

Sphagnum tenellum (Cheney and Evans 1944-Waukesha Co)an Atlantic species not occurring in Wisconsin (Cruro 1975)

The nomenclature and systematic arrangement of hornwort 00 hepatic species in the atlas follow Stotler and Crandall-5totler (1977) and of moss species fonow Crum Steere and Anderson (1973) except in such instances as the JreptlIfockIdus fonns whiGh follow Wijk et aL (19S9middot1973) Species reported in the literature under names which are now listed as synonyms on either of these lists were converted to the accepted name A dot is placed in each county of Wisconsin from which the taxon has been reported The numbers after the name of the taxon correspond to the numbered references in the bibliography which giye ~act location data OEat least location data -s~fficient to place it in-particular counties in the state The numbers in parentheses indicate references which attribute the taxon to Wisconsin without providing exact loltation data

Fossil and subfossil reports are summarized Slfparately in our atlas only six species (Aulacomnium turgidum lJ1achythecium turgidum Ditrichum Jlelficaule Hypnum bambergti Scorpidium 6fJisities and Tortel14 1rIgi1i$) are known exclUsively from the fossil record ~

6

GEOGRAPHIC CLIMATIC AND BOTANICAL FEATURES OF WISCONSIN

Wisconsin is a north central state bounded on the east by Lake Michigan on the north by Michigan and Lake Superior on the south by Illinois and on the west by the Mississippi and St Croix rivers It is located midway between the Prime Meridian (0deg) of Longitude and 1800 Longitude and midway between the equator and the north pole thus placing it in the center of the northern half of the western hemisphere

The topography of the state is relatively flat a few hills exceed 1900 feet with Thns Hill at 1952 feet forming the highest point in Price County Lake Michigan represents the lowest point at 580 feet The northern and eastern two-thirds of the state were heavily glaciated during the last stage of the Pleistocene and now contain many of the best examples of continental glaciation topOgraphy including considerable ground moraine large terminal recessional and intetlobate moraines kames eskers drumlins and small depressions or kettle-holes Most of the nearly 9000 lakes of 20 acres or more lie within this part of the state The southwestern third of the state is the Driftless Area It includes most of the large flat generally sandy bed of Glacial Lake Wisconsin plus the highly dissected ridge and valley section with numerous cliffs and rock-outcroppings (Figure 1)

Much of the bedrock in the state lies below glacial till but exposures include igneous and metamiddot morphic rocks primarily granitic especially in the north as well as sedimentary layers primarily sandshystones and dolomitic limestones especially along Lake Michigan and in the DriftlesS Area

The climate of Wisconsin is classified as humid-temperate with cold winters The annual precipimiddot tation averages approximately 30 inches which is sufficiently even in its seasonal distribution to support forest vegetation throughout most of the state Nevertheless according to Curtis (I 959) the pre-settlement vegetation of the state (J 830-1850) included about 27 prairie and oak-savanna plus about 64 forests

The state is presently divided into seventy-two counties Marathon County is the largest (1586 sq mi) the smallest counties include Pepin (235 sq mi) Ozaukee (236 sq mi) andMilw8ukee (237 sq mi) We included Menominee County though its future status as a county is in doubt since it regained its Indian reservation status and rights in 1973 This county was established April 30 1961 from the Menomishynee Indian reservation which consisted of parts of Shawano and Oconto counties Wisconsin ranks 26th in the United States in area with 54464 square miles and 16th in population with 4651000

More ecological research is needed on the bryophyte flora of Wisconsin especially to relate it to the vegetation and to such other environmental features as soils climate and glaCial deposits Plant communishyties in Wisconsin are many and diverse these are summarized on Figure 2 from studies conducted by Curtis (1959) The state is divided into northern and southern floristic provinces by a tension zone deshyscribed by Curtis (1959) This tension zone is an eeotonal region vatying from 10middot31 mt wide where the ranges of many vascular plants overlap or terminatej it runs in an S-curve northeast from Milwaukee to Hudson in St Croix County (Figure 2) The principal plant communities of the northern province apshyproaching their southern boundaries within the zone include the northern conifermiddothardwood forests of sugar maple white pine hemlock and yellow birch and the Sphagnum bogs with tamarack and black spruce Southern communities at or near their northern limits within this zone include prairies oak and jack pine savannas and hardwood forests of maples oaks ash ehn and cottonwood

We hope to encourage research on cryptogamic ecology with our checklist We know that some species are northern while others are southern (eg cimacium) fits the tension zone concept Freckmann (I977) while Bryoxiphium is only found in the Driftless Area of the state Steere (1937) and Love and Love (1953) The tension zone theory may prove to be acceptable for bryophytes We speculate that certain bryophytes can indicate particular plant communities since many bryophytes are restricted to preshycise environmental requirements Many papers or theses have dealt with cryptogamic bark and rock ecology in Wisconsin (Culberson 1955 Foote 1962 Hale 1952 1955 and Cole 1975)

We hope that this atlas will stimulate collecting in counties poorly sampled so that our knowledge of the bryophytes can be increased It should aid anyone interested in floras for preservation environmental impact statements and distributional studies

7

A numerical summary of the bryophyte flora of the state based on our literature search is presented in Table 1 (below) This includes 517 taxa 70 families and 176 genera This numerical summary is comshypared with the published floras of Michigan mosses (Crum 1976) and Minnesota hepatics (Schuster 1953) Thus we surmise that further collecting could lead to the same approximate numbers for Wisconsin bryoshyphytes

Table 1 Comparison of Recorded Bryophytes of Wisconsin with Michigan and Minnesota and Summary of Wisconsin Taxa

Wisconsin Michigan (Crum 1976)

Minnesota (Schuster 1955)

Hepaticsamp Hornworts

Taxa Families Genera

137 28 47

174 32 52

Mosses

Taxa Families Genera

380 42

129

503 42

149

8

-

~

~ tIt

Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map

ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area

9

Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959

tit beol Forest e Phillie

Northernmiddot hardwood forest reg Pine 5cMInna

copy Southernmiddot hordwood forest CD Oak Savanna

10

SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES

ANTHOCEROTAE (Hornworts)

1 Anthocerotaceae Anthoceros 1 emnd Prosk

macounii M A Howe punctatum 1

Phaeoceros Prosk laevis (1) Prosk subsp laevis

Notothylas Sul1 orbicularis (Schwein) Sullo

HEPATICAE (liverworts)

2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum

trlchophyllum (L) Dum subsp trlchophylluni

3 Ptilidiaceae Ptilidium Nees

ci1iare (L) Hampe pulcherrimum (G Web) Hampe

4 Trichocoleaceae Trichocolea Dum nom cons

tomentella (Ehrh) Dum

5 Lepidoziaceae Kunia von Martens

setacea (Web) GroUe

Lepidozia (Dum) Dum reptans (1) Dum

Bazzania S Gray trilobata (1) S Gray var triZobata

6 Calypogejaceae 011ypogeja Raddi

flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda

11

7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum

biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps

Noweilia Mitt cunifolilz (Dicks) Mitt

8 Adelanthaceae Odontoschimuz (Dwn) Dum

denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw

c

9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$

divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD

10 Geo~c~ larptuttllus Nees

scutatus (Web et Mohr) Spruce

Geocalyx Nees grGVeolens (Schrad) Nees

11 Lophocoleaceae Lophocolea (Dum) Dum

heteropllyll4 (Schrad) Own minorNees

Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees

12 PlagiochiJaceae Plagio(hila Dum nom cons

tlBpienioides (L) Dum var asplenioidu

13 Jungermanniaceae Btriilophoril Loeske

attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske

12

Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones

Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg

Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum

Mylill S Gray nom cons anomala (Hook) S Gray

Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis

Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook

14 Scapaniaceae Diplophyllum (Dum) Dum nom cons

apiculatum (Evans) Steph var apiculatum

Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta

13

cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata

15 Radulaceae Raduia Dum nom COllS

compianata (L) Dum obconica Sull

16 Porellaceae Parella L

pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb

17 Jubulaceae FruUanill Raddi

bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt

18 lejeuneaceae Calolejeunea (Spruce) Schlffn

biddlecomiae (Aust) Evans

Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch

19 Codoniaceae Fossombronia Raddi

foveolata Lindb

20 Pa11aviciniaceae Paklvieinlil S Gray nom cons

lyeiii (Hook) Carruth

14

Moerckia Gott hibemica (Hook) Gott

21 PeWaceae Pellfa Raddi nom cons propos

endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr

22 Metzgeriaceae Metzgerla Raddi

conjugata Undb

23 Blasiacae B L

pulilm L

24 Aneurareae AneumDum

pinguis (L) Dum

Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth

25 Aytoniaceae Reboulill Raddi nom conI

hemllphaerica (L) Radeli

ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe

26 Conocephalaceae Conocephalum Wigg nom cons

conicum (L) Lindb

27 Marchantiaceae Preiuia Corda

quadrata (Scop) Nees

Marchantia L polymorpha L

28 Ricciaceae Riccwcarpus Corda

natans (L) Corda

Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm

15

i

CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust

MUSCI (Mosses)

29 Sphagnaceae SpIutgnum L

capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm

WtlI7IItorjii Russ wuljfanum Gira

30 Andreaeaceae AndNrrerI Hedw

rupeitrlr Oedw var pest

31 Fissident~e Fluid_ lledw

trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u

16

ONEIDA

LINCOLN

TAYLOR

t---------- MARATHON C1l-RK

SAu~

t

~

~ C i-W=AS_-r7-1

l=itHLANO

)

lOCK

INGTb

lt

WALWORTH

5

INTRODUCfION

This atlas of the bryophyte flora of Wisconsin is an attempt to show the distribution of all bryoshyphyte species known to occur in the state by county occurrenoes based on published reports in the literashyture It is not an annot~ted check list (only a feW problematic taxa were examined) but is a literature search intended as a foundation for a complete flora or for otherbryological research

The literature of btYo1ogy is large and spread over numerous journals reports monographs and manuals It would be almost impossible to search all sources for every reference attributing bryophyte species to Wisconsin and to particular localities in the state Wehav checked many inlportant references and paid special attention to works dealing with Wisconsiit botany and the bryologyof the upper Midwest Several bibliographies and indices were checked thoroughly including Green an~ Curtis (1955) the Bry ologist indices (Crum 1959 and Miller 1976) andt)le Wisconsin Academy of Sciences Arts and Letters indices

Only those published repots which app~red to)e substanti2ted by herbarium specimens and which gave- ~fficient location dat to P~CetheW(On in amiddot particula(county were-used in the initial compilation Species described in manuals in generalized way implying their occurrence in Wisconsin (Without menmiddot tioning Wisconsin specifically or citing a particular location in the state) were excluded eg Gnmmia raui (Wynne 1943) and Pohlia annotinll var decipiens (Crum 1976) Ecological papers which were either too general in taxonomic determinations (eg Sphagnum sp Polytrichum sp) or in location data were also excluded Some other taxa attributed to Wisconsin which have been excluded are

Cylindrothectum compr~ (Cheney 189tmiddotDane Co) identified later by Grout as Entodon seducshytrtx

Dicranell4 humilis (Holzjne~ 1925middotBuffalo Co) probablyD vllJia since Dhumilis is not found in North Arnerl~

Didymodon triarium (Foote 1 966-SW Visconsin) identified by Fred Hennann as Barbulaacuta Funoria microstoma (anonymous 1975middotDane Co) excluded because it is outside of its geographic

range and the packe~ QaMot be located Phil(motis calcarea (Cheney and Evans 1944-Dane Co) excluded in Crum Steere and Anderson

(1973) Plagiobryum demissum and P zierl (Cheney and Evans 1944-various counties) are both Arcticshy

alpine species not likely to be found in isconsin Two of Cheneys collections of Pl1giobyrum have been referred to Bryum

Pogonatum alpinum var septentrionale (Che~ey and Evans 1944-Chippewa Co) was misidentified (Ireland 1960) and is P umigerum

Rhizomnium fAPpoJachIanum and R punctalu1fl ssp chlorophyDosum (Koponen 1973middotseveral counshy ties) excluded North American bryologiflts have not recognized these taxa in America

Sphagnum tenellum (Cheney and Evans 1944-Waukesha Co)an Atlantic species not occurring in Wisconsin (Cruro 1975)

The nomenclature and systematic arrangement of hornwort 00 hepatic species in the atlas follow Stotler and Crandall-5totler (1977) and of moss species fonow Crum Steere and Anderson (1973) except in such instances as the JreptlIfockIdus fonns whiGh follow Wijk et aL (19S9middot1973) Species reported in the literature under names which are now listed as synonyms on either of these lists were converted to the accepted name A dot is placed in each county of Wisconsin from which the taxon has been reported The numbers after the name of the taxon correspond to the numbered references in the bibliography which giye ~act location data OEat least location data -s~fficient to place it in-particular counties in the state The numbers in parentheses indicate references which attribute the taxon to Wisconsin without providing exact loltation data

Fossil and subfossil reports are summarized Slfparately in our atlas only six species (Aulacomnium turgidum lJ1achythecium turgidum Ditrichum Jlelficaule Hypnum bambergti Scorpidium 6fJisities and Tortel14 1rIgi1i$) are known exclUsively from the fossil record ~

6

GEOGRAPHIC CLIMATIC AND BOTANICAL FEATURES OF WISCONSIN

Wisconsin is a north central state bounded on the east by Lake Michigan on the north by Michigan and Lake Superior on the south by Illinois and on the west by the Mississippi and St Croix rivers It is located midway between the Prime Meridian (0deg) of Longitude and 1800 Longitude and midway between the equator and the north pole thus placing it in the center of the northern half of the western hemisphere

The topography of the state is relatively flat a few hills exceed 1900 feet with Thns Hill at 1952 feet forming the highest point in Price County Lake Michigan represents the lowest point at 580 feet The northern and eastern two-thirds of the state were heavily glaciated during the last stage of the Pleistocene and now contain many of the best examples of continental glaciation topOgraphy including considerable ground moraine large terminal recessional and intetlobate moraines kames eskers drumlins and small depressions or kettle-holes Most of the nearly 9000 lakes of 20 acres or more lie within this part of the state The southwestern third of the state is the Driftless Area It includes most of the large flat generally sandy bed of Glacial Lake Wisconsin plus the highly dissected ridge and valley section with numerous cliffs and rock-outcroppings (Figure 1)

Much of the bedrock in the state lies below glacial till but exposures include igneous and metamiddot morphic rocks primarily granitic especially in the north as well as sedimentary layers primarily sandshystones and dolomitic limestones especially along Lake Michigan and in the DriftlesS Area

The climate of Wisconsin is classified as humid-temperate with cold winters The annual precipimiddot tation averages approximately 30 inches which is sufficiently even in its seasonal distribution to support forest vegetation throughout most of the state Nevertheless according to Curtis (I 959) the pre-settlement vegetation of the state (J 830-1850) included about 27 prairie and oak-savanna plus about 64 forests

The state is presently divided into seventy-two counties Marathon County is the largest (1586 sq mi) the smallest counties include Pepin (235 sq mi) Ozaukee (236 sq mi) andMilw8ukee (237 sq mi) We included Menominee County though its future status as a county is in doubt since it regained its Indian reservation status and rights in 1973 This county was established April 30 1961 from the Menomishynee Indian reservation which consisted of parts of Shawano and Oconto counties Wisconsin ranks 26th in the United States in area with 54464 square miles and 16th in population with 4651000

More ecological research is needed on the bryophyte flora of Wisconsin especially to relate it to the vegetation and to such other environmental features as soils climate and glaCial deposits Plant communishyties in Wisconsin are many and diverse these are summarized on Figure 2 from studies conducted by Curtis (1959) The state is divided into northern and southern floristic provinces by a tension zone deshyscribed by Curtis (1959) This tension zone is an eeotonal region vatying from 10middot31 mt wide where the ranges of many vascular plants overlap or terminatej it runs in an S-curve northeast from Milwaukee to Hudson in St Croix County (Figure 2) The principal plant communities of the northern province apshyproaching their southern boundaries within the zone include the northern conifermiddothardwood forests of sugar maple white pine hemlock and yellow birch and the Sphagnum bogs with tamarack and black spruce Southern communities at or near their northern limits within this zone include prairies oak and jack pine savannas and hardwood forests of maples oaks ash ehn and cottonwood

We hope to encourage research on cryptogamic ecology with our checklist We know that some species are northern while others are southern (eg cimacium) fits the tension zone concept Freckmann (I977) while Bryoxiphium is only found in the Driftless Area of the state Steere (1937) and Love and Love (1953) The tension zone theory may prove to be acceptable for bryophytes We speculate that certain bryophytes can indicate particular plant communities since many bryophytes are restricted to preshycise environmental requirements Many papers or theses have dealt with cryptogamic bark and rock ecology in Wisconsin (Culberson 1955 Foote 1962 Hale 1952 1955 and Cole 1975)

We hope that this atlas will stimulate collecting in counties poorly sampled so that our knowledge of the bryophytes can be increased It should aid anyone interested in floras for preservation environmental impact statements and distributional studies

7

A numerical summary of the bryophyte flora of the state based on our literature search is presented in Table 1 (below) This includes 517 taxa 70 families and 176 genera This numerical summary is comshypared with the published floras of Michigan mosses (Crum 1976) and Minnesota hepatics (Schuster 1953) Thus we surmise that further collecting could lead to the same approximate numbers for Wisconsin bryoshyphytes

Table 1 Comparison of Recorded Bryophytes of Wisconsin with Michigan and Minnesota and Summary of Wisconsin Taxa

Wisconsin Michigan (Crum 1976)

Minnesota (Schuster 1955)

Hepaticsamp Hornworts

Taxa Families Genera

137 28 47

174 32 52

Mosses

Taxa Families Genera

380 42

129

503 42

149

8

-

~

~ tIt

Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map

ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area

9

Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959

tit beol Forest e Phillie

Northernmiddot hardwood forest reg Pine 5cMInna

copy Southernmiddot hordwood forest CD Oak Savanna

10

SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES

ANTHOCEROTAE (Hornworts)

1 Anthocerotaceae Anthoceros 1 emnd Prosk

macounii M A Howe punctatum 1

Phaeoceros Prosk laevis (1) Prosk subsp laevis

Notothylas Sul1 orbicularis (Schwein) Sullo

HEPATICAE (liverworts)

2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum

trlchophyllum (L) Dum subsp trlchophylluni

3 Ptilidiaceae Ptilidium Nees

ci1iare (L) Hampe pulcherrimum (G Web) Hampe

4 Trichocoleaceae Trichocolea Dum nom cons

tomentella (Ehrh) Dum

5 Lepidoziaceae Kunia von Martens

setacea (Web) GroUe

Lepidozia (Dum) Dum reptans (1) Dum

Bazzania S Gray trilobata (1) S Gray var triZobata

6 Calypogejaceae 011ypogeja Raddi

flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda

11

7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum

biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps

Noweilia Mitt cunifolilz (Dicks) Mitt

8 Adelanthaceae Odontoschimuz (Dwn) Dum

denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw

c

9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$

divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD

10 Geo~c~ larptuttllus Nees

scutatus (Web et Mohr) Spruce

Geocalyx Nees grGVeolens (Schrad) Nees

11 Lophocoleaceae Lophocolea (Dum) Dum

heteropllyll4 (Schrad) Own minorNees

Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees

12 PlagiochiJaceae Plagio(hila Dum nom cons

tlBpienioides (L) Dum var asplenioidu

13 Jungermanniaceae Btriilophoril Loeske

attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske

12

Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones

Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg

Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum

Mylill S Gray nom cons anomala (Hook) S Gray

Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis

Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook

14 Scapaniaceae Diplophyllum (Dum) Dum nom cons

apiculatum (Evans) Steph var apiculatum

Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta

13

cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata

15 Radulaceae Raduia Dum nom COllS

compianata (L) Dum obconica Sull

16 Porellaceae Parella L

pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb

17 Jubulaceae FruUanill Raddi

bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt

18 lejeuneaceae Calolejeunea (Spruce) Schlffn

biddlecomiae (Aust) Evans

Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch

19 Codoniaceae Fossombronia Raddi

foveolata Lindb

20 Pa11aviciniaceae Paklvieinlil S Gray nom cons

lyeiii (Hook) Carruth

14

Moerckia Gott hibemica (Hook) Gott

21 PeWaceae Pellfa Raddi nom cons propos

endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr

22 Metzgeriaceae Metzgerla Raddi

conjugata Undb

23 Blasiacae B L

pulilm L

24 Aneurareae AneumDum

pinguis (L) Dum

Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth

25 Aytoniaceae Reboulill Raddi nom conI

hemllphaerica (L) Radeli

ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe

26 Conocephalaceae Conocephalum Wigg nom cons

conicum (L) Lindb

27 Marchantiaceae Preiuia Corda

quadrata (Scop) Nees

Marchantia L polymorpha L

28 Ricciaceae Riccwcarpus Corda

natans (L) Corda

Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm

15

i

CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust

MUSCI (Mosses)

29 Sphagnaceae SpIutgnum L

capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm

WtlI7IItorjii Russ wuljfanum Gira

30 Andreaeaceae AndNrrerI Hedw

rupeitrlr Oedw var pest

31 Fissident~e Fluid_ lledw

trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u

16

INTRODUCfION

This atlas of the bryophyte flora of Wisconsin is an attempt to show the distribution of all bryoshyphyte species known to occur in the state by county occurrenoes based on published reports in the literashyture It is not an annot~ted check list (only a feW problematic taxa were examined) but is a literature search intended as a foundation for a complete flora or for otherbryological research

The literature of btYo1ogy is large and spread over numerous journals reports monographs and manuals It would be almost impossible to search all sources for every reference attributing bryophyte species to Wisconsin and to particular localities in the state Wehav checked many inlportant references and paid special attention to works dealing with Wisconsiit botany and the bryologyof the upper Midwest Several bibliographies and indices were checked thoroughly including Green an~ Curtis (1955) the Bry ologist indices (Crum 1959 and Miller 1976) andt)le Wisconsin Academy of Sciences Arts and Letters indices

Only those published repots which app~red to)e substanti2ted by herbarium specimens and which gave- ~fficient location dat to P~CetheW(On in amiddot particula(county were-used in the initial compilation Species described in manuals in generalized way implying their occurrence in Wisconsin (Without menmiddot tioning Wisconsin specifically or citing a particular location in the state) were excluded eg Gnmmia raui (Wynne 1943) and Pohlia annotinll var decipiens (Crum 1976) Ecological papers which were either too general in taxonomic determinations (eg Sphagnum sp Polytrichum sp) or in location data were also excluded Some other taxa attributed to Wisconsin which have been excluded are

Cylindrothectum compr~ (Cheney 189tmiddotDane Co) identified later by Grout as Entodon seducshytrtx

Dicranell4 humilis (Holzjne~ 1925middotBuffalo Co) probablyD vllJia since Dhumilis is not found in North Arnerl~

Didymodon triarium (Foote 1 966-SW Visconsin) identified by Fred Hennann as Barbulaacuta Funoria microstoma (anonymous 1975middotDane Co) excluded because it is outside of its geographic

range and the packe~ QaMot be located Phil(motis calcarea (Cheney and Evans 1944-Dane Co) excluded in Crum Steere and Anderson

(1973) Plagiobryum demissum and P zierl (Cheney and Evans 1944-various counties) are both Arcticshy

alpine species not likely to be found in isconsin Two of Cheneys collections of Pl1giobyrum have been referred to Bryum

Pogonatum alpinum var septentrionale (Che~ey and Evans 1944-Chippewa Co) was misidentified (Ireland 1960) and is P umigerum

Rhizomnium fAPpoJachIanum and R punctalu1fl ssp chlorophyDosum (Koponen 1973middotseveral counshy ties) excluded North American bryologiflts have not recognized these taxa in America

Sphagnum tenellum (Cheney and Evans 1944-Waukesha Co)an Atlantic species not occurring in Wisconsin (Cruro 1975)

The nomenclature and systematic arrangement of hornwort 00 hepatic species in the atlas follow Stotler and Crandall-5totler (1977) and of moss species fonow Crum Steere and Anderson (1973) except in such instances as the JreptlIfockIdus fonns whiGh follow Wijk et aL (19S9middot1973) Species reported in the literature under names which are now listed as synonyms on either of these lists were converted to the accepted name A dot is placed in each county of Wisconsin from which the taxon has been reported The numbers after the name of the taxon correspond to the numbered references in the bibliography which giye ~act location data OEat least location data -s~fficient to place it in-particular counties in the state The numbers in parentheses indicate references which attribute the taxon to Wisconsin without providing exact loltation data

Fossil and subfossil reports are summarized Slfparately in our atlas only six species (Aulacomnium turgidum lJ1achythecium turgidum Ditrichum Jlelficaule Hypnum bambergti Scorpidium 6fJisities and Tortel14 1rIgi1i$) are known exclUsively from the fossil record ~

6

GEOGRAPHIC CLIMATIC AND BOTANICAL FEATURES OF WISCONSIN

Wisconsin is a north central state bounded on the east by Lake Michigan on the north by Michigan and Lake Superior on the south by Illinois and on the west by the Mississippi and St Croix rivers It is located midway between the Prime Meridian (0deg) of Longitude and 1800 Longitude and midway between the equator and the north pole thus placing it in the center of the northern half of the western hemisphere

The topography of the state is relatively flat a few hills exceed 1900 feet with Thns Hill at 1952 feet forming the highest point in Price County Lake Michigan represents the lowest point at 580 feet The northern and eastern two-thirds of the state were heavily glaciated during the last stage of the Pleistocene and now contain many of the best examples of continental glaciation topOgraphy including considerable ground moraine large terminal recessional and intetlobate moraines kames eskers drumlins and small depressions or kettle-holes Most of the nearly 9000 lakes of 20 acres or more lie within this part of the state The southwestern third of the state is the Driftless Area It includes most of the large flat generally sandy bed of Glacial Lake Wisconsin plus the highly dissected ridge and valley section with numerous cliffs and rock-outcroppings (Figure 1)

Much of the bedrock in the state lies below glacial till but exposures include igneous and metamiddot morphic rocks primarily granitic especially in the north as well as sedimentary layers primarily sandshystones and dolomitic limestones especially along Lake Michigan and in the DriftlesS Area

The climate of Wisconsin is classified as humid-temperate with cold winters The annual precipimiddot tation averages approximately 30 inches which is sufficiently even in its seasonal distribution to support forest vegetation throughout most of the state Nevertheless according to Curtis (I 959) the pre-settlement vegetation of the state (J 830-1850) included about 27 prairie and oak-savanna plus about 64 forests

The state is presently divided into seventy-two counties Marathon County is the largest (1586 sq mi) the smallest counties include Pepin (235 sq mi) Ozaukee (236 sq mi) andMilw8ukee (237 sq mi) We included Menominee County though its future status as a county is in doubt since it regained its Indian reservation status and rights in 1973 This county was established April 30 1961 from the Menomishynee Indian reservation which consisted of parts of Shawano and Oconto counties Wisconsin ranks 26th in the United States in area with 54464 square miles and 16th in population with 4651000

More ecological research is needed on the bryophyte flora of Wisconsin especially to relate it to the vegetation and to such other environmental features as soils climate and glaCial deposits Plant communishyties in Wisconsin are many and diverse these are summarized on Figure 2 from studies conducted by Curtis (1959) The state is divided into northern and southern floristic provinces by a tension zone deshyscribed by Curtis (1959) This tension zone is an eeotonal region vatying from 10middot31 mt wide where the ranges of many vascular plants overlap or terminatej it runs in an S-curve northeast from Milwaukee to Hudson in St Croix County (Figure 2) The principal plant communities of the northern province apshyproaching their southern boundaries within the zone include the northern conifermiddothardwood forests of sugar maple white pine hemlock and yellow birch and the Sphagnum bogs with tamarack and black spruce Southern communities at or near their northern limits within this zone include prairies oak and jack pine savannas and hardwood forests of maples oaks ash ehn and cottonwood

We hope to encourage research on cryptogamic ecology with our checklist We know that some species are northern while others are southern (eg cimacium) fits the tension zone concept Freckmann (I977) while Bryoxiphium is only found in the Driftless Area of the state Steere (1937) and Love and Love (1953) The tension zone theory may prove to be acceptable for bryophytes We speculate that certain bryophytes can indicate particular plant communities since many bryophytes are restricted to preshycise environmental requirements Many papers or theses have dealt with cryptogamic bark and rock ecology in Wisconsin (Culberson 1955 Foote 1962 Hale 1952 1955 and Cole 1975)

We hope that this atlas will stimulate collecting in counties poorly sampled so that our knowledge of the bryophytes can be increased It should aid anyone interested in floras for preservation environmental impact statements and distributional studies

7

A numerical summary of the bryophyte flora of the state based on our literature search is presented in Table 1 (below) This includes 517 taxa 70 families and 176 genera This numerical summary is comshypared with the published floras of Michigan mosses (Crum 1976) and Minnesota hepatics (Schuster 1953) Thus we surmise that further collecting could lead to the same approximate numbers for Wisconsin bryoshyphytes

Table 1 Comparison of Recorded Bryophytes of Wisconsin with Michigan and Minnesota and Summary of Wisconsin Taxa

Wisconsin Michigan (Crum 1976)

Minnesota (Schuster 1955)

Hepaticsamp Hornworts

Taxa Families Genera

137 28 47

174 32 52

Mosses

Taxa Families Genera

380 42

129

503 42

149

8

-

~

~ tIt

Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map

ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area

9

Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959

tit beol Forest e Phillie

Northernmiddot hardwood forest reg Pine 5cMInna

copy Southernmiddot hordwood forest CD Oak Savanna

10

SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES

ANTHOCEROTAE (Hornworts)

1 Anthocerotaceae Anthoceros 1 emnd Prosk

macounii M A Howe punctatum 1

Phaeoceros Prosk laevis (1) Prosk subsp laevis

Notothylas Sul1 orbicularis (Schwein) Sullo

HEPATICAE (liverworts)

2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum

trlchophyllum (L) Dum subsp trlchophylluni

3 Ptilidiaceae Ptilidium Nees

ci1iare (L) Hampe pulcherrimum (G Web) Hampe

4 Trichocoleaceae Trichocolea Dum nom cons

tomentella (Ehrh) Dum

5 Lepidoziaceae Kunia von Martens

setacea (Web) GroUe

Lepidozia (Dum) Dum reptans (1) Dum

Bazzania S Gray trilobata (1) S Gray var triZobata

6 Calypogejaceae 011ypogeja Raddi

flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda

11

7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum

biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps

Noweilia Mitt cunifolilz (Dicks) Mitt

8 Adelanthaceae Odontoschimuz (Dwn) Dum

denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw

c

9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$

divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD

10 Geo~c~ larptuttllus Nees

scutatus (Web et Mohr) Spruce

Geocalyx Nees grGVeolens (Schrad) Nees

11 Lophocoleaceae Lophocolea (Dum) Dum

heteropllyll4 (Schrad) Own minorNees

Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees

12 PlagiochiJaceae Plagio(hila Dum nom cons

tlBpienioides (L) Dum var asplenioidu

13 Jungermanniaceae Btriilophoril Loeske

attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske

12

Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones

Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg

Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum

Mylill S Gray nom cons anomala (Hook) S Gray

Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis

Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook

14 Scapaniaceae Diplophyllum (Dum) Dum nom cons

apiculatum (Evans) Steph var apiculatum

Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta

13

cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata

15 Radulaceae Raduia Dum nom COllS

compianata (L) Dum obconica Sull

16 Porellaceae Parella L

pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb

17 Jubulaceae FruUanill Raddi

bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt

18 lejeuneaceae Calolejeunea (Spruce) Schlffn

biddlecomiae (Aust) Evans

Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch

19 Codoniaceae Fossombronia Raddi

foveolata Lindb

20 Pa11aviciniaceae Paklvieinlil S Gray nom cons

lyeiii (Hook) Carruth

14

Moerckia Gott hibemica (Hook) Gott

21 PeWaceae Pellfa Raddi nom cons propos

endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr

22 Metzgeriaceae Metzgerla Raddi

conjugata Undb

23 Blasiacae B L

pulilm L

24 Aneurareae AneumDum

pinguis (L) Dum

Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth

25 Aytoniaceae Reboulill Raddi nom conI

hemllphaerica (L) Radeli

ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe

26 Conocephalaceae Conocephalum Wigg nom cons

conicum (L) Lindb

27 Marchantiaceae Preiuia Corda

quadrata (Scop) Nees

Marchantia L polymorpha L

28 Ricciaceae Riccwcarpus Corda

natans (L) Corda

Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm

15

i

CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust

MUSCI (Mosses)

29 Sphagnaceae SpIutgnum L

capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm

WtlI7IItorjii Russ wuljfanum Gira

30 Andreaeaceae AndNrrerI Hedw

rupeitrlr Oedw var pest

31 Fissident~e Fluid_ lledw

trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u

16

GEOGRAPHIC CLIMATIC AND BOTANICAL FEATURES OF WISCONSIN

Wisconsin is a north central state bounded on the east by Lake Michigan on the north by Michigan and Lake Superior on the south by Illinois and on the west by the Mississippi and St Croix rivers It is located midway between the Prime Meridian (0deg) of Longitude and 1800 Longitude and midway between the equator and the north pole thus placing it in the center of the northern half of the western hemisphere

The topography of the state is relatively flat a few hills exceed 1900 feet with Thns Hill at 1952 feet forming the highest point in Price County Lake Michigan represents the lowest point at 580 feet The northern and eastern two-thirds of the state were heavily glaciated during the last stage of the Pleistocene and now contain many of the best examples of continental glaciation topOgraphy including considerable ground moraine large terminal recessional and intetlobate moraines kames eskers drumlins and small depressions or kettle-holes Most of the nearly 9000 lakes of 20 acres or more lie within this part of the state The southwestern third of the state is the Driftless Area It includes most of the large flat generally sandy bed of Glacial Lake Wisconsin plus the highly dissected ridge and valley section with numerous cliffs and rock-outcroppings (Figure 1)

Much of the bedrock in the state lies below glacial till but exposures include igneous and metamiddot morphic rocks primarily granitic especially in the north as well as sedimentary layers primarily sandshystones and dolomitic limestones especially along Lake Michigan and in the DriftlesS Area

The climate of Wisconsin is classified as humid-temperate with cold winters The annual precipimiddot tation averages approximately 30 inches which is sufficiently even in its seasonal distribution to support forest vegetation throughout most of the state Nevertheless according to Curtis (I 959) the pre-settlement vegetation of the state (J 830-1850) included about 27 prairie and oak-savanna plus about 64 forests

The state is presently divided into seventy-two counties Marathon County is the largest (1586 sq mi) the smallest counties include Pepin (235 sq mi) Ozaukee (236 sq mi) andMilw8ukee (237 sq mi) We included Menominee County though its future status as a county is in doubt since it regained its Indian reservation status and rights in 1973 This county was established April 30 1961 from the Menomishynee Indian reservation which consisted of parts of Shawano and Oconto counties Wisconsin ranks 26th in the United States in area with 54464 square miles and 16th in population with 4651000

More ecological research is needed on the bryophyte flora of Wisconsin especially to relate it to the vegetation and to such other environmental features as soils climate and glaCial deposits Plant communishyties in Wisconsin are many and diverse these are summarized on Figure 2 from studies conducted by Curtis (1959) The state is divided into northern and southern floristic provinces by a tension zone deshyscribed by Curtis (1959) This tension zone is an eeotonal region vatying from 10middot31 mt wide where the ranges of many vascular plants overlap or terminatej it runs in an S-curve northeast from Milwaukee to Hudson in St Croix County (Figure 2) The principal plant communities of the northern province apshyproaching their southern boundaries within the zone include the northern conifermiddothardwood forests of sugar maple white pine hemlock and yellow birch and the Sphagnum bogs with tamarack and black spruce Southern communities at or near their northern limits within this zone include prairies oak and jack pine savannas and hardwood forests of maples oaks ash ehn and cottonwood

We hope to encourage research on cryptogamic ecology with our checklist We know that some species are northern while others are southern (eg cimacium) fits the tension zone concept Freckmann (I977) while Bryoxiphium is only found in the Driftless Area of the state Steere (1937) and Love and Love (1953) The tension zone theory may prove to be acceptable for bryophytes We speculate that certain bryophytes can indicate particular plant communities since many bryophytes are restricted to preshycise environmental requirements Many papers or theses have dealt with cryptogamic bark and rock ecology in Wisconsin (Culberson 1955 Foote 1962 Hale 1952 1955 and Cole 1975)

We hope that this atlas will stimulate collecting in counties poorly sampled so that our knowledge of the bryophytes can be increased It should aid anyone interested in floras for preservation environmental impact statements and distributional studies

7

A numerical summary of the bryophyte flora of the state based on our literature search is presented in Table 1 (below) This includes 517 taxa 70 families and 176 genera This numerical summary is comshypared with the published floras of Michigan mosses (Crum 1976) and Minnesota hepatics (Schuster 1953) Thus we surmise that further collecting could lead to the same approximate numbers for Wisconsin bryoshyphytes

Table 1 Comparison of Recorded Bryophytes of Wisconsin with Michigan and Minnesota and Summary of Wisconsin Taxa

Wisconsin Michigan (Crum 1976)

Minnesota (Schuster 1955)

Hepaticsamp Hornworts

Taxa Families Genera

137 28 47

174 32 52

Mosses

Taxa Families Genera

380 42

129

503 42

149

8

-

~

~ tIt

Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map

ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area

9

Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959

tit beol Forest e Phillie

Northernmiddot hardwood forest reg Pine 5cMInna

copy Southernmiddot hordwood forest CD Oak Savanna

10

SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES

ANTHOCEROTAE (Hornworts)

1 Anthocerotaceae Anthoceros 1 emnd Prosk

macounii M A Howe punctatum 1

Phaeoceros Prosk laevis (1) Prosk subsp laevis

Notothylas Sul1 orbicularis (Schwein) Sullo

HEPATICAE (liverworts)

2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum

trlchophyllum (L) Dum subsp trlchophylluni

3 Ptilidiaceae Ptilidium Nees

ci1iare (L) Hampe pulcherrimum (G Web) Hampe

4 Trichocoleaceae Trichocolea Dum nom cons

tomentella (Ehrh) Dum

5 Lepidoziaceae Kunia von Martens

setacea (Web) GroUe

Lepidozia (Dum) Dum reptans (1) Dum

Bazzania S Gray trilobata (1) S Gray var triZobata

6 Calypogejaceae 011ypogeja Raddi

flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda

11

7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum

biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps

Noweilia Mitt cunifolilz (Dicks) Mitt

8 Adelanthaceae Odontoschimuz (Dwn) Dum

denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw

c

9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$

divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD

10 Geo~c~ larptuttllus Nees

scutatus (Web et Mohr) Spruce

Geocalyx Nees grGVeolens (Schrad) Nees

11 Lophocoleaceae Lophocolea (Dum) Dum

heteropllyll4 (Schrad) Own minorNees

Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees

12 PlagiochiJaceae Plagio(hila Dum nom cons

tlBpienioides (L) Dum var asplenioidu

13 Jungermanniaceae Btriilophoril Loeske

attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske

12

Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones

Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg

Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum

Mylill S Gray nom cons anomala (Hook) S Gray

Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis

Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook

14 Scapaniaceae Diplophyllum (Dum) Dum nom cons

apiculatum (Evans) Steph var apiculatum

Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta

13

cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata

15 Radulaceae Raduia Dum nom COllS

compianata (L) Dum obconica Sull

16 Porellaceae Parella L

pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb

17 Jubulaceae FruUanill Raddi

bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt

18 lejeuneaceae Calolejeunea (Spruce) Schlffn

biddlecomiae (Aust) Evans

Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch

19 Codoniaceae Fossombronia Raddi

foveolata Lindb

20 Pa11aviciniaceae Paklvieinlil S Gray nom cons

lyeiii (Hook) Carruth

14

Moerckia Gott hibemica (Hook) Gott

21 PeWaceae Pellfa Raddi nom cons propos

endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr

22 Metzgeriaceae Metzgerla Raddi

conjugata Undb

23 Blasiacae B L

pulilm L

24 Aneurareae AneumDum

pinguis (L) Dum

Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth

25 Aytoniaceae Reboulill Raddi nom conI

hemllphaerica (L) Radeli

ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe

26 Conocephalaceae Conocephalum Wigg nom cons

conicum (L) Lindb

27 Marchantiaceae Preiuia Corda

quadrata (Scop) Nees

Marchantia L polymorpha L

28 Ricciaceae Riccwcarpus Corda

natans (L) Corda

Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm

15

i

CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust

MUSCI (Mosses)

29 Sphagnaceae SpIutgnum L

capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm

WtlI7IItorjii Russ wuljfanum Gira

30 Andreaeaceae AndNrrerI Hedw

rupeitrlr Oedw var pest

31 Fissident~e Fluid_ lledw

trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u

16

A numerical summary of the bryophyte flora of the state based on our literature search is presented in Table 1 (below) This includes 517 taxa 70 families and 176 genera This numerical summary is comshypared with the published floras of Michigan mosses (Crum 1976) and Minnesota hepatics (Schuster 1953) Thus we surmise that further collecting could lead to the same approximate numbers for Wisconsin bryoshyphytes

Table 1 Comparison of Recorded Bryophytes of Wisconsin with Michigan and Minnesota and Summary of Wisconsin Taxa

Wisconsin Michigan (Crum 1976)

Minnesota (Schuster 1955)

Hepaticsamp Hornworts

Taxa Families Genera

137 28 47

174 32 52

Mosses

Taxa Families Genera

380 42

129

503 42

149

8

-

~

~ tIt

Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map

ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area

9

Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959

tit beol Forest e Phillie

Northernmiddot hardwood forest reg Pine 5cMInna

copy Southernmiddot hordwood forest CD Oak Savanna

10

SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES

ANTHOCEROTAE (Hornworts)

1 Anthocerotaceae Anthoceros 1 emnd Prosk

macounii M A Howe punctatum 1

Phaeoceros Prosk laevis (1) Prosk subsp laevis

Notothylas Sul1 orbicularis (Schwein) Sullo

HEPATICAE (liverworts)

2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum

trlchophyllum (L) Dum subsp trlchophylluni

3 Ptilidiaceae Ptilidium Nees

ci1iare (L) Hampe pulcherrimum (G Web) Hampe

4 Trichocoleaceae Trichocolea Dum nom cons

tomentella (Ehrh) Dum

5 Lepidoziaceae Kunia von Martens

setacea (Web) GroUe

Lepidozia (Dum) Dum reptans (1) Dum

Bazzania S Gray trilobata (1) S Gray var triZobata

6 Calypogejaceae 011ypogeja Raddi

flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda

11

7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum

biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps

Noweilia Mitt cunifolilz (Dicks) Mitt

8 Adelanthaceae Odontoschimuz (Dwn) Dum

denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw

c

9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$

divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD

10 Geo~c~ larptuttllus Nees

scutatus (Web et Mohr) Spruce

Geocalyx Nees grGVeolens (Schrad) Nees

11 Lophocoleaceae Lophocolea (Dum) Dum

heteropllyll4 (Schrad) Own minorNees

Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees

12 PlagiochiJaceae Plagio(hila Dum nom cons

tlBpienioides (L) Dum var asplenioidu

13 Jungermanniaceae Btriilophoril Loeske

attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske

12

Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones

Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg

Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum

Mylill S Gray nom cons anomala (Hook) S Gray

Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis

Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook

14 Scapaniaceae Diplophyllum (Dum) Dum nom cons

apiculatum (Evans) Steph var apiculatum

Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta

13

cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata

15 Radulaceae Raduia Dum nom COllS

compianata (L) Dum obconica Sull

16 Porellaceae Parella L

pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb

17 Jubulaceae FruUanill Raddi

bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt

18 lejeuneaceae Calolejeunea (Spruce) Schlffn

biddlecomiae (Aust) Evans

Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch

19 Codoniaceae Fossombronia Raddi

foveolata Lindb

20 Pa11aviciniaceae Paklvieinlil S Gray nom cons

lyeiii (Hook) Carruth

14

Moerckia Gott hibemica (Hook) Gott

21 PeWaceae Pellfa Raddi nom cons propos

endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr

22 Metzgeriaceae Metzgerla Raddi

conjugata Undb

23 Blasiacae B L

pulilm L

24 Aneurareae AneumDum

pinguis (L) Dum

Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth

25 Aytoniaceae Reboulill Raddi nom conI

hemllphaerica (L) Radeli

ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe

26 Conocephalaceae Conocephalum Wigg nom cons

conicum (L) Lindb

27 Marchantiaceae Preiuia Corda

quadrata (Scop) Nees

Marchantia L polymorpha L

28 Ricciaceae Riccwcarpus Corda

natans (L) Corda

Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm

15

i

CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust

MUSCI (Mosses)

29 Sphagnaceae SpIutgnum L

capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm

WtlI7IItorjii Russ wuljfanum Gira

30 Andreaeaceae AndNrrerI Hedw

rupeitrlr Oedw var pest

31 Fissident~e Fluid_ lledw

trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u

16

-

~

~ tIt

Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map

ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area

9

Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959

tit beol Forest e Phillie

Northernmiddot hardwood forest reg Pine 5cMInna

copy Southernmiddot hordwood forest CD Oak Savanna

10

SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES

ANTHOCEROTAE (Hornworts)

1 Anthocerotaceae Anthoceros 1 emnd Prosk

macounii M A Howe punctatum 1

Phaeoceros Prosk laevis (1) Prosk subsp laevis

Notothylas Sul1 orbicularis (Schwein) Sullo

HEPATICAE (liverworts)

2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum

trlchophyllum (L) Dum subsp trlchophylluni

3 Ptilidiaceae Ptilidium Nees

ci1iare (L) Hampe pulcherrimum (G Web) Hampe

4 Trichocoleaceae Trichocolea Dum nom cons

tomentella (Ehrh) Dum

5 Lepidoziaceae Kunia von Martens

setacea (Web) GroUe

Lepidozia (Dum) Dum reptans (1) Dum

Bazzania S Gray trilobata (1) S Gray var triZobata

6 Calypogejaceae 011ypogeja Raddi

flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda

11

7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum

biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps

Noweilia Mitt cunifolilz (Dicks) Mitt

8 Adelanthaceae Odontoschimuz (Dwn) Dum

denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw

c

9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$

divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD

10 Geo~c~ larptuttllus Nees

scutatus (Web et Mohr) Spruce

Geocalyx Nees grGVeolens (Schrad) Nees

11 Lophocoleaceae Lophocolea (Dum) Dum

heteropllyll4 (Schrad) Own minorNees

Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees

12 PlagiochiJaceae Plagio(hila Dum nom cons

tlBpienioides (L) Dum var asplenioidu

13 Jungermanniaceae Btriilophoril Loeske

attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske

12

Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones

Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg

Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum

Mylill S Gray nom cons anomala (Hook) S Gray

Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis

Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook

14 Scapaniaceae Diplophyllum (Dum) Dum nom cons

apiculatum (Evans) Steph var apiculatum

Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta

13

cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata

15 Radulaceae Raduia Dum nom COllS

compianata (L) Dum obconica Sull

16 Porellaceae Parella L

pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb

17 Jubulaceae FruUanill Raddi

bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt

18 lejeuneaceae Calolejeunea (Spruce) Schlffn

biddlecomiae (Aust) Evans

Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch

19 Codoniaceae Fossombronia Raddi

foveolata Lindb

20 Pa11aviciniaceae Paklvieinlil S Gray nom cons

lyeiii (Hook) Carruth

14

Moerckia Gott hibemica (Hook) Gott

21 PeWaceae Pellfa Raddi nom cons propos

endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr

22 Metzgeriaceae Metzgerla Raddi

conjugata Undb

23 Blasiacae B L

pulilm L

24 Aneurareae AneumDum

pinguis (L) Dum

Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth

25 Aytoniaceae Reboulill Raddi nom conI

hemllphaerica (L) Radeli

ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe

26 Conocephalaceae Conocephalum Wigg nom cons

conicum (L) Lindb

27 Marchantiaceae Preiuia Corda

quadrata (Scop) Nees

Marchantia L polymorpha L

28 Ricciaceae Riccwcarpus Corda

natans (L) Corda

Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm

15

i

CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust

MUSCI (Mosses)

29 Sphagnaceae SpIutgnum L

capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm

WtlI7IItorjii Russ wuljfanum Gira

30 Andreaeaceae AndNrrerI Hedw

rupeitrlr Oedw var pest

31 Fissident~e Fluid_ lledw

trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u

16

Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959

tit beol Forest e Phillie

Northernmiddot hardwood forest reg Pine 5cMInna

copy Southernmiddot hordwood forest CD Oak Savanna

10

SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES

ANTHOCEROTAE (Hornworts)

1 Anthocerotaceae Anthoceros 1 emnd Prosk

macounii M A Howe punctatum 1

Phaeoceros Prosk laevis (1) Prosk subsp laevis

Notothylas Sul1 orbicularis (Schwein) Sullo

HEPATICAE (liverworts)

2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum

trlchophyllum (L) Dum subsp trlchophylluni

3 Ptilidiaceae Ptilidium Nees

ci1iare (L) Hampe pulcherrimum (G Web) Hampe

4 Trichocoleaceae Trichocolea Dum nom cons

tomentella (Ehrh) Dum

5 Lepidoziaceae Kunia von Martens

setacea (Web) GroUe

Lepidozia (Dum) Dum reptans (1) Dum

Bazzania S Gray trilobata (1) S Gray var triZobata

6 Calypogejaceae 011ypogeja Raddi

flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda

11

7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum

biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps

Noweilia Mitt cunifolilz (Dicks) Mitt

8 Adelanthaceae Odontoschimuz (Dwn) Dum

denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw

c

9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$

divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD

10 Geo~c~ larptuttllus Nees

scutatus (Web et Mohr) Spruce

Geocalyx Nees grGVeolens (Schrad) Nees

11 Lophocoleaceae Lophocolea (Dum) Dum

heteropllyll4 (Schrad) Own minorNees

Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees

12 PlagiochiJaceae Plagio(hila Dum nom cons

tlBpienioides (L) Dum var asplenioidu

13 Jungermanniaceae Btriilophoril Loeske

attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske

12

Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones

Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg

Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum

Mylill S Gray nom cons anomala (Hook) S Gray

Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis

Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook

14 Scapaniaceae Diplophyllum (Dum) Dum nom cons

apiculatum (Evans) Steph var apiculatum

Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta

13

cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata

15 Radulaceae Raduia Dum nom COllS

compianata (L) Dum obconica Sull

16 Porellaceae Parella L

pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb

17 Jubulaceae FruUanill Raddi

bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt

18 lejeuneaceae Calolejeunea (Spruce) Schlffn

biddlecomiae (Aust) Evans

Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch

19 Codoniaceae Fossombronia Raddi

foveolata Lindb

20 Pa11aviciniaceae Paklvieinlil S Gray nom cons

lyeiii (Hook) Carruth

14

Moerckia Gott hibemica (Hook) Gott

21 PeWaceae Pellfa Raddi nom cons propos

endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr

22 Metzgeriaceae Metzgerla Raddi

conjugata Undb

23 Blasiacae B L

pulilm L

24 Aneurareae AneumDum

pinguis (L) Dum

Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth

25 Aytoniaceae Reboulill Raddi nom conI

hemllphaerica (L) Radeli

ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe

26 Conocephalaceae Conocephalum Wigg nom cons

conicum (L) Lindb

27 Marchantiaceae Preiuia Corda

quadrata (Scop) Nees

Marchantia L polymorpha L

28 Ricciaceae Riccwcarpus Corda

natans (L) Corda

Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm

15

i

CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust

MUSCI (Mosses)

29 Sphagnaceae SpIutgnum L

capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm

WtlI7IItorjii Russ wuljfanum Gira

30 Andreaeaceae AndNrrerI Hedw

rupeitrlr Oedw var pest

31 Fissident~e Fluid_ lledw

trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u

16

SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES

ANTHOCEROTAE (Hornworts)

1 Anthocerotaceae Anthoceros 1 emnd Prosk

macounii M A Howe punctatum 1

Phaeoceros Prosk laevis (1) Prosk subsp laevis

Notothylas Sul1 orbicularis (Schwein) Sullo

HEPATICAE (liverworts)

2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum

trlchophyllum (L) Dum subsp trlchophylluni

3 Ptilidiaceae Ptilidium Nees

ci1iare (L) Hampe pulcherrimum (G Web) Hampe

4 Trichocoleaceae Trichocolea Dum nom cons

tomentella (Ehrh) Dum

5 Lepidoziaceae Kunia von Martens

setacea (Web) GroUe

Lepidozia (Dum) Dum reptans (1) Dum

Bazzania S Gray trilobata (1) S Gray var triZobata

6 Calypogejaceae 011ypogeja Raddi

flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda

11

7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum

biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps

Noweilia Mitt cunifolilz (Dicks) Mitt

8 Adelanthaceae Odontoschimuz (Dwn) Dum

denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw

c

9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$

divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD

10 Geo~c~ larptuttllus Nees

scutatus (Web et Mohr) Spruce

Geocalyx Nees grGVeolens (Schrad) Nees

11 Lophocoleaceae Lophocolea (Dum) Dum

heteropllyll4 (Schrad) Own minorNees

Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees

12 PlagiochiJaceae Plagio(hila Dum nom cons

tlBpienioides (L) Dum var asplenioidu

13 Jungermanniaceae Btriilophoril Loeske

attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske

12

Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones

Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg

Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum

Mylill S Gray nom cons anomala (Hook) S Gray

Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis

Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook

14 Scapaniaceae Diplophyllum (Dum) Dum nom cons

apiculatum (Evans) Steph var apiculatum

Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta

13

cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata

15 Radulaceae Raduia Dum nom COllS

compianata (L) Dum obconica Sull

16 Porellaceae Parella L

pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb

17 Jubulaceae FruUanill Raddi

bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt

18 lejeuneaceae Calolejeunea (Spruce) Schlffn

biddlecomiae (Aust) Evans

Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch

19 Codoniaceae Fossombronia Raddi

foveolata Lindb

20 Pa11aviciniaceae Paklvieinlil S Gray nom cons

lyeiii (Hook) Carruth

14

Moerckia Gott hibemica (Hook) Gott

21 PeWaceae Pellfa Raddi nom cons propos

endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr

22 Metzgeriaceae Metzgerla Raddi

conjugata Undb

23 Blasiacae B L

pulilm L

24 Aneurareae AneumDum

pinguis (L) Dum

Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth

25 Aytoniaceae Reboulill Raddi nom conI

hemllphaerica (L) Radeli

ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe

26 Conocephalaceae Conocephalum Wigg nom cons

conicum (L) Lindb

27 Marchantiaceae Preiuia Corda

quadrata (Scop) Nees

Marchantia L polymorpha L

28 Ricciaceae Riccwcarpus Corda

natans (L) Corda

Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm

15

i

CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust

MUSCI (Mosses)

29 Sphagnaceae SpIutgnum L

capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm

WtlI7IItorjii Russ wuljfanum Gira

30 Andreaeaceae AndNrrerI Hedw

rupeitrlr Oedw var pest

31 Fissident~e Fluid_ lledw

trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u

16

7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum

biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps

Noweilia Mitt cunifolilz (Dicks) Mitt

8 Adelanthaceae Odontoschimuz (Dwn) Dum

denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw

c

9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$

divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD

10 Geo~c~ larptuttllus Nees

scutatus (Web et Mohr) Spruce

Geocalyx Nees grGVeolens (Schrad) Nees

11 Lophocoleaceae Lophocolea (Dum) Dum

heteropllyll4 (Schrad) Own minorNees

Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees

12 PlagiochiJaceae Plagio(hila Dum nom cons

tlBpienioides (L) Dum var asplenioidu

13 Jungermanniaceae Btriilophoril Loeske

attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske

12

Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones

Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg

Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum

Mylill S Gray nom cons anomala (Hook) S Gray

Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis

Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook

14 Scapaniaceae Diplophyllum (Dum) Dum nom cons

apiculatum (Evans) Steph var apiculatum

Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta

13

cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata

15 Radulaceae Raduia Dum nom COllS

compianata (L) Dum obconica Sull

16 Porellaceae Parella L

pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb

17 Jubulaceae FruUanill Raddi

bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt

18 lejeuneaceae Calolejeunea (Spruce) Schlffn

biddlecomiae (Aust) Evans

Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch

19 Codoniaceae Fossombronia Raddi

foveolata Lindb

20 Pa11aviciniaceae Paklvieinlil S Gray nom cons

lyeiii (Hook) Carruth

14

Moerckia Gott hibemica (Hook) Gott

21 PeWaceae Pellfa Raddi nom cons propos

endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr

22 Metzgeriaceae Metzgerla Raddi

conjugata Undb

23 Blasiacae B L

pulilm L

24 Aneurareae AneumDum

pinguis (L) Dum

Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth

25 Aytoniaceae Reboulill Raddi nom conI

hemllphaerica (L) Radeli

ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe

26 Conocephalaceae Conocephalum Wigg nom cons

conicum (L) Lindb

27 Marchantiaceae Preiuia Corda

quadrata (Scop) Nees

Marchantia L polymorpha L

28 Ricciaceae Riccwcarpus Corda

natans (L) Corda

Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm

15

i

CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust

MUSCI (Mosses)

29 Sphagnaceae SpIutgnum L

capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm

WtlI7IItorjii Russ wuljfanum Gira

30 Andreaeaceae AndNrrerI Hedw

rupeitrlr Oedw var pest

31 Fissident~e Fluid_ lledw

trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u

16

Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones

Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg

Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum

Mylill S Gray nom cons anomala (Hook) S Gray

Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis

Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook

14 Scapaniaceae Diplophyllum (Dum) Dum nom cons

apiculatum (Evans) Steph var apiculatum

Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta

13

cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata

15 Radulaceae Raduia Dum nom COllS

compianata (L) Dum obconica Sull

16 Porellaceae Parella L

pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb

17 Jubulaceae FruUanill Raddi

bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt

18 lejeuneaceae Calolejeunea (Spruce) Schlffn

biddlecomiae (Aust) Evans

Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch

19 Codoniaceae Fossombronia Raddi

foveolata Lindb

20 Pa11aviciniaceae Paklvieinlil S Gray nom cons

lyeiii (Hook) Carruth

14

Moerckia Gott hibemica (Hook) Gott

21 PeWaceae Pellfa Raddi nom cons propos

endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr

22 Metzgeriaceae Metzgerla Raddi

conjugata Undb

23 Blasiacae B L

pulilm L

24 Aneurareae AneumDum

pinguis (L) Dum

Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth

25 Aytoniaceae Reboulill Raddi nom conI

hemllphaerica (L) Radeli

ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe

26 Conocephalaceae Conocephalum Wigg nom cons

conicum (L) Lindb

27 Marchantiaceae Preiuia Corda

quadrata (Scop) Nees

Marchantia L polymorpha L

28 Ricciaceae Riccwcarpus Corda

natans (L) Corda

Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm

15

i

CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust

MUSCI (Mosses)

29 Sphagnaceae SpIutgnum L

capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm

WtlI7IItorjii Russ wuljfanum Gira

30 Andreaeaceae AndNrrerI Hedw

rupeitrlr Oedw var pest

31 Fissident~e Fluid_ lledw

trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u

16

cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata

15 Radulaceae Raduia Dum nom COllS

compianata (L) Dum obconica Sull

16 Porellaceae Parella L

pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb

17 Jubulaceae FruUanill Raddi

bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt

18 lejeuneaceae Calolejeunea (Spruce) Schlffn

biddlecomiae (Aust) Evans

Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch

19 Codoniaceae Fossombronia Raddi

foveolata Lindb

20 Pa11aviciniaceae Paklvieinlil S Gray nom cons

lyeiii (Hook) Carruth

14

Moerckia Gott hibemica (Hook) Gott

21 PeWaceae Pellfa Raddi nom cons propos

endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr

22 Metzgeriaceae Metzgerla Raddi

conjugata Undb

23 Blasiacae B L

pulilm L

24 Aneurareae AneumDum

pinguis (L) Dum

Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth

25 Aytoniaceae Reboulill Raddi nom conI

hemllphaerica (L) Radeli

ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe

26 Conocephalaceae Conocephalum Wigg nom cons

conicum (L) Lindb

27 Marchantiaceae Preiuia Corda

quadrata (Scop) Nees

Marchantia L polymorpha L

28 Ricciaceae Riccwcarpus Corda

natans (L) Corda

Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm

15

i

CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust

MUSCI (Mosses)

29 Sphagnaceae SpIutgnum L

capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm

WtlI7IItorjii Russ wuljfanum Gira

30 Andreaeaceae AndNrrerI Hedw

rupeitrlr Oedw var pest

31 Fissident~e Fluid_ lledw

trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u

16

Moerckia Gott hibemica (Hook) Gott

21 PeWaceae Pellfa Raddi nom cons propos

endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr

22 Metzgeriaceae Metzgerla Raddi

conjugata Undb

23 Blasiacae B L

pulilm L

24 Aneurareae AneumDum

pinguis (L) Dum

Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth

25 Aytoniaceae Reboulill Raddi nom conI

hemllphaerica (L) Radeli

ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe

26 Conocephalaceae Conocephalum Wigg nom cons

conicum (L) Lindb

27 Marchantiaceae Preiuia Corda

quadrata (Scop) Nees

Marchantia L polymorpha L

28 Ricciaceae Riccwcarpus Corda

natans (L) Corda

Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm

15

i

CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust

MUSCI (Mosses)

29 Sphagnaceae SpIutgnum L

capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm

WtlI7IItorjii Russ wuljfanum Gira

30 Andreaeaceae AndNrrerI Hedw

rupeitrlr Oedw var pest

31 Fissident~e Fluid_ lledw

trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u

16

CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust

MUSCI (Mosses)

29 Sphagnaceae SpIutgnum L

capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm

WtlI7IItorjii Russ wuljfanum Gira

30 Andreaeaceae AndNrrerI Hedw

rupeitrlr Oedw var pest

31 Fissident~e Fluid_ lledw

trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u

16

32 Ditrichaceae Pleuridium Brid

subulatum (Hedw) Rabenh

Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb

~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe

Saelania Lindb gIlucescens (Hedw) Bomanss et Broth

Ceratodon Brid purpureus (He(lw) Brid var prupurew

Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG

33 Bryoxipbiaceae Bryoxlphium Mitt

norvegicum (Brid) Mitt

34 Seligeriaceae Seligeria BSG

cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG

Bindia BSG acuta (Hedw) BSG

35 Dicranaceae BlUehia Schwegr

lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust

Trematodon Michx ambiguus (Hedw) Hornsch

Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata

heteromalla (Hedw) Schimp ~ rufescens (With) Schimp

schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp

Dicranodontium BSG denudatum (Brid) Britt ex Williams

Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium

17

Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr

Hyophila Brid involuta (Hoole) Jaeg et Sauerb

Didymodon Hedw rigidulus Hedw

BryoerythrophyUum Chen recurvirostnon (Hedw) Chen

Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw

Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum

PJl~lm Hedw CUSPidtltum Hedw

De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp

T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis

39 Grimmiaceae Grlmmill Hedw

alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr

18

Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr

Hyophila Brid involuta (Hoole) Jaeg et Sauerb

Didymodon Hedw rigidulus Hedw

BryoerythrophyUum Chen recurvirostnon (Hedw) Chen

Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw

Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum

PJl~lm Hedw CUSPidtltum Hedw

De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp

T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis

39 Grimmiaceae Grlmmill Hedw

alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr

18

Orothera C Muell leana (Sull) C Muell

Rfwbdoweisia crispata (With) Undb

Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb

Dichodontium Schimp pellucidum (Hedw) Schimp

Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)

Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb

36 Leucobryaceae Leucobryum Hampe

glaucum (Hedw) Angstr ex Fr

37 Encalyptaceae Encalypta Hedw

ciliata Hedw var ciliata procera Bruch

38 Pottiaceae Astomum Hampe

muhlenbergianum (Sw) Grout

Weissia Hedw controversa Hedw var controversa

Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum

Trichostomum Bruch spirale Prout

19

40 Ephemeraceae Ephemerum Hampe

cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp

41 Funariaceae AphflllolThegma SuD

serratum (1 Hook et Wils ex Drumm) Sullo

Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe

Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica

42 Splachnaceae Splachnum Hedw

ampullaceum Hedw

43 Schistostegaceae Schistostega Mohr

pennata (Hedw) Web el Mohr

44 Bryaceae Pohlia Hedw

annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr

Leptobryum (B S G) Wits pyriforme (Hedw) Wils

Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum

20

40 Ephemeraceae Ephemerum Hampe

cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp

41 Funariaceae AphflllolThegma SuD

serratum (1 Hook et Wils ex Drumm) Sullo

Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe

Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica

42 Splachnaceae Splachnum Hedw

ampullaceum Hedw

43 Schistostegaceae Schistostega Mohr

pennata (Hedw) Web el Mohr

44 Bryaceae Pohlia Hedw

annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr

Leptobryum (B S G) Wits pyriforme (Hedw) Wils

Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum

20

pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng

Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr

45 Mniaceae Mnium Hedw

ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw

46 Aulacomruaceae Au1acomnium Schwaegr

androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre

47 Meesiaceae Amblyodon B S G

deillbotus (Hedw) B S G

Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw

48 Bartramiaceae Plagiopus Brid

oederiana (Sw) Umpr

Bartramia Hedw pomiformis Hedw var pomiformis

Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid

21

~

~

49 Timmiaceae Timmia Hedw

megapolitana Hedw var megapolitana

50 Orthotrichaceae Zygodon Hook et Tayl

viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm

Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp

Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv

Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid

Drummondill Hook ex Drumm prorepens (Hedw) Brltt

51 Fontinalaceae Fantinalis Hedw

antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll

missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S

DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G

52 Climaciaceae aimecium Web et Mohr ex Mohr

americllnum Brid dedroides (Hedw) Web et Mohr

22

S3 Hedwigiaceae Hedwigia P Beauv

ciliata (Hedw) P Beauv

54 Cryphaeaceae Forsstroemia lindh

ohioense (Sull) lindh

55 Leucodontaceae Leucodon Schwaegr

brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU

56 Neckeraceae Neckera Hedw

pennata Hedw var pennata

1fomalia (Brid) B S G trichomanoides (Hedw) B S G

57 Theliaceae Thelia Sull

asprellll Sullo hirtella (Hedw) Sullo lescurii SuU

Myurella B SG julacea (Schwaegr) B S G

sibrica (C MueU) Reim var sibrica

58 Fabroniaceae Fabronio Raddi

ciliaris (Brid) Bnd

Anacamptodon Brid splachnoides (Groet ex Brid) Brid

Schwetschkeopsis Broth fabronia (Schwaegr) Broth

59 Leskeaceae Lindbergia Kindb

brachyptera (Mitt) Kindb

Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw

Leskeella (limpr) Loeske nervosa (Brid) Loeske

23

Pterlgy1lllndrum Hedw filiforme Hedw

60 Thuidiaceae Haplohymenfum Dozy et MoIk

trlste (Ces ex De Not) Kiodb

Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl

Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull

Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust

Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum

Steere et Anderson paludomm (Sull) AuS

61 Amblystegiaceae Cratoneuron (Sull) Spruce

commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum

Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum

Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth

Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile

24

fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn

Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium

Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum

Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst

Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt

Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb

Calliergonella Loeske cuspidata (Hedw) Loeske

Calliergidium (Ren) Grout bakeri (Ren) Grout

Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr

62 Brachytbeciaceae Tomenthypnum loeske

nitens (Hedw) Loeske var mtens

25

Homalotheciella (Card) Broth subcapillata (Hedw) Broth

Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum

Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout

Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb

Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske

EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich

Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum

63 Entodontaceae Entodon C Muell

brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix

Pleurozium Mitt schreberl (8rid) Mitt var schreberi

64 Plagiotheciaceae Pltlglothecium BSG

CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG

26

6S SernatophyUaceae Brotherella Loeske ex Fleisch

recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch

Sematophyllum Mitt demissum (Wils) Mitt

66 Hypnaceae Pl(itygyrium BSG

repens (Brid) BSG

PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson

Homomallium (Schimp) Loeske adnatum (Hedw) Broth

Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid

Calicladium Crum haldanianum (Grev) Crum

sopterygiopsis Iwats muelleriana (Schimp) Iwats

Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt

Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch

Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats

hilium De Not crista-castrenna (Hedw) De Not

27

67 Rhytidiaceae Rhytidium (Sull) Kindb

rugosum (Hedw) Kindb

Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus

68 Hylocomiaceae Hylocomium BSG

pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens

69 Buxbaumiaceae Buxbaumia Hedw

aphyllll Hedw

Diphyscium Mohr follosum (Hedw) Mohr

70 Tetraphidaceae Tetraphi3

pellucida Hedw

71 Polytrichaceae Atrichum PmiddotBeauv

angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum

Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv

Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid

28

MAPS OF BRYOPHYTE TAXA

REPORTED FOR WISCONSIN

HORNWORTS AND LIVERWORTS

29

bull bull WSHeV~N S vE II

0 ~

e~RRON ~USK

(J~ (j -

CHIPPE A DUNN

0 o0 poundAU CLAIRE

~ TREMshyPEAshyLEAU

MICE

ONEIDA

Q LINCOLN

TAYLOR

Q t-~----- MOIRA THON

CLARK

o

10

LA FAVETTE

Hornwor ts amp Liverworts

- None - 1 - 25 taxa

- 26 - 50 tRXshy

- 51 - 75 tax)

- 7c - 100 taxa

FONDOU j LAC ~

DODGE

i O JEfFERSON

WALWORTH

30

Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~

46 88 136 141 Evans 38 46 141

Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn

89 21 384687 88 89 90 (138) (145)

Anthoceros macounil MA Howe ~oceros punctatus L

21 (45) (144) 34

31

Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~

24 141 Loeske 28 39 46 89 90 141

Bazzania trilobata (1) S Gray

1 26 39 46 (57) (59) 79 88 89

90 119 141

BlePbarostoma trichoffiyllum (1) Dumort

1 8 26 39 46 89 90 137 141

Blasia pusilla 1

1 39 46 90

Calypogeja fissa (1) Raddi

90

32

$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull

88 89 89

CallpoundO~eja neesiana (Mass amp Carest) K Mull

39 46 89 140 (141)

~pogeja sJlapound2icola (H Arnell tl J bull

Perss) Warnst et Loeske (119) 141

CeIalozia biscuspidata (L) Dum

1 37 I 46 90 t 137 141

CalYpogej~ trichomanis (L) Corda

26 39 (119) 137

33

CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb

39 46 79 141 39 4679 88 (89) 90 119 141

Cephalozia lacinulata Jack ~ Spruce

141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141

Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141

34

49 Timmiaceae Timmia Hedw

megapolitana Hedw var megapolitana

50 Orthotrichaceae Zygodon Hook et Tayl

viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm

Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp

Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv

Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid

Drummondill Hook ex Drumm prorepens (Hedw) Brltt

51 Fontinalaceae Fantinalis Hedw

antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll

missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S

DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G

52 Climaciaceae aimecium Web et Mohr ex Mohr

americllnum Brid dedroides (Hedw) Web et Mohr

22

S3 Hedwigiaceae Hedwigia P Beauv

ciliata (Hedw) P Beauv

54 Cryphaeaceae Forsstroemia lindh

ohioense (Sull) lindh

55 Leucodontaceae Leucodon Schwaegr

brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU

56 Neckeraceae Neckera Hedw

pennata Hedw var pennata

1fomalia (Brid) B S G trichomanoides (Hedw) B S G

57 Theliaceae Thelia Sull

asprellll Sullo hirtella (Hedw) Sullo lescurii SuU

Myurella B SG julacea (Schwaegr) B S G

sibrica (C MueU) Reim var sibrica

58 Fabroniaceae Fabronio Raddi

ciliaris (Brid) Bnd

Anacamptodon Brid splachnoides (Groet ex Brid) Brid

Schwetschkeopsis Broth fabronia (Schwaegr) Broth

59 Leskeaceae Lindbergia Kindb

brachyptera (Mitt) Kindb

Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw

Leskeella (limpr) Loeske nervosa (Brid) Loeske

23

Pterlgy1lllndrum Hedw filiforme Hedw

60 Thuidiaceae Haplohymenfum Dozy et MoIk

trlste (Ces ex De Not) Kiodb

Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl

Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull

Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust

Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum

Steere et Anderson paludomm (Sull) AuS

61 Amblystegiaceae Cratoneuron (Sull) Spruce

commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum

Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum

Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth

Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile

24

fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn

Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium

Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum

Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst

Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt

Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb

Calliergonella Loeske cuspidata (Hedw) Loeske

Calliergidium (Ren) Grout bakeri (Ren) Grout

Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr

62 Brachytbeciaceae Tomenthypnum loeske

nitens (Hedw) Loeske var mtens

25

Homalotheciella (Card) Broth subcapillata (Hedw) Broth

Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum

Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout

Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb

Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske

EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich

Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum

63 Entodontaceae Entodon C Muell

brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix

Pleurozium Mitt schreberl (8rid) Mitt var schreberi

64 Plagiotheciaceae Pltlglothecium BSG

CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG

26

6S SernatophyUaceae Brotherella Loeske ex Fleisch

recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch

Sematophyllum Mitt demissum (Wils) Mitt

66 Hypnaceae Pl(itygyrium BSG

repens (Brid) BSG

PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson

Homomallium (Schimp) Loeske adnatum (Hedw) Broth

Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid

Calicladium Crum haldanianum (Grev) Crum

sopterygiopsis Iwats muelleriana (Schimp) Iwats

Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt

Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch

Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats

hilium De Not crista-castrenna (Hedw) De Not

27

67 Rhytidiaceae Rhytidium (Sull) Kindb

rugosum (Hedw) Kindb

Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus

68 Hylocomiaceae Hylocomium BSG

pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens

69 Buxbaumiaceae Buxbaumia Hedw

aphyllll Hedw

Diphyscium Mohr follosum (Hedw) Mohr

70 Tetraphidaceae Tetraphi3

pellucida Hedw

71 Polytrichaceae Atrichum PmiddotBeauv

angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum

Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv

Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid

28

MAPS OF BRYOPHYTE TAXA

REPORTED FOR WISCONSIN

HORNWORTS AND LIVERWORTS

29

bull bull WSHeV~N S vE II

0 ~

e~RRON ~USK

(J~ (j -

CHIPPE A DUNN

0 o0 poundAU CLAIRE

~ TREMshyPEAshyLEAU

MICE

ONEIDA

Q LINCOLN

TAYLOR

Q t-~----- MOIRA THON

CLARK

o

10

LA FAVETTE

Hornwor ts amp Liverworts

- None - 1 - 25 taxa

- 26 - 50 tRXshy

- 51 - 75 tax)

- 7c - 100 taxa

FONDOU j LAC ~

DODGE

i O JEfFERSON

WALWORTH

30

Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~

46 88 136 141 Evans 38 46 141

Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn

89 21 384687 88 89 90 (138) (145)

Anthoceros macounil MA Howe ~oceros punctatus L

21 (45) (144) 34

31

Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~

24 141 Loeske 28 39 46 89 90 141

Bazzania trilobata (1) S Gray

1 26 39 46 (57) (59) 79 88 89

90 119 141

BlePbarostoma trichoffiyllum (1) Dumort

1 8 26 39 46 89 90 137 141

Blasia pusilla 1

1 39 46 90

Calypogeja fissa (1) Raddi

90

32

$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull

88 89 89

CallpoundO~eja neesiana (Mass amp Carest) K Mull

39 46 89 140 (141)

~pogeja sJlapound2icola (H Arnell tl J bull

Perss) Warnst et Loeske (119) 141

CeIalozia biscuspidata (L) Dum

1 37 I 46 90 t 137 141

CalYpogej~ trichomanis (L) Corda

26 39 (119) 137

33

CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb

39 46 79 141 39 4679 88 (89) 90 119 141

Cephalozia lacinulata Jack ~ Spruce

141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141

Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141

34

S3 Hedwigiaceae Hedwigia P Beauv

ciliata (Hedw) P Beauv

54 Cryphaeaceae Forsstroemia lindh

ohioense (Sull) lindh

55 Leucodontaceae Leucodon Schwaegr

brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU

56 Neckeraceae Neckera Hedw

pennata Hedw var pennata

1fomalia (Brid) B S G trichomanoides (Hedw) B S G

57 Theliaceae Thelia Sull

asprellll Sullo hirtella (Hedw) Sullo lescurii SuU

Myurella B SG julacea (Schwaegr) B S G

sibrica (C MueU) Reim var sibrica

58 Fabroniaceae Fabronio Raddi

ciliaris (Brid) Bnd

Anacamptodon Brid splachnoides (Groet ex Brid) Brid

Schwetschkeopsis Broth fabronia (Schwaegr) Broth

59 Leskeaceae Lindbergia Kindb

brachyptera (Mitt) Kindb

Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw

Leskeella (limpr) Loeske nervosa (Brid) Loeske

23

Pterlgy1lllndrum Hedw filiforme Hedw

60 Thuidiaceae Haplohymenfum Dozy et MoIk

trlste (Ces ex De Not) Kiodb

Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl

Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull

Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust

Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum

Steere et Anderson paludomm (Sull) AuS

61 Amblystegiaceae Cratoneuron (Sull) Spruce

commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum

Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum

Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth

Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile

24

fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn

Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium

Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum

Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst

Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt

Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb

Calliergonella Loeske cuspidata (Hedw) Loeske

Calliergidium (Ren) Grout bakeri (Ren) Grout

Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr

62 Brachytbeciaceae Tomenthypnum loeske

nitens (Hedw) Loeske var mtens

25

Homalotheciella (Card) Broth subcapillata (Hedw) Broth

Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum

Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout

Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb

Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske

EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich

Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum

63 Entodontaceae Entodon C Muell

brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix

Pleurozium Mitt schreberl (8rid) Mitt var schreberi

64 Plagiotheciaceae Pltlglothecium BSG

CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG

26

6S SernatophyUaceae Brotherella Loeske ex Fleisch

recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch

Sematophyllum Mitt demissum (Wils) Mitt

66 Hypnaceae Pl(itygyrium BSG

repens (Brid) BSG

PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson

Homomallium (Schimp) Loeske adnatum (Hedw) Broth

Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid

Calicladium Crum haldanianum (Grev) Crum

sopterygiopsis Iwats muelleriana (Schimp) Iwats

Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt

Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch

Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats

hilium De Not crista-castrenna (Hedw) De Not

27

67 Rhytidiaceae Rhytidium (Sull) Kindb

rugosum (Hedw) Kindb

Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus

68 Hylocomiaceae Hylocomium BSG

pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens

69 Buxbaumiaceae Buxbaumia Hedw

aphyllll Hedw

Diphyscium Mohr follosum (Hedw) Mohr

70 Tetraphidaceae Tetraphi3

pellucida Hedw

71 Polytrichaceae Atrichum PmiddotBeauv

angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum

Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv

Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid

28

MAPS OF BRYOPHYTE TAXA

REPORTED FOR WISCONSIN

HORNWORTS AND LIVERWORTS

29

bull bull WSHeV~N S vE II

0 ~

e~RRON ~USK

(J~ (j -

CHIPPE A DUNN

0 o0 poundAU CLAIRE

~ TREMshyPEAshyLEAU

MICE

ONEIDA

Q LINCOLN

TAYLOR

Q t-~----- MOIRA THON

CLARK

o

10

LA FAVETTE

Hornwor ts amp Liverworts

- None - 1 - 25 taxa

- 26 - 50 tRXshy

- 51 - 75 tax)

- 7c - 100 taxa

FONDOU j LAC ~

DODGE

i O JEfFERSON

WALWORTH

30

Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~

46 88 136 141 Evans 38 46 141

Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn

89 21 384687 88 89 90 (138) (145)

Anthoceros macounil MA Howe ~oceros punctatus L

21 (45) (144) 34

31

Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~

24 141 Loeske 28 39 46 89 90 141

Bazzania trilobata (1) S Gray

1 26 39 46 (57) (59) 79 88 89

90 119 141

BlePbarostoma trichoffiyllum (1) Dumort

1 8 26 39 46 89 90 137 141

Blasia pusilla 1

1 39 46 90

Calypogeja fissa (1) Raddi

90

32

$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull

88 89 89

CallpoundO~eja neesiana (Mass amp Carest) K Mull

39 46 89 140 (141)

~pogeja sJlapound2icola (H Arnell tl J bull

Perss) Warnst et Loeske (119) 141

CeIalozia biscuspidata (L) Dum

1 37 I 46 90 t 137 141

CalYpogej~ trichomanis (L) Corda

26 39 (119) 137

33

CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb

39 46 79 141 39 4679 88 (89) 90 119 141

Cephalozia lacinulata Jack ~ Spruce

141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141

Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141

34

Pterlgy1lllndrum Hedw filiforme Hedw

60 Thuidiaceae Haplohymenfum Dozy et MoIk

trlste (Ces ex De Not) Kiodb

Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl

Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull

Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust

Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum

Steere et Anderson paludomm (Sull) AuS

61 Amblystegiaceae Cratoneuron (Sull) Spruce

commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum

Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum

Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth

Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile

24

fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn

Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium

Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum

Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst

Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt

Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb

Calliergonella Loeske cuspidata (Hedw) Loeske

Calliergidium (Ren) Grout bakeri (Ren) Grout

Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr

62 Brachytbeciaceae Tomenthypnum loeske

nitens (Hedw) Loeske var mtens

25

Homalotheciella (Card) Broth subcapillata (Hedw) Broth

Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum

Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout

Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb

Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske

EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich

Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum

63 Entodontaceae Entodon C Muell

brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix

Pleurozium Mitt schreberl (8rid) Mitt var schreberi

64 Plagiotheciaceae Pltlglothecium BSG

CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG

26

6S SernatophyUaceae Brotherella Loeske ex Fleisch

recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch

Sematophyllum Mitt demissum (Wils) Mitt

66 Hypnaceae Pl(itygyrium BSG

repens (Brid) BSG

PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson

Homomallium (Schimp) Loeske adnatum (Hedw) Broth

Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid

Calicladium Crum haldanianum (Grev) Crum

sopterygiopsis Iwats muelleriana (Schimp) Iwats

Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt

Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch

Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats

hilium De Not crista-castrenna (Hedw) De Not

27

67 Rhytidiaceae Rhytidium (Sull) Kindb

rugosum (Hedw) Kindb

Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus

68 Hylocomiaceae Hylocomium BSG

pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens

69 Buxbaumiaceae Buxbaumia Hedw

aphyllll Hedw

Diphyscium Mohr follosum (Hedw) Mohr

70 Tetraphidaceae Tetraphi3

pellucida Hedw

71 Polytrichaceae Atrichum PmiddotBeauv

angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum

Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv

Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid

28

MAPS OF BRYOPHYTE TAXA

REPORTED FOR WISCONSIN

HORNWORTS AND LIVERWORTS

29

bull bull WSHeV~N S vE II

0 ~

e~RRON ~USK

(J~ (j -

CHIPPE A DUNN

0 o0 poundAU CLAIRE

~ TREMshyPEAshyLEAU

MICE

ONEIDA

Q LINCOLN

TAYLOR

Q t-~----- MOIRA THON

CLARK

o

10

LA FAVETTE

Hornwor ts amp Liverworts

- None - 1 - 25 taxa

- 26 - 50 tRXshy

- 51 - 75 tax)

- 7c - 100 taxa

FONDOU j LAC ~

DODGE

i O JEfFERSON

WALWORTH

30

Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~

46 88 136 141 Evans 38 46 141

Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn

89 21 384687 88 89 90 (138) (145)

Anthoceros macounil MA Howe ~oceros punctatus L

21 (45) (144) 34

31

Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~

24 141 Loeske 28 39 46 89 90 141

Bazzania trilobata (1) S Gray

1 26 39 46 (57) (59) 79 88 89

90 119 141

BlePbarostoma trichoffiyllum (1) Dumort

1 8 26 39 46 89 90 137 141

Blasia pusilla 1

1 39 46 90

Calypogeja fissa (1) Raddi

90

32

$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull

88 89 89

CallpoundO~eja neesiana (Mass amp Carest) K Mull

39 46 89 140 (141)

~pogeja sJlapound2icola (H Arnell tl J bull

Perss) Warnst et Loeske (119) 141

CeIalozia biscuspidata (L) Dum

1 37 I 46 90 t 137 141

CalYpogej~ trichomanis (L) Corda

26 39 (119) 137

33

CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb

39 46 79 141 39 4679 88 (89) 90 119 141

Cephalozia lacinulata Jack ~ Spruce

141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141

Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141

34

fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn

Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium

Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum

Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst

Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt

Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb

Calliergonella Loeske cuspidata (Hedw) Loeske

Calliergidium (Ren) Grout bakeri (Ren) Grout

Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr

62 Brachytbeciaceae Tomenthypnum loeske

nitens (Hedw) Loeske var mtens

25

Homalotheciella (Card) Broth subcapillata (Hedw) Broth

Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum

Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout

Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb

Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske

EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich

Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum

63 Entodontaceae Entodon C Muell

brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix

Pleurozium Mitt schreberl (8rid) Mitt var schreberi

64 Plagiotheciaceae Pltlglothecium BSG

CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG

26

6S SernatophyUaceae Brotherella Loeske ex Fleisch

recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch

Sematophyllum Mitt demissum (Wils) Mitt

66 Hypnaceae Pl(itygyrium BSG

repens (Brid) BSG

PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson

Homomallium (Schimp) Loeske adnatum (Hedw) Broth

Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid

Calicladium Crum haldanianum (Grev) Crum

sopterygiopsis Iwats muelleriana (Schimp) Iwats

Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt

Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch

Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats

hilium De Not crista-castrenna (Hedw) De Not

27

67 Rhytidiaceae Rhytidium (Sull) Kindb

rugosum (Hedw) Kindb

Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus

68 Hylocomiaceae Hylocomium BSG

pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens

69 Buxbaumiaceae Buxbaumia Hedw

aphyllll Hedw

Diphyscium Mohr follosum (Hedw) Mohr

70 Tetraphidaceae Tetraphi3

pellucida Hedw

71 Polytrichaceae Atrichum PmiddotBeauv

angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum

Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv

Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid

28

MAPS OF BRYOPHYTE TAXA

REPORTED FOR WISCONSIN

HORNWORTS AND LIVERWORTS

29

bull bull WSHeV~N S vE II

0 ~

e~RRON ~USK

(J~ (j -

CHIPPE A DUNN

0 o0 poundAU CLAIRE

~ TREMshyPEAshyLEAU

MICE

ONEIDA

Q LINCOLN

TAYLOR

Q t-~----- MOIRA THON

CLARK

o

10

LA FAVETTE

Hornwor ts amp Liverworts

- None - 1 - 25 taxa

- 26 - 50 tRXshy

- 51 - 75 tax)

- 7c - 100 taxa

FONDOU j LAC ~

DODGE

i O JEfFERSON

WALWORTH

30

Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~

46 88 136 141 Evans 38 46 141

Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn

89 21 384687 88 89 90 (138) (145)

Anthoceros macounil MA Howe ~oceros punctatus L

21 (45) (144) 34

31

Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~

24 141 Loeske 28 39 46 89 90 141

Bazzania trilobata (1) S Gray

1 26 39 46 (57) (59) 79 88 89

90 119 141

BlePbarostoma trichoffiyllum (1) Dumort

1 8 26 39 46 89 90 137 141

Blasia pusilla 1

1 39 46 90

Calypogeja fissa (1) Raddi

90

32

$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull

88 89 89

CallpoundO~eja neesiana (Mass amp Carest) K Mull

39 46 89 140 (141)

~pogeja sJlapound2icola (H Arnell tl J bull

Perss) Warnst et Loeske (119) 141

CeIalozia biscuspidata (L) Dum

1 37 I 46 90 t 137 141

CalYpogej~ trichomanis (L) Corda

26 39 (119) 137

33

CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb

39 46 79 141 39 4679 88 (89) 90 119 141

Cephalozia lacinulata Jack ~ Spruce

141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141

Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141

34

Homalotheciella (Card) Broth subcapillata (Hedw) Broth

Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum

Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout

Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb

Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske

EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich

Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum

63 Entodontaceae Entodon C Muell

brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix

Pleurozium Mitt schreberl (8rid) Mitt var schreberi

64 Plagiotheciaceae Pltlglothecium BSG

CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG

26

6S SernatophyUaceae Brotherella Loeske ex Fleisch

recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch

Sematophyllum Mitt demissum (Wils) Mitt

66 Hypnaceae Pl(itygyrium BSG

repens (Brid) BSG

PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson

Homomallium (Schimp) Loeske adnatum (Hedw) Broth

Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid

Calicladium Crum haldanianum (Grev) Crum

sopterygiopsis Iwats muelleriana (Schimp) Iwats

Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt

Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch

Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats

hilium De Not crista-castrenna (Hedw) De Not

27

67 Rhytidiaceae Rhytidium (Sull) Kindb

rugosum (Hedw) Kindb

Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus

68 Hylocomiaceae Hylocomium BSG

pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens

69 Buxbaumiaceae Buxbaumia Hedw

aphyllll Hedw

Diphyscium Mohr follosum (Hedw) Mohr

70 Tetraphidaceae Tetraphi3

pellucida Hedw

71 Polytrichaceae Atrichum PmiddotBeauv

angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum

Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv

Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid

28

MAPS OF BRYOPHYTE TAXA

REPORTED FOR WISCONSIN

HORNWORTS AND LIVERWORTS

29

bull bull WSHeV~N S vE II

0 ~

e~RRON ~USK

(J~ (j -

CHIPPE A DUNN

0 o0 poundAU CLAIRE

~ TREMshyPEAshyLEAU

MICE

ONEIDA

Q LINCOLN

TAYLOR

Q t-~----- MOIRA THON

CLARK

o

10

LA FAVETTE

Hornwor ts amp Liverworts

- None - 1 - 25 taxa

- 26 - 50 tRXshy

- 51 - 75 tax)

- 7c - 100 taxa

FONDOU j LAC ~

DODGE

i O JEfFERSON

WALWORTH

30

Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~

46 88 136 141 Evans 38 46 141

Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn

89 21 384687 88 89 90 (138) (145)

Anthoceros macounil MA Howe ~oceros punctatus L

21 (45) (144) 34

31

Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~

24 141 Loeske 28 39 46 89 90 141

Bazzania trilobata (1) S Gray

1 26 39 46 (57) (59) 79 88 89

90 119 141

BlePbarostoma trichoffiyllum (1) Dumort

1 8 26 39 46 89 90 137 141

Blasia pusilla 1

1 39 46 90

Calypogeja fissa (1) Raddi

90

32

$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull

88 89 89

CallpoundO~eja neesiana (Mass amp Carest) K Mull

39 46 89 140 (141)

~pogeja sJlapound2icola (H Arnell tl J bull

Perss) Warnst et Loeske (119) 141

CeIalozia biscuspidata (L) Dum

1 37 I 46 90 t 137 141

CalYpogej~ trichomanis (L) Corda

26 39 (119) 137

33

CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb

39 46 79 141 39 4679 88 (89) 90 119 141

Cephalozia lacinulata Jack ~ Spruce

141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141

Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141

34

6S SernatophyUaceae Brotherella Loeske ex Fleisch

recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch

Sematophyllum Mitt demissum (Wils) Mitt

66 Hypnaceae Pl(itygyrium BSG

repens (Brid) BSG

PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson

Homomallium (Schimp) Loeske adnatum (Hedw) Broth

Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid

Calicladium Crum haldanianum (Grev) Crum

sopterygiopsis Iwats muelleriana (Schimp) Iwats

Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt

Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch

Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats

hilium De Not crista-castrenna (Hedw) De Not

27

67 Rhytidiaceae Rhytidium (Sull) Kindb

rugosum (Hedw) Kindb

Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus

68 Hylocomiaceae Hylocomium BSG

pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens

69 Buxbaumiaceae Buxbaumia Hedw

aphyllll Hedw

Diphyscium Mohr follosum (Hedw) Mohr

70 Tetraphidaceae Tetraphi3

pellucida Hedw

71 Polytrichaceae Atrichum PmiddotBeauv

angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum

Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv

Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid

28

MAPS OF BRYOPHYTE TAXA

REPORTED FOR WISCONSIN

HORNWORTS AND LIVERWORTS

29

bull bull WSHeV~N S vE II

0 ~

e~RRON ~USK

(J~ (j -

CHIPPE A DUNN

0 o0 poundAU CLAIRE

~ TREMshyPEAshyLEAU

MICE

ONEIDA

Q LINCOLN

TAYLOR

Q t-~----- MOIRA THON

CLARK

o

10

LA FAVETTE

Hornwor ts amp Liverworts

- None - 1 - 25 taxa

- 26 - 50 tRXshy

- 51 - 75 tax)

- 7c - 100 taxa

FONDOU j LAC ~

DODGE

i O JEfFERSON

WALWORTH

30

Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~

46 88 136 141 Evans 38 46 141

Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn

89 21 384687 88 89 90 (138) (145)

Anthoceros macounil MA Howe ~oceros punctatus L

21 (45) (144) 34

31

Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~

24 141 Loeske 28 39 46 89 90 141

Bazzania trilobata (1) S Gray

1 26 39 46 (57) (59) 79 88 89

90 119 141

BlePbarostoma trichoffiyllum (1) Dumort

1 8 26 39 46 89 90 137 141

Blasia pusilla 1

1 39 46 90

Calypogeja fissa (1) Raddi

90

32

$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull

88 89 89

CallpoundO~eja neesiana (Mass amp Carest) K Mull

39 46 89 140 (141)

~pogeja sJlapound2icola (H Arnell tl J bull

Perss) Warnst et Loeske (119) 141

CeIalozia biscuspidata (L) Dum

1 37 I 46 90 t 137 141

CalYpogej~ trichomanis (L) Corda

26 39 (119) 137

33

CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb

39 46 79 141 39 4679 88 (89) 90 119 141

Cephalozia lacinulata Jack ~ Spruce

141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141

Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141

34

67 Rhytidiaceae Rhytidium (Sull) Kindb

rugosum (Hedw) Kindb

Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus

68 Hylocomiaceae Hylocomium BSG

pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens

69 Buxbaumiaceae Buxbaumia Hedw

aphyllll Hedw

Diphyscium Mohr follosum (Hedw) Mohr

70 Tetraphidaceae Tetraphi3

pellucida Hedw

71 Polytrichaceae Atrichum PmiddotBeauv

angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum

Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv

Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid

28

MAPS OF BRYOPHYTE TAXA

REPORTED FOR WISCONSIN

HORNWORTS AND LIVERWORTS

29

bull bull WSHeV~N S vE II

0 ~

e~RRON ~USK

(J~ (j -

CHIPPE A DUNN

0 o0 poundAU CLAIRE

~ TREMshyPEAshyLEAU

MICE

ONEIDA

Q LINCOLN

TAYLOR

Q t-~----- MOIRA THON

CLARK

o

10

LA FAVETTE

Hornwor ts amp Liverworts

- None - 1 - 25 taxa

- 26 - 50 tRXshy

- 51 - 75 tax)

- 7c - 100 taxa

FONDOU j LAC ~

DODGE

i O JEfFERSON

WALWORTH

30

Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~

46 88 136 141 Evans 38 46 141

Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn

89 21 384687 88 89 90 (138) (145)

Anthoceros macounil MA Howe ~oceros punctatus L

21 (45) (144) 34

31

Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~

24 141 Loeske 28 39 46 89 90 141

Bazzania trilobata (1) S Gray

1 26 39 46 (57) (59) 79 88 89

90 119 141

BlePbarostoma trichoffiyllum (1) Dumort

1 8 26 39 46 89 90 137 141

Blasia pusilla 1

1 39 46 90

Calypogeja fissa (1) Raddi

90

32

$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull

88 89 89

CallpoundO~eja neesiana (Mass amp Carest) K Mull

39 46 89 140 (141)

~pogeja sJlapound2icola (H Arnell tl J bull

Perss) Warnst et Loeske (119) 141

CeIalozia biscuspidata (L) Dum

1 37 I 46 90 t 137 141

CalYpogej~ trichomanis (L) Corda

26 39 (119) 137

33

CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb

39 46 79 141 39 4679 88 (89) 90 119 141

Cephalozia lacinulata Jack ~ Spruce

141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141

Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141

34

MAPS OF BRYOPHYTE TAXA

REPORTED FOR WISCONSIN

HORNWORTS AND LIVERWORTS

29

bull bull WSHeV~N S vE II

0 ~

e~RRON ~USK

(J~ (j -

CHIPPE A DUNN

0 o0 poundAU CLAIRE

~ TREMshyPEAshyLEAU

MICE

ONEIDA

Q LINCOLN

TAYLOR

Q t-~----- MOIRA THON

CLARK

o

10

LA FAVETTE

Hornwor ts amp Liverworts

- None - 1 - 25 taxa

- 26 - 50 tRXshy

- 51 - 75 tax)

- 7c - 100 taxa

FONDOU j LAC ~

DODGE

i O JEfFERSON

WALWORTH

30

Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~

46 88 136 141 Evans 38 46 141

Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn

89 21 384687 88 89 90 (138) (145)

Anthoceros macounil MA Howe ~oceros punctatus L

21 (45) (144) 34

31

Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~

24 141 Loeske 28 39 46 89 90 141

Bazzania trilobata (1) S Gray

1 26 39 46 (57) (59) 79 88 89

90 119 141

BlePbarostoma trichoffiyllum (1) Dumort

1 8 26 39 46 89 90 137 141

Blasia pusilla 1

1 39 46 90

Calypogeja fissa (1) Raddi

90

32

$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull

88 89 89

CallpoundO~eja neesiana (Mass amp Carest) K Mull

39 46 89 140 (141)

~pogeja sJlapound2icola (H Arnell tl J bull

Perss) Warnst et Loeske (119) 141

CeIalozia biscuspidata (L) Dum

1 37 I 46 90 t 137 141

CalYpogej~ trichomanis (L) Corda

26 39 (119) 137

33

CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb

39 46 79 141 39 4679 88 (89) 90 119 141

Cephalozia lacinulata Jack ~ Spruce

141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141

Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141

34

bull bull WSHeV~N S vE II

0 ~

e~RRON ~USK

(J~ (j -

CHIPPE A DUNN

0 o0 poundAU CLAIRE

~ TREMshyPEAshyLEAU

MICE

ONEIDA

Q LINCOLN

TAYLOR

Q t-~----- MOIRA THON

CLARK

o

10

LA FAVETTE

Hornwor ts amp Liverworts

- None - 1 - 25 taxa

- 26 - 50 tRXshy

- 51 - 75 tax)

- 7c - 100 taxa

FONDOU j LAC ~

DODGE

i O JEfFERSON

WALWORTH

30

Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~

46 88 136 141 Evans 38 46 141

Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn

89 21 384687 88 89 90 (138) (145)

Anthoceros macounil MA Howe ~oceros punctatus L

21 (45) (144) 34

31

Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~

24 141 Loeske 28 39 46 89 90 141

Bazzania trilobata (1) S Gray

1 26 39 46 (57) (59) 79 88 89

90 119 141

BlePbarostoma trichoffiyllum (1) Dumort

1 8 26 39 46 89 90 137 141

Blasia pusilla 1

1 39 46 90

Calypogeja fissa (1) Raddi

90

32

$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull

88 89 89

CallpoundO~eja neesiana (Mass amp Carest) K Mull

39 46 89 140 (141)

~pogeja sJlapound2icola (H Arnell tl J bull

Perss) Warnst et Loeske (119) 141

CeIalozia biscuspidata (L) Dum

1 37 I 46 90 t 137 141

CalYpogej~ trichomanis (L) Corda

26 39 (119) 137

33

CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb

39 46 79 141 39 4679 88 (89) 90 119 141

Cephalozia lacinulata Jack ~ Spruce

141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141

Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141

34

Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~

46 88 136 141 Evans 38 46 141

Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn

89 21 384687 88 89 90 (138) (145)

Anthoceros macounil MA Howe ~oceros punctatus L

21 (45) (144) 34

31

Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~

24 141 Loeske 28 39 46 89 90 141

Bazzania trilobata (1) S Gray

1 26 39 46 (57) (59) 79 88 89

90 119 141

BlePbarostoma trichoffiyllum (1) Dumort

1 8 26 39 46 89 90 137 141

Blasia pusilla 1

1 39 46 90

Calypogeja fissa (1) Raddi

90

32

$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull

88 89 89

CallpoundO~eja neesiana (Mass amp Carest) K Mull

39 46 89 140 (141)

~pogeja sJlapound2icola (H Arnell tl J bull

Perss) Warnst et Loeske (119) 141

CeIalozia biscuspidata (L) Dum

1 37 I 46 90 t 137 141

CalYpogej~ trichomanis (L) Corda

26 39 (119) 137

33

CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb

39 46 79 141 39 4679 88 (89) 90 119 141

Cephalozia lacinulata Jack ~ Spruce

141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141

Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141

34

Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~

24 141 Loeske 28 39 46 89 90 141

Bazzania trilobata (1) S Gray

1 26 39 46 (57) (59) 79 88 89

90 119 141

BlePbarostoma trichoffiyllum (1) Dumort

1 8 26 39 46 89 90 137 141

Blasia pusilla 1

1 39 46 90

Calypogeja fissa (1) Raddi

90

32

$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull

88 89 89

CallpoundO~eja neesiana (Mass amp Carest) K Mull

39 46 89 140 (141)

~pogeja sJlapound2icola (H Arnell tl J bull

Perss) Warnst et Loeske (119) 141

CeIalozia biscuspidata (L) Dum

1 37 I 46 90 t 137 141

CalYpogej~ trichomanis (L) Corda

26 39 (119) 137

33

CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb

39 46 79 141 39 4679 88 (89) 90 119 141

Cephalozia lacinulata Jack ~ Spruce

141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141

Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141

34

$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull

88 89 89

CallpoundO~eja neesiana (Mass amp Carest) K Mull

39 46 89 140 (141)

~pogeja sJlapound2icola (H Arnell tl J bull

Perss) Warnst et Loeske (119) 141

CeIalozia biscuspidata (L) Dum

1 37 I 46 90 t 137 141

CalYpogej~ trichomanis (L) Corda

26 39 (119) 137

33

CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb

39 46 79 141 39 4679 88 (89) 90 119 141

Cephalozia lacinulata Jack ~ Spruce

141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141

Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141

34

CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb

39 46 79 141 39 4679 88 (89) 90 119 141

Cephalozia lacinulata Jack ~ Spruce

141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141

Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141

34

Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn

46 90 46 (119) 141

Cephaloziella hampeana (Nees) Schiffn

46

Cephaloziella rubella (Nees) Warnst

46 39

Cephaloziella rubella var sullivantii

(Aust) K Mull 46 39

Cephaloziella rubella var bifida (Lindb) Douin

35

46

Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis

39J 46 88 89J 90 (Roth) K Milll 46

Chiloscyphus E9lyanthos (L) Corda

1 24 39 46

Chiloscyphus polyanthos vax rivularis

(Schrad) Nees 39 46

poundpoundnocepha1um conicum (1) Lindb

12 28 39 46 (49) 87 88 8990 120 121 148

Cololejeunea biddlecomiae (Aust) Evans

39 46 62 87 88 89 90

36

Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb

(35)41 46 90 140 1 t 38 (39) t 46

Frullania bolanderi Aust

13 46 62 88 90 139 140

Frullanla ari ttoniae Evans

39 46 69 (71) 87 88 89 90

Frullania eboracensis Gott

1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148

Frullania inflata Gott

46 87 88 89 (139)

Frullania oakesiana Aust Frullania plana Sullo

(35) 42 46 87 (139) (140) 92

Frullania ri paria Hampe ~ Lehm

46 69 71 87 89

Frullania ~elwynianna Pears

46 89

Geocalyx SEaveolens (Schrad) Nees

394688 8990 (119) 140

Frullania tamarisci (L) Dum subsp sPagrayana

(Mont) Hatt 28 34 l~6 t (57) 88 89 90

38

Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph

(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141

Jungermannia confertissima Nees

137 141

Jungermannia crenuliform1s Aust

79 137 141

Jungermannia ~il1ima Sm

46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~

Vana (35) 46 79 141

39

Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle

40 46 79 136 141 39 46 79 88 90 1)7 141

Jungermannia IXlaxis Lindb Jungermannia pumila With

46 67 79 (139) (140) 141 )9 46 90 1)6 141

Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle

4679136140IIH 136

Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum

1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88

lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees

1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136

Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust

46 90 137 141 136 140 141

41

loIilozia badensls (Gatt ~ Rabenh)

46 141

lophozia blcrenata (Schmid sectA Hoffm)

Dum 46 141

lophozia collaris (Nees) Dum

39 46 141

lophozla excisa (Dicks) Dum

27 4679 141

lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)

36 39 40 46 136 (140) 141 Evans 39 46 136 141

42

141

Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum

27 39 46 89 141 8 27 31 46 79 90 140 141

LoQhozia longidens Macoun Lophozia ventricosa

1 46 79 90 141

Lophozia ventricosa var silvicola

(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)

Macoun 39 46 79 141

43

~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark

46 (139) (140) 9 46 79 116 139

Mannia triandra (Scop) Gro 11e

46 (140)

Marchantia pound91ymorpha L

1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147

Mo~rckia hibernica (HookJ Gott

46 89

Metzgeria conjugata Lindb

44

46

Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo

39 46 141 (35)45 46

Nowellia curvifolia (Dicks) Mitt

17 27 3846 88 89 90 141

Odontoschisma denudatuJI1 (Nees ~ Mart)

Dum 46 81 137 141

Pallavicinia lyellii (Hook~ Carruth

90

Odontoschisma macounii (Aust) Under

45 46 (66) (139) 140 141

45

=

Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum

27 39 46 90 13724 39 46

Pellia neesiana Limp bull Phaeoceros

39 46 89 27 39 45 46 97

Porella pinnata L

13 27 I 46 90 Plagiochila asplenioides (L) Dum

27 I 39 1+6 88 89 I 90 137

46

~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb

17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77

Preissia quadrata (Seop) Nees

1 36 3946 (71) 88 89 90 121 123

137

Ptllidium ciliare (L) Hampe

1 27 39 46 132 141

Dum

(22) 28 3946 (57) 87 88 89 90 92

Ptilidium ~lcherrium (G Web) Hampe

(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141

47

Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis

39J 46 88 89J 90 (Roth) K Milll 46

Chiloscyphus E9lyanthos (L) Corda

1 24 39 46

Chiloscyphus polyanthos vax rivularis

(Schrad) Nees 39 46

poundpoundnocepha1um conicum (1) Lindb

12 28 39 46 (49) 87 88 8990 120 121 148

Cololejeunea biddlecomiae (Aust) Evans

39 46 62 87 88 89 90

36

Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb

(35)41 46 90 140 1 t 38 (39) t 46

Frullania bolanderi Aust

13 46 62 88 90 139 140

Frullanla ari ttoniae Evans

39 46 69 (71) 87 88 89 90

Frullania eboracensis Gott

1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148

Frullania inflata Gott

46 87 88 89 (139)

Frullania oakesiana Aust Frullania plana Sullo

(35) 42 46 87 (139) (140) 92

Frullania ri paria Hampe ~ Lehm

46 69 71 87 89

Frullania ~elwynianna Pears

46 89

Geocalyx SEaveolens (Schrad) Nees

394688 8990 (119) 140

Frullania tamarisci (L) Dum subsp sPagrayana

(Mont) Hatt 28 34 l~6 t (57) 88 89 90

38

Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph

(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141

Jungermannia confertissima Nees

137 141

Jungermannia crenuliform1s Aust

79 137 141

Jungermannia ~il1ima Sm

46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~

Vana (35) 46 79 141

39

Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle

40 46 79 136 141 39 46 79 88 90 1)7 141

Jungermannia IXlaxis Lindb Jungermannia pumila With

46 67 79 (139) (140) 141 )9 46 90 1)6 141

Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle

4679136140IIH 136

Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum

1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88

lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees

1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136

Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust

46 90 137 141 136 140 141

41

loIilozia badensls (Gatt ~ Rabenh)

46 141

lophozia blcrenata (Schmid sectA Hoffm)

Dum 46 141

lophozia collaris (Nees) Dum

39 46 141

lophozla excisa (Dicks) Dum

27 4679 141

lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)

36 39 40 46 136 (140) 141 Evans 39 46 136 141

42

141

Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum

27 39 46 89 141 8 27 31 46 79 90 140 141

LoQhozia longidens Macoun Lophozia ventricosa

1 46 79 90 141

Lophozia ventricosa var silvicola

(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)

Macoun 39 46 79 141

43

~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark

46 (139) (140) 9 46 79 116 139

Mannia triandra (Scop) Gro 11e

46 (140)

Marchantia pound91ymorpha L

1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147

Mo~rckia hibernica (HookJ Gott

46 89

Metzgeria conjugata Lindb

44

46

Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo

39 46 141 (35)45 46

Nowellia curvifolia (Dicks) Mitt

17 27 3846 88 89 90 141

Odontoschisma denudatuJI1 (Nees ~ Mart)

Dum 46 81 137 141

Pallavicinia lyellii (Hook~ Carruth

90

Odontoschisma macounii (Aust) Under

45 46 (66) (139) 140 141

45

=

Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum

27 39 46 90 13724 39 46

Pellia neesiana Limp bull Phaeoceros

39 46 89 27 39 45 46 97

Porella pinnata L

13 27 I 46 90 Plagiochila asplenioides (L) Dum

27 I 39 1+6 88 89 I 90 137

46

~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb

17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77

Preissia quadrata (Seop) Nees

1 36 3946 (71) 88 89 90 121 123

137

Ptllidium ciliare (L) Hampe

1 27 39 46 132 141

Dum

(22) 28 3946 (57) 87 88 89 90 92

Ptilidium ~lcherrium (G Web) Hampe

(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141

47

Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb

(35)41 46 90 140 1 t 38 (39) t 46

Frullania bolanderi Aust

13 46 62 88 90 139 140

Frullanla ari ttoniae Evans

39 46 69 (71) 87 88 89 90

Frullania eboracensis Gott

1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148

Frullania inflata Gott

46 87 88 89 (139)

Frullania oakesiana Aust Frullania plana Sullo

(35) 42 46 87 (139) (140) 92

Frullania ri paria Hampe ~ Lehm

46 69 71 87 89

Frullania ~elwynianna Pears

46 89

Geocalyx SEaveolens (Schrad) Nees

394688 8990 (119) 140

Frullania tamarisci (L) Dum subsp sPagrayana

(Mont) Hatt 28 34 l~6 t (57) 88 89 90

38

Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph

(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141

Jungermannia confertissima Nees

137 141

Jungermannia crenuliform1s Aust

79 137 141

Jungermannia ~il1ima Sm

46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~

Vana (35) 46 79 141

39

Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle

40 46 79 136 141 39 46 79 88 90 1)7 141

Jungermannia IXlaxis Lindb Jungermannia pumila With

46 67 79 (139) (140) 141 )9 46 90 1)6 141

Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle

4679136140IIH 136

Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum

1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88

lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees

1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136

Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust

46 90 137 141 136 140 141

41

loIilozia badensls (Gatt ~ Rabenh)

46 141

lophozia blcrenata (Schmid sectA Hoffm)

Dum 46 141

lophozia collaris (Nees) Dum

39 46 141

lophozla excisa (Dicks) Dum

27 4679 141

lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)

36 39 40 46 136 (140) 141 Evans 39 46 136 141

42

141

Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum

27 39 46 89 141 8 27 31 46 79 90 140 141

LoQhozia longidens Macoun Lophozia ventricosa

1 46 79 90 141

Lophozia ventricosa var silvicola

(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)

Macoun 39 46 79 141

43

~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark

46 (139) (140) 9 46 79 116 139

Mannia triandra (Scop) Gro 11e

46 (140)

Marchantia pound91ymorpha L

1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147

Mo~rckia hibernica (HookJ Gott

46 89

Metzgeria conjugata Lindb

44

46

Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo

39 46 141 (35)45 46

Nowellia curvifolia (Dicks) Mitt

17 27 3846 88 89 90 141

Odontoschisma denudatuJI1 (Nees ~ Mart)

Dum 46 81 137 141

Pallavicinia lyellii (Hook~ Carruth

90

Odontoschisma macounii (Aust) Under

45 46 (66) (139) 140 141

45

=

Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum

27 39 46 90 13724 39 46

Pellia neesiana Limp bull Phaeoceros

39 46 89 27 39 45 46 97

Porella pinnata L

13 27 I 46 90 Plagiochila asplenioides (L) Dum

27 I 39 1+6 88 89 I 90 137

46

~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb

17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77

Preissia quadrata (Seop) Nees

1 36 3946 (71) 88 89 90 121 123

137

Ptllidium ciliare (L) Hampe

1 27 39 46 132 141

Dum

(22) 28 3946 (57) 87 88 89 90 92

Ptilidium ~lcherrium (G Web) Hampe

(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141

47

Frullania oakesiana Aust Frullania plana Sullo

(35) 42 46 87 (139) (140) 92

Frullania ri paria Hampe ~ Lehm

46 69 71 87 89

Frullania ~elwynianna Pears

46 89

Geocalyx SEaveolens (Schrad) Nees

394688 8990 (119) 140

Frullania tamarisci (L) Dum subsp sPagrayana

(Mont) Hatt 28 34 l~6 t (57) 88 89 90

38

Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph

(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141

Jungermannia confertissima Nees

137 141

Jungermannia crenuliform1s Aust

79 137 141

Jungermannia ~il1ima Sm

46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~

Vana (35) 46 79 141

39

Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle

40 46 79 136 141 39 46 79 88 90 1)7 141

Jungermannia IXlaxis Lindb Jungermannia pumila With

46 67 79 (139) (140) 141 )9 46 90 1)6 141

Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle

4679136140IIH 136

Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum

1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88

lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees

1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136

Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust

46 90 137 141 136 140 141

41

loIilozia badensls (Gatt ~ Rabenh)

46 141

lophozia blcrenata (Schmid sectA Hoffm)

Dum 46 141

lophozia collaris (Nees) Dum

39 46 141

lophozla excisa (Dicks) Dum

27 4679 141

lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)

36 39 40 46 136 (140) 141 Evans 39 46 136 141

42

141

Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum

27 39 46 89 141 8 27 31 46 79 90 140 141

LoQhozia longidens Macoun Lophozia ventricosa

1 46 79 90 141

Lophozia ventricosa var silvicola

(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)

Macoun 39 46 79 141

43

~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark

46 (139) (140) 9 46 79 116 139

Mannia triandra (Scop) Gro 11e

46 (140)

Marchantia pound91ymorpha L

1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147

Mo~rckia hibernica (HookJ Gott

46 89

Metzgeria conjugata Lindb

44

46

Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo

39 46 141 (35)45 46

Nowellia curvifolia (Dicks) Mitt

17 27 3846 88 89 90 141

Odontoschisma denudatuJI1 (Nees ~ Mart)

Dum 46 81 137 141

Pallavicinia lyellii (Hook~ Carruth

90

Odontoschisma macounii (Aust) Under

45 46 (66) (139) 140 141

45

=

Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum

27 39 46 90 13724 39 46

Pellia neesiana Limp bull Phaeoceros

39 46 89 27 39 45 46 97

Porella pinnata L

13 27 I 46 90 Plagiochila asplenioides (L) Dum

27 I 39 1+6 88 89 I 90 137

46

~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb

17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77

Preissia quadrata (Seop) Nees

1 36 3946 (71) 88 89 90 121 123

137

Ptllidium ciliare (L) Hampe

1 27 39 46 132 141

Dum

(22) 28 3946 (57) 87 88 89 90 92

Ptilidium ~lcherrium (G Web) Hampe

(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141

47

Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph

(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141

Jungermannia confertissima Nees

137 141

Jungermannia crenuliform1s Aust

79 137 141

Jungermannia ~il1ima Sm

46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~

Vana (35) 46 79 141

39

Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle

40 46 79 136 141 39 46 79 88 90 1)7 141

Jungermannia IXlaxis Lindb Jungermannia pumila With

46 67 79 (139) (140) 141 )9 46 90 1)6 141

Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle

4679136140IIH 136

Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum

1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88

lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees

1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136

Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust

46 90 137 141 136 140 141

41

loIilozia badensls (Gatt ~ Rabenh)

46 141

lophozia blcrenata (Schmid sectA Hoffm)

Dum 46 141

lophozia collaris (Nees) Dum

39 46 141

lophozla excisa (Dicks) Dum

27 4679 141

lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)

36 39 40 46 136 (140) 141 Evans 39 46 136 141

42

141

Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum

27 39 46 89 141 8 27 31 46 79 90 140 141

LoQhozia longidens Macoun Lophozia ventricosa

1 46 79 90 141

Lophozia ventricosa var silvicola

(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)

Macoun 39 46 79 141

43

~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark

46 (139) (140) 9 46 79 116 139

Mannia triandra (Scop) Gro 11e

46 (140)

Marchantia pound91ymorpha L

1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147

Mo~rckia hibernica (HookJ Gott

46 89

Metzgeria conjugata Lindb

44

46

Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo

39 46 141 (35)45 46

Nowellia curvifolia (Dicks) Mitt

17 27 3846 88 89 90 141

Odontoschisma denudatuJI1 (Nees ~ Mart)

Dum 46 81 137 141

Pallavicinia lyellii (Hook~ Carruth

90

Odontoschisma macounii (Aust) Under

45 46 (66) (139) 140 141

45

=

Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum

27 39 46 90 13724 39 46

Pellia neesiana Limp bull Phaeoceros

39 46 89 27 39 45 46 97

Porella pinnata L

13 27 I 46 90 Plagiochila asplenioides (L) Dum

27 I 39 1+6 88 89 I 90 137

46

~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb

17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77

Preissia quadrata (Seop) Nees

1 36 3946 (71) 88 89 90 121 123

137

Ptllidium ciliare (L) Hampe

1 27 39 46 132 141

Dum

(22) 28 3946 (57) 87 88 89 90 92

Ptilidium ~lcherrium (G Web) Hampe

(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141

47

Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle

40 46 79 136 141 39 46 79 88 90 1)7 141

Jungermannia IXlaxis Lindb Jungermannia pumila With

46 67 79 (139) (140) 141 )9 46 90 1)6 141

Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle

4679136140IIH 136

Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum

1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88

lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees

1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136

Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust

46 90 137 141 136 140 141

41

loIilozia badensls (Gatt ~ Rabenh)

46 141

lophozia blcrenata (Schmid sectA Hoffm)

Dum 46 141

lophozia collaris (Nees) Dum

39 46 141

lophozla excisa (Dicks) Dum

27 4679 141

lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)

36 39 40 46 136 (140) 141 Evans 39 46 136 141

42

141

Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum

27 39 46 89 141 8 27 31 46 79 90 140 141

LoQhozia longidens Macoun Lophozia ventricosa

1 46 79 90 141

Lophozia ventricosa var silvicola

(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)

Macoun 39 46 79 141

43

~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark

46 (139) (140) 9 46 79 116 139

Mannia triandra (Scop) Gro 11e

46 (140)

Marchantia pound91ymorpha L

1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147

Mo~rckia hibernica (HookJ Gott

46 89

Metzgeria conjugata Lindb

44

46

Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo

39 46 141 (35)45 46

Nowellia curvifolia (Dicks) Mitt

17 27 3846 88 89 90 141

Odontoschisma denudatuJI1 (Nees ~ Mart)

Dum 46 81 137 141

Pallavicinia lyellii (Hook~ Carruth

90

Odontoschisma macounii (Aust) Under

45 46 (66) (139) 140 141

45

=

Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum

27 39 46 90 13724 39 46

Pellia neesiana Limp bull Phaeoceros

39 46 89 27 39 45 46 97

Porella pinnata L

13 27 I 46 90 Plagiochila asplenioides (L) Dum

27 I 39 1+6 88 89 I 90 137

46

~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb

17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77

Preissia quadrata (Seop) Nees

1 36 3946 (71) 88 89 90 121 123

137

Ptllidium ciliare (L) Hampe

1 27 39 46 132 141

Dum

(22) 28 3946 (57) 87 88 89 90 92

Ptilidium ~lcherrium (G Web) Hampe

(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141

47

Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum

1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88

lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees

1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136

Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust

46 90 137 141 136 140 141

41

loIilozia badensls (Gatt ~ Rabenh)

46 141

lophozia blcrenata (Schmid sectA Hoffm)

Dum 46 141

lophozia collaris (Nees) Dum

39 46 141

lophozla excisa (Dicks) Dum

27 4679 141

lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)

36 39 40 46 136 (140) 141 Evans 39 46 136 141

42

141

Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum

27 39 46 89 141 8 27 31 46 79 90 140 141

LoQhozia longidens Macoun Lophozia ventricosa

1 46 79 90 141

Lophozia ventricosa var silvicola

(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)

Macoun 39 46 79 141

43

~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark

46 (139) (140) 9 46 79 116 139

Mannia triandra (Scop) Gro 11e

46 (140)

Marchantia pound91ymorpha L

1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147

Mo~rckia hibernica (HookJ Gott

46 89

Metzgeria conjugata Lindb

44

46

Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo

39 46 141 (35)45 46

Nowellia curvifolia (Dicks) Mitt

17 27 3846 88 89 90 141

Odontoschisma denudatuJI1 (Nees ~ Mart)

Dum 46 81 137 141

Pallavicinia lyellii (Hook~ Carruth

90

Odontoschisma macounii (Aust) Under

45 46 (66) (139) 140 141

45

=

Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum

27 39 46 90 13724 39 46

Pellia neesiana Limp bull Phaeoceros

39 46 89 27 39 45 46 97

Porella pinnata L

13 27 I 46 90 Plagiochila asplenioides (L) Dum

27 I 39 1+6 88 89 I 90 137

46

~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb

17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77

Preissia quadrata (Seop) Nees

1 36 3946 (71) 88 89 90 121 123

137

Ptllidium ciliare (L) Hampe

1 27 39 46 132 141

Dum

(22) 28 3946 (57) 87 88 89 90 92

Ptilidium ~lcherrium (G Web) Hampe

(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141

47

loIilozia badensls (Gatt ~ Rabenh)

46 141

lophozia blcrenata (Schmid sectA Hoffm)

Dum 46 141

lophozia collaris (Nees) Dum

39 46 141

lophozla excisa (Dicks) Dum

27 4679 141

lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)

36 39 40 46 136 (140) 141 Evans 39 46 136 141

42

141

Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum

27 39 46 89 141 8 27 31 46 79 90 140 141

LoQhozia longidens Macoun Lophozia ventricosa

1 46 79 90 141

Lophozia ventricosa var silvicola

(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)

Macoun 39 46 79 141

43

~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark

46 (139) (140) 9 46 79 116 139

Mannia triandra (Scop) Gro 11e

46 (140)

Marchantia pound91ymorpha L

1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147

Mo~rckia hibernica (HookJ Gott

46 89

Metzgeria conjugata Lindb

44

46

Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo

39 46 141 (35)45 46

Nowellia curvifolia (Dicks) Mitt

17 27 3846 88 89 90 141

Odontoschisma denudatuJI1 (Nees ~ Mart)

Dum 46 81 137 141

Pallavicinia lyellii (Hook~ Carruth

90

Odontoschisma macounii (Aust) Under

45 46 (66) (139) 140 141

45

=

Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum

27 39 46 90 13724 39 46

Pellia neesiana Limp bull Phaeoceros

39 46 89 27 39 45 46 97

Porella pinnata L

13 27 I 46 90 Plagiochila asplenioides (L) Dum

27 I 39 1+6 88 89 I 90 137

46

~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb

17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77

Preissia quadrata (Seop) Nees

1 36 3946 (71) 88 89 90 121 123

137

Ptllidium ciliare (L) Hampe

1 27 39 46 132 141

Dum

(22) 28 3946 (57) 87 88 89 90 92

Ptilidium ~lcherrium (G Web) Hampe

(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141

47

141

Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum

27 39 46 89 141 8 27 31 46 79 90 140 141

LoQhozia longidens Macoun Lophozia ventricosa

1 46 79 90 141

Lophozia ventricosa var silvicola

(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)

Macoun 39 46 79 141

43

~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark

46 (139) (140) 9 46 79 116 139

Mannia triandra (Scop) Gro 11e

46 (140)

Marchantia pound91ymorpha L

1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147

Mo~rckia hibernica (HookJ Gott

46 89

Metzgeria conjugata Lindb

44

46

Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo

39 46 141 (35)45 46

Nowellia curvifolia (Dicks) Mitt

17 27 3846 88 89 90 141

Odontoschisma denudatuJI1 (Nees ~ Mart)

Dum 46 81 137 141

Pallavicinia lyellii (Hook~ Carruth

90

Odontoschisma macounii (Aust) Under

45 46 (66) (139) 140 141

45

=

Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum

27 39 46 90 13724 39 46

Pellia neesiana Limp bull Phaeoceros

39 46 89 27 39 45 46 97

Porella pinnata L

13 27 I 46 90 Plagiochila asplenioides (L) Dum

27 I 39 1+6 88 89 I 90 137

46

~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb

17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77

Preissia quadrata (Seop) Nees

1 36 3946 (71) 88 89 90 121 123

137

Ptllidium ciliare (L) Hampe

1 27 39 46 132 141

Dum

(22) 28 3946 (57) 87 88 89 90 92

Ptilidium ~lcherrium (G Web) Hampe

(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141

47

~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark

46 (139) (140) 9 46 79 116 139

Mannia triandra (Scop) Gro 11e

46 (140)

Marchantia pound91ymorpha L

1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147

Mo~rckia hibernica (HookJ Gott

46 89

Metzgeria conjugata Lindb

44

46

Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo

39 46 141 (35)45 46

Nowellia curvifolia (Dicks) Mitt

17 27 3846 88 89 90 141

Odontoschisma denudatuJI1 (Nees ~ Mart)

Dum 46 81 137 141

Pallavicinia lyellii (Hook~ Carruth

90

Odontoschisma macounii (Aust) Under

45 46 (66) (139) 140 141

45

=

Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum

27 39 46 90 13724 39 46

Pellia neesiana Limp bull Phaeoceros

39 46 89 27 39 45 46 97

Porella pinnata L

13 27 I 46 90 Plagiochila asplenioides (L) Dum

27 I 39 1+6 88 89 I 90 137

46

~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb

17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77

Preissia quadrata (Seop) Nees

1 36 3946 (71) 88 89 90 121 123

137

Ptllidium ciliare (L) Hampe

1 27 39 46 132 141

Dum

(22) 28 3946 (57) 87 88 89 90 92

Ptilidium ~lcherrium (G Web) Hampe

(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141

47

Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo

39 46 141 (35)45 46

Nowellia curvifolia (Dicks) Mitt

17 27 3846 88 89 90 141

Odontoschisma denudatuJI1 (Nees ~ Mart)

Dum 46 81 137 141

Pallavicinia lyellii (Hook~ Carruth

90

Odontoschisma macounii (Aust) Under

45 46 (66) (139) 140 141

45

=

Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum

27 39 46 90 13724 39 46

Pellia neesiana Limp bull Phaeoceros

39 46 89 27 39 45 46 97

Porella pinnata L

13 27 I 46 90 Plagiochila asplenioides (L) Dum

27 I 39 1+6 88 89 I 90 137

46

~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb

17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77

Preissia quadrata (Seop) Nees

1 36 3946 (71) 88 89 90 121 123

137

Ptllidium ciliare (L) Hampe

1 27 39 46 132 141

Dum

(22) 28 3946 (57) 87 88 89 90 92

Ptilidium ~lcherrium (G Web) Hampe

(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141

47

=

Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum

27 39 46 90 13724 39 46

Pellia neesiana Limp bull Phaeoceros

39 46 89 27 39 45 46 97

Porella pinnata L

13 27 I 46 90 Plagiochila asplenioides (L) Dum

27 I 39 1+6 88 89 I 90 137

46

~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb

17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77

Preissia quadrata (Seop) Nees

1 36 3946 (71) 88 89 90 121 123

137

Ptllidium ciliare (L) Hampe

1 27 39 46 132 141

Dum

(22) 28 3946 (57) 87 88 89 90 92

Ptilidium ~lcherrium (G Web) Hampe

(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141

47

~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb

17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77

Preissia quadrata (Seop) Nees

1 36 3946 (71) 88 89 90 121 123

137

Ptllidium ciliare (L) Hampe

1 27 39 46 132 141

Dum

(22) 28 3946 (57) 87 88 89 90 92

Ptilidium ~lcherrium (G Web) Hampe

(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141

47

Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi

39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)

Riccardia ~atifrons Lindb

1 27 39 46 89

Riccardia multifi~ (L) S Gray

39 46 88 145

Riccia beyrichiana Hampe ~ Lehm

46 Riccardia palmata (Hedw) Carruth

39 46 90

48

Riccia bifurca Hoffm Riccia cavernosa Hoffm

48 13739 46 79 137

Riccia fluitans L

242731454679

Riccla frostli Aust

48 79

Riccia sullicantii Aust

79 (137) Riccia sorocarpa Bisch 46 (137)

49

Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce

13 27 46 79 89 110 39 46 (139) 141

ScaJltll1ia carinthica Jack ~ Lindb

137 t 141

Scapania ~ (Mart) Durn

39467990137141

Scapania digenil Schiffn

90

ScaEsectflia cuspiduligera (Nees) K Mull

46 (139) (140) (141)

50

Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal

39 46 (139) 141 41 44 46 64 (139) (140) 141

ScaP3poundlia irrigua Gott ~ almiddot

7 39 46 79 137 (140L 141

Seapania nemorosa (L) Dum

1 39 46 89 90 137

Scapmi~ ~~ta Smiddotlch

137 141 51

~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust

46 (66) 141 136 141

Sc~~ scandica (H Arnell ~ Buch) Macv

139 140

Scapania subalpina (Nees) Durn

39 40 46 79 137 141

Scapania undulata (L) Dum

1 42 46 79 141

Scapania umbrosa (Schrad) Dum

137 140 141

52

Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn

27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141

Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch

39 -t6 89 90 136 140 141 7 39 46 141

Trltomaria scitula (Tayl) Joerg

46 136 (139) 141

53

Riccia bifurca Hoffm Riccia cavernosa Hoffm

48 13739 46 79 137

Riccia fluitans L

242731454679

Riccla frostli Aust

48 79

Riccia sullicantii Aust

79 (137) Riccia sorocarpa Bisch 46 (137)

49

Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce

13 27 46 79 89 110 39 46 (139) 141

ScaJltll1ia carinthica Jack ~ Lindb

137 t 141

Scapania ~ (Mart) Durn

39467990137141

Scapania digenil Schiffn

90

ScaEsectflia cuspiduligera (Nees) K Mull

46 (139) (140) (141)

50

Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal

39 46 (139) 141 41 44 46 64 (139) (140) 141

ScaP3poundlia irrigua Gott ~ almiddot

7 39 46 79 137 (140L 141

Seapania nemorosa (L) Dum

1 39 46 89 90 137

Scapmi~ ~~ta Smiddotlch

137 141 51

~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust

46 (66) 141 136 141

Sc~~ scandica (H Arnell ~ Buch) Macv

139 140

Scapania subalpina (Nees) Durn

39 40 46 79 137 141

Scapania undulata (L) Dum

1 42 46 79 141

Scapania umbrosa (Schrad) Dum

137 140 141

52

Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn

27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141

Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch

39 -t6 89 90 136 140 141 7 39 46 141

Trltomaria scitula (Tayl) Joerg

46 136 (139) 141

53

Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce

13 27 46 79 89 110 39 46 (139) 141

ScaJltll1ia carinthica Jack ~ Lindb

137 t 141

Scapania ~ (Mart) Durn

39467990137141

Scapania digenil Schiffn

90

ScaEsectflia cuspiduligera (Nees) K Mull

46 (139) (140) (141)

50

Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal

39 46 (139) 141 41 44 46 64 (139) (140) 141

ScaP3poundlia irrigua Gott ~ almiddot

7 39 46 79 137 (140L 141

Seapania nemorosa (L) Dum

1 39 46 89 90 137

Scapmi~ ~~ta Smiddotlch

137 141 51

~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust

46 (66) 141 136 141

Sc~~ scandica (H Arnell ~ Buch) Macv

139 140

Scapania subalpina (Nees) Durn

39 40 46 79 137 141

Scapania undulata (L) Dum

1 42 46 79 141

Scapania umbrosa (Schrad) Dum

137 140 141

52

Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn

27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141

Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch

39 -t6 89 90 136 140 141 7 39 46 141

Trltomaria scitula (Tayl) Joerg

46 136 (139) 141

53

Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal

39 46 (139) 141 41 44 46 64 (139) (140) 141

ScaP3poundlia irrigua Gott ~ almiddot

7 39 46 79 137 (140L 141

Seapania nemorosa (L) Dum

1 39 46 89 90 137

Scapmi~ ~~ta Smiddotlch

137 141 51

~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust

46 (66) 141 136 141

Sc~~ scandica (H Arnell ~ Buch) Macv

139 140

Scapania subalpina (Nees) Durn

39 40 46 79 137 141

Scapania undulata (L) Dum

1 42 46 79 141

Scapania umbrosa (Schrad) Dum

137 140 141

52

Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn

27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141

Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch

39 -t6 89 90 136 140 141 7 39 46 141

Trltomaria scitula (Tayl) Joerg

46 136 (139) 141

53

~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust

46 (66) 141 136 141

Sc~~ scandica (H Arnell ~ Buch) Macv

139 140

Scapania subalpina (Nees) Durn

39 40 46 79 137 141

Scapania undulata (L) Dum

1 42 46 79 141

Scapania umbrosa (Schrad) Dum

137 140 141

52

Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn

27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141

Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch

39 -t6 89 90 136 140 141 7 39 46 141

Trltomaria scitula (Tayl) Joerg

46 136 (139) 141

53

Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn

27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141

Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch

39 -t6 89 90 136 140 141 7 39 46 141

Trltomaria scitula (Tayl) Joerg

46 136 (139) 141

53