neo-tropical costus - gingers. · •costus – 100+ species, mostly neo-tropical, about 20 ......

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Neo-Tropical Costus Dave Skinner www.gingersrus.com

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Page 1: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Neo-Tropical Costus

Dave Skinner

www.gingersrus.com

Page 2: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Plant Family Costaceae

• Costus – 100+ species, mostly neo-tropical, about 20

species in Africa

• Chamaecostus – 7 species, South America

• Cheilocostus – 4+ species, Asia-Australia

• Paracostus – 2 species, Africa and Borneo

• Monocostus – 1 specie, Peru

• Dimerocostus – 2 species, Central & South America

• Tapeinochilos – 16 species, New Guinea-Australia

Page 3: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Bract Types Appendaged Not Appendaged

Page 4: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Bract Colors

Red Bracts Green, Yellow, Orange

Exposed part vs. covered

Page 5: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Terminal or Basal Inflorescence Basal Inflorescence Terminal

Inflorescence

Page 6: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Flower Types Tubular Not Tubular

Page 7: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Flower Parts Calyx Length

Page 8: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Bracteole Types Boat Shaped Bicarinate

Page 9: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Ligule Length and Shape

Page 10: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Other Characters

• Leaf base (cordate, cuneate, rounded, etc)

• Leaf apex (long or short acuminate)

• Leaf dimensions and size (linear vs. wide)

• Bract margins (fibrous or entire)

• Indument (hairiness) NOTE: this tends to

be variable within most species!

Page 11: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Four Main Categories

• Non-Appendaged Bracts with Tubular

Flowers

• Non-Appendaged Bracts with Non-Tubular

Flowers

• Appendaged Bracts with Tubular Flowers

• Appendaged Bracts with Non-Tubular

Flowers

Page 12: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus species with

Non-Appendaged Bracts

and

Tubular Flowers

Page 13: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus scaber (cultivated plant)

Page 14: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

The “real” Costus spicatus (Tagged C. cylindricus at Smithsonian)

Page 15: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus scaber from Osa Peninsula, Costa Rica

Page 16: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus scaber from Cerro Nara, Costa Rica

Page 17: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus scaber near Gualaquiza, Ecuador

Page 18: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus aff. scaber (Alto Mayo, Peru)

Page 19: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus woodsonii near Parque Nacionál Cahuita, Costa Rica

Page 20: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Parque Nacionál Cahuita, Costa Rica

Page 21: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus wilsonii left, Reserva Durika Cord. de Talamanca (2000

meters) Costa Rica, above from Carla Black’s

garden. Volcán, Panamá

Page 22: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus pulverulentus cultivated plant

Page 23: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus pulverulentus near Parque Nacionál La Cangreja Costa Rica

Page 24: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus pulverulentus in Ecuador

Page 25: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus pulverulentus (X villosissimus?)

near Rancho Mastatal Costa Rica

Page 26: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus chartaceus Costus erythrothyrsus cultivated plants

Page 27: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus plicatus Costus lasius Osa Peninsula, Costa Rica

Page 28: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus lasius Urco Mirano, Peru

Page 29: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus vinosus at Marie Selby Gardens

Page 30: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus stenophyllus Estación La Gamba Costa Rica

Page 31: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus erythrocoryne near Iquitos, Peru

Page 32: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus vargassii Carla Black Garden, Panama

Page 33: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus species with

Non-Appendaged Bracts

and

Non-Tubular Flowers

(Broad Labellum)

Page 34: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus laevis (Osa Peninsula, Costa Rica)

Page 35: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus laevis with characteristic

silvery midrib on young foliage

Page 36: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus laevis near Pocosol, Cordillera de Tilarán, Costa Rica

Page 37: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus laevis El Copé, Panama

Page 38: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus laevis with orange flower near Rancho Mastatal, Costa Rica

Page 39: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus pictus (cultivated plants)

Page 40: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus arabicus (cultivated plant)

Page 41: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus arabicus Quebrada Cayapoza, Peru

Cordate Leaf Base

Page 42: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus amazonicus near Gualaquiza,

Morona-Santiago Province, Ecuador

Page 43: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus malortieanus in Cord. de Tilarán near Poco Sol

Page 44: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Cerro Nara Costus glaucus Osa Peninsula

P.N. La Cangreja

Page 45: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus species with

Appendaged Bracts

and

Tubular Flowers

Page 46: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus barbatus (hort. at Marie Selby Gardens)

Page 47: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Maas’description does not fit this plant!

1. The ligules are short and truncate - Maas describes as 15-40 mm) and lobed

2. It is often found producing BASAL instead of terminal inflorescences.

3. The margins of the bracts, bracteoles and calyx are entire, NOT “delacerating into fibers”

4. The corolla lobes of the plant in horticulture are NOT puberulous, a character of the real Costus barbatus, but rarely found in Costus

5. None of the plant parts are particularly hairy, as mentioned to distinguish this from Costus montanus.

Page 48: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus montanus near P. N. Tapantí, Costa Rica

Page 49: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus productus

Page 50: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus curvibracteatus Rio Piro, Costa Rica

Page 51: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus aff. curvibracteatus near Poco Sol, Cord. de Tilarán, Costa Rica

Page 52: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus lima Osa Peninsula, Costa Rica

Page 53: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus lima Carla Black Garden

Panama

Page 54: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza
Page 55: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus aff. ricus Cerro Mariposa, Panama

Page 56: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Boat Shaped Bracteole Bicarinate (2 keeled) Bracteole

Costus lima Costus ricus

Comparison of Bracteole Types

Page 57: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus lima Costus ricus

Costus lima and Costus Ricus

Page 58: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus ricus Cerro Nara, Costa Rica

Page 59: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus aff. Lima (hybrid?) Rio Rincon, Osa Peninsula, Costa Rica

Page 60: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus osae in P. N. Piedras Blancas Rio Bonito and Quebrada Negra

Page 61: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus sp. Cabo Matapalo

on the southern tip of the Osa Peninsula, Costa Rica

Page 62: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus sp. Cabo Matapalo habitat

Page 63: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus species with

Appendaged Bracts

and

Non-Tubular Flowers

(Broad Labellum)

Page 64: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus

guanaiensis (various locations in

Costa Rica)

Page 65: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus

guanaiensis near La Fortuna,

Panama

Page 66: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus guanaiensis

var. guanaiensis

(photos from Ecuador and Colombia)

Page 67: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus guanaiensis

var. macrostrobilus (?) (Ecuador, near Reserva Bilsa)

Costus guanaiensis

var. tarmicus (common in Ecuador)

Page 68: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus guanaiensis (plicate leaf form found at Reserva Bilsa, Ecuador)

Page 69: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus sp. found near

Guayzimi, Zamora, Ecuador

Page 70: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus bracteatus near Shiroles, Costa Rica

Page 71: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus erythrophyllus (cultivated plant)

Page 72: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus varzearum (Lyon Arboretum)

Page 73: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

(above from near Pachijal, Ecuador)

Costus leucanthus (right from Waimea Arboretum)

Page 74: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus longebracteolatus (cultivated plant)

Page 75: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus longebracteolatus (San Pedro, Peru)

Page 76: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus villosissimus (near Rancho Mastatal, Costa Rica)

Page 77: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus asplundii Lyon Arboretum

Page 78: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Unidentified Costus

Two Possible New Species from

Cordillera del Condor area

Southern Ecuador

Page 79: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Basal flowering Costus from Rio Nangaritza

Cord. del Condor area of southern Ecuador

Page 80: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

?Costus zamoranus?

John Mood Garden, Hawaii

Page 81: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

Costus from Rio Numbaime

Cord. del Condor area of southern Ecuador

Page 82: Neo-Tropical Costus - gingers. · •Costus – 100+ species, mostly neo-tropical, about 20 ... Cordillera del Condor area Southern Ecuador . Basal flowering Costus from Rio Nangaritza

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