Atlas of the Bryophytes of WisconsinJeff Rose

Department of BotanyUniversity of Wisconsin-Madison

07 November, 2016

Contents

Diversity of Bryophytes by County 2

History of Bryology in Wisconsin 3

Atlas Methodology 3

Mosses 4Floristics . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4About the Maps & Nomenclature . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4

Hornworts and Liverworts 52Floristics . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 52About the Maps & Nomenclature . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 52

1

Diversity of Bryophytes by County

number of species

under 99 − 4747 − 7373 − 100100 − 146146 − 190190 − 218218 − 254over 254

125

253

126

313

3929

58

41

39

31

92

53218

32

216

283

8

76

63

89

90

190

61

65

141

156

113

99

130

31

26

33

73

32178

91

198

78

27

53

172

43

84

181

70

136

047

135

190

47

65

209

113

42

254

133

88

87

980

77

159

215

100

53

95

146

60

38 7

84

2

History of Bryology in Wisconsin

The earilest collections of Wisconsin bryophytes date back to those of Increase Lapham from the 1830’s. Otherimportant collectors or students of Wisconsin Bryophytes include C. Barnes (WIS), F. Bowers (UWSP), L.Cheney (WIS), G. Conklin (WIS), J. Christy (MPM), R. Evans (WIS), S. Freckmann (UWSP), H. Iltis(WIS), R. Koch (UWL), and M. Nee (WIS). As seen in chart below, the peak activity in brophyte collectingwas between 1960-1980 with over 10,000 collections in that time frame.

year

Col

lect

ions

010

0020

0030

0040

0050

00

1800 1840 1880 1920 1960 2000

Atlas Methodology

Records were downloaded from the Consortium of North American Bryophyte Herbaria websiteand supplemented by the database at the Milwaukee Public Museum (MPM). Currently, 35127 Wisconsincollections are databased. Names were then examined to check for synonomy, remove unidentified specimens,and remove specimens lacking locality data. When possible, dubious specimens at WIS have been checked.However, specimens from other herbria have not been checked unless they are present in duplicate at WIS.In cases where the identifications are highly dubious, these collecteions have not been maped and the speciesexcluded from the flora for the time being.

3

Mosses

Floristics

Mosses (Bryophyta) can be found in all terrestrial and aquatic habitats throughout Wisconsin. Habitatsparticulalry well-suited for large amounts of moss diversity include cliff faces, wetlands (bogs, fens) andparticularly white cedar (Thuja occidentalis) swamps. The distribution and diversity of mosses is far fromuniform throughout the states (see map). This is due to a combiantion of non-uniform distributuion ofhabitats and collection bias towards more “scenic” areas. The problem of collection bias is further hamperedby a lack of bryologists acitvley collecting in Wisconsin in the last several decades. Areas of the highestmoss diversity in the state are (1) the Driftless Area, particularly the Baraboo Hills, (2) the Wisconsin RiverDrainage, (3) the Lake Superior shore, and (4) the Door Peninsula. Conversley, areas of low diversity are(1) West-Central Wisconsin and (2) the Lake Winnebago Area. High divertsity areas (1), (2), and (4) arealso well-known for their high diversity of vascular plant species.

The Baraboo Hills contains species that likely persisted in the Driftless Area during Pleistocene glaciation,but also contains some northern species disjunct from the northern part of the state. Driftless area specialitiesinclude Bryoxiphium norvegicum, Dicranodontium denudatum and Jaffueliobryum spp. Another floristicallyimportant feature of the Driftless Area is several Ozarkian species that enter the state in Grant Countyincluding Forsstroemia trichomitria and Zygodon viridissimus. Associated with the diversity of the Driftlessarea is a noticible increase in diversity along the Wisconsin River (and apparent slight decrease in theAdams/Jeauneau County areas of glacial Lake Wisconsin). This seems to be sometimes associated withspecies that are more widespread in the northern portion of the state but follow the length of the river to theMississippi River. Examples of this pattern include Homalia trichomanoides, Isopterygiopsis muelleriana,Mnium lycopodioides, and Saelania glaucescens.

Door County is signifigant in its mixture of northern and southern habitats, but also supports several GreatLakes dune species sich as Niphotrichum cansecens.

Areas of low diversity are misleading in that the exceptionally low numbers are more than likely due toundercollection (for example, Pepin County likely has at least one moss species), although habitat loss dueto agriculture and/or relative habitat uniformity is likely to be an additional factor. These areas shouldspecifically be targeted for additional records, especially the west-central part of the state, where species ofa slightly more western distribution may yet be found.

About the Maps & Nomenclature

The moss flora of Wisconsin presently contains 415 species in 45 families. Nomenclature follows Flora ofNorth America volumes 27 and 28. The following taxa have been excluded from the flora due to probablemisidentifications (specimens are not at WIS), nomenclatural issues, or problems with georeferencing:

Amphidium mougeotii, Aulacomnium turgidum, Brachelyma subulatum, Brachythecium mildeanum, Bryumsandbergii, Bryum turbinatum, Campylophyllum halleri, Claopodium whippleanum, Dichodontium olympicum,Dicranoweisia cirrata, Dicranum crassinerve, Dicranum elongatum, Dicranum majus, Dicranum ouiscon-sinense, Dicranum spurium, Dicranum strictum, Dicranum tauricum, Grimmia donniana, Hennediellaheimii, Homalothecium aureum, Hygrohypnum smithii, Hypnum circinale, Hypnum ravaudii, Leucobryumantillarum, Mnium hornum, Oncophorus rauei, Orthotrichum affine, Orthotrichum cupulatum, Philonotisgracillima, Philonotis longiseta, Plagiothecium micans, Pogonatum brachyphyllum, Rhytidiadelphus loreus,Scleropodium obtusifolium, Scopelophila ligulata, Sphagnum torreyanum, Sphagnum wilsonii var. quinquefar-ium, Taxiphyllum geophilum, Tetraplodon mnioides, Thamnobryum alleghaniense, Thuidium tamariscinum,Tortula muralis

4

Abietinella abietina(Hedw.) M. Fleisch.

Thuidiaceae

Acaulon muticum(Schreb.) Mull. Hal.

Pottiaceae

Amblyodon dealbatus(Sw.) Bruch & Schimp.

Meesiaceae

Amblystegium serpens(Hedw.) Schimp.Amblystegiaceae

Amphidium lapponicum(Hedw.) Schimp.Orthotrichaceae

Anacamptodon splachnoides(Froel.) Brid.Fabroniaceae

Andreaea rupestrisHedw.

Andreaeaceae

Anomobryum concinnatum(Schrad.) Schimp.

Bryaceae

Anomobryum julaceum(Schrad.) Schimp.

Bryaceae

5

Anomodon attenuatus(Hedw.) Hub.Thuidiaceae

Anomodon minor(Hedw.) Lindb.Thuidiaceae

Anomodon rostratus(Hedw.) Schimp.

Thuidiaceae

Anomodon rugelii(Mull. Hal.) Keissl.

Thuidiaceae

Anomodon tristis(Ces.) Sull. & Lesq.

Thuidiaceae

Anomodon viticulosus(Hedw.) Hook. & Taylor

Thuidiaceae

Aphanorrhegma serratum(Hook. & Wilson) Sull.

Funariaceae

Archidium alternifolium(Dicks.) Mitt.Archidiaceae

Atrichum altecristatum(Ren. & Card.) Smyth & Smyth

Polytrichaceae

6

Atrichum angustatum(Brid.) Bruch & Schimp.

Polytrichaceae

Atrichum crispum(James) Sull.

Polytrichaceae

Atrichum oerstedianum(Mull. Hal.) Mitt.Polytrichaceae

Atrichum tenellum(Rahl.) Bruch & Schimp.

Polytrichaceae

Atrichum undulatum(Hedw.) P. Beauv.

Polytrichaceae

Aulacomnium androgynum(Hedw.) Schwagr.Aulacomniaceae

Aulacomnium heterostichum(Hedw.) Bruch & Schimp.

Aulacomniaceae

Aulacomnium palustre(Hedw.) Schwagr.Aulacomniaceae

Barbula convolutaHedw.

Pottiaceae

7

Barbula unguiculataHedw.

Pottiaceae

Bartramia pomiformisHedw.

Bartramiaceae

Blindia acuta(Hedw.) Bruch & Schimp.

Seligeriaceae

Brachytheciastrum velutinum(Hedw.) Ignatov & Huttunen

Brachytheciaceae

Brachythecium acuminatum(Hedw.) Aust.

Brachytheciaceae

Brachythecium acutum(Mitt.) Sull.

Brachytheciaceae

Brachythecium campestre(Mull. Hal.) Schimp.Brachytheciaceae

Brachythecium erythrorrhizonSchimp.

Brachytheciaceae

Brachythecium laetum(Brid.) Schimp.

Brachytheciaceae

8

Brachythecium rivulareSchimp.

Brachytheciaceae

Brachythecium rotaeanumDe Not.

Brachytheciaceae

Brachythecium rutabulum(Hedw.) Schimp.Brachytheciaceae

Brachythecium turgidum(Hartm.) Kindb.

Brachytheciaceae

Brothera leana(Sull.) Mull. Hal.

Dicranaceae

Brotherella recurvans(Michx.) M. Fleisch.Sematophyllaceae

Bruchia flexuosa(Schwagr.) Mull. Hal.

Bruchiaceae

Bryhnia graminicolor(Brid.) Grout

Brachytheciaceae

Bryhnia novae−angliae(Sull. & Lesq.) Grout

Brachytheciaceae

9

Bryoerythrophyllum recurvirostre(Hedw.) Chen

Pottiaceae

Bryoxiphium norvegicum(Brid.) Mitt.

Bryoxiphiaceae

Bryum argenteumHedw.

Bryaceae

Bucklandiella heterosticha(Hedw.) Bednarek−Ochyra & Ochyra

Grimmiaceae

Buxbaumia aphyllaHedw.

Buxbaumiaceae

Callicladium haldanianum(Grev.) H.A. Crum

Hypnaceae

Calliergon cordifolium(Hedw.) Kindb.

Amblystegiaceae

Calliergon giganteum(Schimp.) Kindb.Amblystegiaceae

Calliergon richardsonii(Mitt.) Kindb.

Amblystegiaceae

10

Calliergonella cuspidata(Hedw.) Loeske

Hypnaceae

Campyliadelphus chrysophyllus(Brid.) R.S. ChopraAmblystegiaceae

Campylium stellatum(Hedw.) C.E.O. Jensen

Amblystegiaceae

Campylophyllum hispidulum(Brid.) Hedenas

Hypnaceae

Ceratodon purpureus(Hedw.) Brid.Ditrichaceae

Cirriphyllum piliferum(Hedw.) Grout

Brachytheciaceae

Clasmatodon parvulus(Hampe) Sull.Fabroniaceae

Climacium americanumBrid.

Climaciaceae

Climacium dendroides(Hedw.) F. Weber & D. Mohr

Climaciaceae

11

Codriophorus acicularis(Hedw.) P. Beauv.

Grimmiaceae

Conardia compacta(Mull. Hal.) H. Rob.Amblystegiaceae

Coscinodon calyptratus(Drumm.) C.E.O. Jensen

Grimmiaceae

Cratoneuron filicinum(Hedw.) Spruce

Amblystegiaceae

Ctenidium molluscum(Hedw.) Mitt.Hypnaceae

Cynodontium polycarpum(Hedw.) Schimp.

Dicranaceae

Cynodontium strumiferum(Hedw.) Lindb.Dicranaceae

Cynodontium tenellum(Schimp.) Limpr.

Dicranaceae

Cyrtohypnum minutulum(Hedw.) W.R. Buck & H.A. Crum

Thuidiaceae

12

Cyrtohypnum pygmaeum(Schimp.) W.R. Buck & H.A. Crum

Thuidiaceae

Dichelyma capillaceum(With.) Myrin

Fontinalaceae

Dichelyma falcatum(Hedw.) MyrinFontinalaceae

Dichelyma pallescensBruch & Schimp.

Fontinalaceae

Dichodontium pellucidum(Hedw.) Schimp.

Dicranaceae

Dicranella cerviculata(Hedw.) Schimp.

Dicranaceae

Dicranella heteromalla(Hedw.) Schimp.

Dicranaceae

Dicranella palustris(Dicks.) A.C. Crundwell

Dicranaceae

Dicranella rufescens(With.) Schimp.

Dicranaceae

13

Dicranella schreberiana(Hedw.) Hilf.Dicranaceae

Dicranella staphylinaH. WhitehouseDicranaceae

Dicranella subulata(Hedw.) Schimp.

Dicranaceae

Dicranella varia(Hedw.) Schimp.

Dicranaceae

Dicranodontium denudatum(Brid.) E. Britton

Dicranaceae

Dicranoweisia crispula(Hedw.) MildeDicranaceae

Dicranum bonjeaniiDe Not.

Dicranaceae

Dicranum condensatumHedw.

Dicranaceae

Dicranum flagellareHedw.

Dicranaceae

14

Dicranum fulvumHook.

Dicranaceae

Dicranum fuscescensTurner

Dicranaceae

Dicranum montanumHedw.

Dicranaceae

Dicranum muehlenbeckiiBruch & Schimp.

Dicranaceae

Dicranum ontarienseW.L. PetersonDicranaceae

Dicranum polysetumSw.

Dicranaceae

Dicranum scopariumHedw.

Dicranaceae

Dicranum undulatumSchrad.

Dicranaceae

Dicranum viride(Sull. & Lesq.) Lindb.

Dicranaceae

15

Didymodon fallax(Hedw.) R.H. Zander

Pottiaceae

Didymodon rigidulusHedw.

Pottiaceae

Didymodon tophaceus(Brid.) LisaPottiaceae

Didymodon vinealis(Brid.) R.H. Zander

Pottiaceae

Diphyscium foliosum(Hedw.) D. MohrBuxbaumiaceae

Distichium capillaceum(Hedw.) Bruch & Schimp.

Ditrichaceae

Distichium inclinatum(Hedw.) Bruch & Schimp.

Ditrichaceae

Ditrichum flexicaule(Schwagr.) Hampe

Ditrichaceae

Ditrichum heteromallum(Hedw.) E. Britton

Ditrichaceae

16

Ditrichum lineare(Sw.) Lindb.Ditrichaceae

Ditrichum pallidum(Hedw.) Hampe

Ditrichaceae

Ditrichum pusillum(Hedw.) Hampe

Ditrichaceae

Drepanocladus aduncus(Hedw.) Warnst.Amblystegiaceae

Drepanocladus longifolius(Mitt.) Broth.

Amblystegiaceae

Drepanocladus polygamus(Schimp.) HedenasAmblystegiaceae

Drepanocladus sordidus(Mull. Hal.) Hedenas

Amblystegiaceae

Drummondia prorepens(Hedw.) E. BrittonOrthotrichaceae

Elodium blandowii(F. Weber & D. Mohr) Eckel

Thuidiaceae

17

Elodium paludosum(Aust.) Broth.Thuidiaceae

Encalypta ciliataHedw.

Encalyptaceae

Encalypta proceraBruch

Encalyptaceae

Encalypta rhaptocarpaSchwagr.

Encalyptaceae

Entodon brevisetus(Hook. & Wilson) Lindb.

Entodontaceae

Entodon challengeri(Paris) CardotEntodontaceae

Entodon cladorrhizans(Hedw.) Mull. Hal.

Entodontaceae

Entodon seductrix(Hedw.) Mull. Hal.

Entodontaceae

Ephemerum cohaerens(Hedw.) HampeEphemeraceae

18

Ephemerum crassinervium(Schwagr.) Hampe

Ephemeraceae

Ephemerum spinulosumBruch & Schimp.Ephemeraceae

Eucladium verticillatum(Hedw.) Bruch & Schimp.

Pottiaceae

Eurhynchiastrum pulchellum(Hedw.) Ignatov & Huttunen

Brachytheciaceae

Eurhynchium praelongum(Hedw.) Schimp.Brachytheciaceae

Eurhynchium striatum(Schreb.) Schimp.Brachytheciaceae

Fabronia ciliaris(Brid.) Brid.

Fabroniaceae

Fissidens adianthoidesHedw.

Fissidentaceae

Fissidens bryoidesHedw.

Fissidentaceae

19

Fissidens bushii(Cardot & Ther.) Cardot & Ther.

Fissidentaceae

Fissidens dubiusP. Beauv.

Fissidentaceae

Fissidens elegansBrid.

Fissidentaceae

Fissidens exilisHedw.

Fissidentaceae

Fissidens fontanus(Bach. Pyl.) Steud.

Fissidentaceae

Fissidens grandifronsBrid.

Fissidentaceae

Fissidens minutulusSull.

Fissidentaceae

Fissidens obtusifoliusWilson

Fissidentaceae

Fissidens osmundioidesHedw.

Fissidentaceae

20

Fissidens subbasilarisHedw.

Fissidentaceae

Fissidens taxifoliusHedw.

Fissidentaceae

Fontinalis antipyreticaHedw.

Fontinalaceae

Fontinalis dalecarlicaBruch & Schimp.

Fontinalaceae

Fontinalis hypnoidesHartm.

Fontinalaceae

Fontinalis novae−angliaeSull.

Fontinalaceae

Fontinalis sphagnifolia(Mull. Hal.) Wijk & Margad.

Fontinalaceae

Fontinalis sullivantiiLindb.

Fontinalaceae

Forsstroemia trichomitria(Hedw.) Lindb.

Leucodontaceae

21

Funaria americanaLindb.

Funariaceae

Funaria flavicansMichx.

Funariaceae

Funaria hygrometricaHedw.

Funariaceae

Gemmabryum caespiticum(Hedw.) J.R. Spence

Bryaceae

Gemmabryum dichotomum(Hedw.) J.R. Spence & H.P. Ramsay

Bryaceae

Grimmia laevigata(Brid.) Brid.

Grimmiaceae

Grimmia muehlenbeckiiSchimp.

Grimmiaceae

Grimmia olneyiSull.

Grimmiaceae

Grimmia ovalis(Hedw.) Lindb.Grimmiaceae

22

Grimmia piliferaP. Beauv.

Grimmiaceae

Grimmia plagiopodiaHedw.

Grimmiaceae

Grimmia teretinervisLimpr.

Grimmiaceae

Gymnostomum aeruginosumSm.

Pottiaceae

Hamaticaulis vernicosus(Mitt.) Hedenas

Amblystegiaceae

Haplocladium microphyllum(Hedw.) Broth.Thuidiaceae

Haplocladium virginianum(Brid.) Broth.Thuidiaceae

Hedwigia ciliata(Hedw.) P. Beauv.

Hedwigiaceae

Herzogiella striatella(Brid.) Z. Iwats.

Hypnaceae

23

Herzogiella turfacea(Lindb.) Z. Iwats.

Hypnaceae

Homalia trichomanoides(Hedw.) Schimp.

Neckeraceae

Homalotheciella subcapillata(Hedw.) Broth.

Brachytheciaceae

Homomallium adnatum(Hedw.) Broth.

Hypnaceae

Hygroamblystegium varium ssp. noterophilum(Sull. & Lesq.) Vanderp. & Hedenas

Amblystegiaceae

Hygroamblystegium varium ssp. varium(Hedw.) Monk.

Amblystegiaceae

Hygroamblystegium varium var. humile(P. Beauv.) Vanderp. & Hedenas

Amblystegiaceae

Hygrohypnum eugyrium(Schimp.) LoeskeAmblystegiaceae

Hygrohypnum luridum(Hedw.) Jenn.

Amblystegiaceae

24

Hygrohypnum ochraceum(Turner) Loeske

Amblystegiaceae

Hylocomiastrum pyrenaicum(Spruce) M. Fleisch.

Hylocomiaceae

Hylocomium splendens(Hedw.) Schimp.Hylocomiaceae

Hymenostylium recurvirostre(Hedw.) Dix.Pottiaceae

Hyophila involuta(Hook.) A. Jaeger

Pottiaceae

Hypnum cupressiformeHedw.

Hypnaceae

Hypnum curvifoliumHedw.

Hypnaceae

Hypnum faurieiCardot

Hypnaceae

Hypnum imponensHedw.

Hypnaceae

25

Hypnum lindbergii(Mitt.) Hedenas

Hypnaceae

Hypnum pallescens(Hedw.) P. Beauv.

Hypnaceae

Hypnum pratenseKoch

Hypnaceae

Imbribryum miniatum(Lesq.) J.R. Spence

Bryaceae

Isopterygiopsis muelleriana(Schimp.) Z. Iwats.Plagiotheciaceae

Isopterygiopsis pulchella(Hedw.) Z. Iwats.Plagiotheciaceae

Jaffueliobryum raui(Aust.) Ther.Grimmiaceae

Jaffueliobryum wrightii(Sull.) Ther.

Grimmiaceae

Leptobryum pyriforme(Hedw.) Wilson

Bryaceae

26

Leptodictyum riparium(Hedw.) Warnst.Amblystegiaceae

Leskea gracilescensHedw.

Leskeaceae

Leskea obscuraHedw.

Leskeaceae

Leskea polycarpaEhrh.

Leskeaceae

Leskeella nervosa(Brid.) LoeskeLeskeaceae

Leucobryum albidum(Brid.) Lindb.Dicranaceae

Leucobryum glaucum(Hedw.) Angstr.

Dicranaceae

Leucodon brachypusBrid.

Leucodontaceae

Leucodon julaceus(Hedw.) Sull.

Leucodontaceae

27

Lindbergia brachyptera(Mitt.) Kindb.Leskeaceae

Meesia triquetra(L.) Angstr.Meesiaceae

Meesia uliginosaHedw.

Meesiaceae

Micromitrium megalosporumAust.

Ephemeraceae

Mnium lycopodioidesSchwagr.Mniaceae

Mnium marginatum(Dicks.) P. Beauv.

Mniaceae

Mnium spinosum(Voit) Schwagr.

Mniaceae

Mnium spinulosumBruch & Schimp.

Mniaceae

Mnium stellareHedw.

Mniaceae

28

Mnium thomsoniiSchimp.

Mniaceae

Myurella julacea(Schwagr.) Schimp.

Theliaceae

Myurella sibirica(Mull. Hal.) Reimers

Theliaceae

Neckera pennataHedw.

Neckeraceae

Niphotrichum canescens(Hedw.) Bednarek−Ochyra & Ochyra

Grimmiaceae

Oncophorus virens(Hedw.) Brid.Dicranaceae

Oncophorus wahlenbergiiBrid.

Dicranaceae

Orthotrichum anomalumHedw.

Orthotrichaceae

Orthotrichum elegansSchwagr.

Orthotrichaceae

29

Orthotrichum obtusifoliumBrid.

Orthotrichaceae

Orthotrichum ohioenseSull. & Lesq.

Orthotrichaceae

Orthotrichum pumilumSw.

Orthotrichaceae

Orthotrichum pusillumMitt.

Orthotrichaceae

Orthotrichum sordidumSull. & Lesq.

Orthotrichaceae

Orthotrichum speciosumNees

Orthotrichaceae

Orthotrichum stellatumBrid.

Orthotrichaceae

Orthotrichum strangulatumP. Beauv.

Orthotrichaceae

Oxyrrhynchium hians(Hedw.) Loeske

Brachytheciaceae

30

Palustriella falcata(Brid.) Hedenas

Amblystegiaceae

Paraleucobryum longifolium(Ehrh.) LoeskeDicranaceae

Philonotis fontana(Hedw.) Brid.

Bartramiaceae

Philonotis marchica(Hedw.) Brid.

Bartramiaceae

Physcomitrium collenchymatumGier

Funariaceae

Physcomitrium hookeriHampe

Funariaceae

Physcomitrium immersumSull.

Funariaceae

Physcomitrium pyriforme(Hedw.) Hampe

Funariaceae

Plagiomnium ciliare(Mull. Hal.) T.J. Kop.

Mniaceae

31

Plagiomnium cuspidatum(Hedw.) T.J. Kop.

Mniaceae

Plagiomnium drummondii(Bruch & Schimp.) T.J. Kop.

Mniaceae

Plagiomnium ellipticum(Brid.) T.J. Kop.

Mniaceae

Plagiomnium medium(Bruch & Schimp.) T.J. Kop.

Mniaceae

Plagiomnium rostratum(Schrad.) T.J. Kop.

Mniaceae

Plagiopus oederianus(Sw.) H.A. Crum & L.E. Anderson

Bartramiaceae

Plagiothecium cavifolium(Brid.) Z. Iwats.

Plagiotheciaceae

Plagiothecium denticulatum(Hedw.) Schimp.Plagiotheciaceae

Plagiothecium laetumSchimp.

Plagiotheciaceae

32

Plagiothecium latebricolaSchimp.

Plagiotheciaceae

Platydictya confervoides(Brid.) H.A. CrumAmblystegiaceae

Platydictya jungermannioides(Brid.) H.A. CrumAmblystegiaceae

Platydictya minutissimum(Sull. & Lesq.) H.A. Crum

Amblystegiaceae

Platydictya subtilis(Hedw.) H.A. CrumAmblystegiaceae

Platygyrium repens(Brid.) Schimp.

Hypnaceae

Pleuridium subulatum(Hedw.) Rabenh.

Ditrichaceae

Pleurozium schreberi(Willd.) Mitt.

Hylocomiaceae

Pogonatum urnigerum(Hedw.) P. Beauv.

Polytrichaceae

33

Pohlia annotina(Hedw.) Lindb.

Bryaceae

Pohlia bulbifera(Warnst.) Warnst.

Bryaceae

Pohlia cruda(Hedw.) Lindb.

Bryaceae

Pohlia elongataHedw.

Bryaceae

Pohlia filum(Schimp.) Mertensson

Bryaceae

Pohlia lescuriana(Sull.) OchiBryaceae

Pohlia melanodon(Brid.) A.J. Shaw

Bryaceae

Pohlia nutans(Hedw.) Lindb.

Bryaceae

Pohlia obtusifolia(Vill.) L.F. Koch

Bryaceae

34

Pohlia proligera(Kindb.) S.O. Lindberg

Bryaceae

Pohlia wahlenbergii(F. Weber & D. Mohr) A.L. Andrews

Bryaceae

Polytrichastrum alpinum(Hedw.) G.L. Sm.

Polytrichaceae

Polytrichastrum formosum(Hedw.) G.L. Sm.

Polytrichaceae

Polytrichastrum longisetum(Sw.) G.L. Sm.Polytrichaceae

Polytrichastrum ohioense(Ren. & Card.) G.L. Sm.

Polytrichaceae

Polytrichastrum pallidisetum(Funck) G.L. Sm.Polytrichaceae

Polytrichum communeHedw.

Polytrichaceae

Polytrichum juniperinumHedw.

Polytrichaceae

35

Polytrichum piliferumHedw.

Polytrichaceae

Polytrichum strictumBrid.

Polytrichaceae

Pseudobryum cinclidioides(Hub.) T.J. Kop.

Mniaceae

Pseudocalliergon trifarium(F. Weber & D. Mohr) Loeske

Amblystegiaceae

Pseudocalliergon turgescens(T. Jensen) LoeskeAmblystegiaceae

Pseudocampylium radicale(P. Beauv.) Vanderp. & Hedenas

Amblystegiaceae

Pseudoleskea patens(Lindb.) Kindb.Leskeaceae

Pseudoleskeella catenulata(Brid.) Kindb.Leskeaceae

Pseudotaxiphyllum distichaceum(Mitt.) Z. Iwats.

Hypnaceae

36

Pseudotaxiphyllum elegans(Brid.) Z. Iwats.

Hypnaceae

Pterigynandrum filiformeHedw.

Pterigynandraceae

Ptilium crista−castrensis(Hedw.) De Not.

Hypnaceae

Ptychostomum cernuum(Hedw.) Hornsch.

Bryaceae

Ptychostomum creberrimum(Taylor) J.R. Spence & H.P. Ramsay

Bryaceae

Ptychostomum cyclophyllum(Schwagr.) J.R. Spence

Bryaceae

Ptychostomum pallens(Sw.) J.R. Spence

Bryaceae

Ptychostomum pallescens(Schleich.) J.R. Spence

Bryaceae

Ptychostomum pendulumHornsch.Bryaceae

37

Ptychostomum pseudotriquetrum(Hedw.) J.R. Spence & H.P. Ramsay

Bryaceae

Ptychostomum weigelii(Spreng.) J.R. Spence

Bryaceae

Pylaisia intricata(Hedw.) Schimp.

Hypnaceae

Pylaisia polyantha(Hedw.) Schimp.

Hypnaceae

Pylaisia selwyniiKindb.

Hypnaceae

Pylaisiadelpha tenuirostris(Bruch & Schimp.) W.R. Buck

Hypnaceae

Rauiella scita(P. Beauv.) Reimers

Thuidiaceae

Rhabdoweisia crispata(Dicks.) Lindb.

Rhabdoweisiaceae

Rhizomnium appalachianumT.J. Kop.

Mniaceae

38

Rhizomnium magnifolium(Horik.) T.J. Kop.

Mniaceae

Rhizomnium pseudopunctatum(Bruch & Schimp.) T.J. Kop.

Mniaceae

Rhizomnium punctatum(Hedw.) T.J. Kop.

Mniaceae

Rhodobryum ontariense(Kindb.) Paris

Bryaceae

Rhynchostegium aquaticumA. Jaeger

Brachytheciaceae

Rhynchostegium serrulatum(Hedw.) A. JaegerBrachytheciaceae

Rhytidiadelphus subpinnatus(Lindb.) T.J. Kop.Hylocomiaceae

Rhytidiadelphus triquetrus(Hedw.) Warnst.Hylocomiaceae

Rhytidium rugosum(Ehrh.) Kindb.

Hylocomiaceae

39

Rosulabryum capillare(Hedw.) J.R. Spence

Bryaceae

Saelania glaucescens(Hedw.) Broth.Ditrichaceae

Sanionia uncinata(Hedw.) Loeske

Amblystegiaceae

Sarmetypnum exannulatum(Schimp.) HedenasAmblystegiaceae

Schistidium agassiziiSull. & Lesq.Grimmiaceae

Schistidium apocarpum(Hedw.) Bruch & Schimp.

Grimmiaceae

Schistidium appalachianumH. H. Blom.

Grimmiaceae

Schistidium atrofuscum(Schimp.) Limpr.

Grimmiaceae

Schistidium confertum(Funck) Bruch & Schimp.

Grimmiaceae

40

Schistidium crassitheciumH.H. Blom

Grimmiaceae

Schistidium dupretii(Ther.) W.A. Weber

Grimmiaceae

Schistidium pulchrumH.H. Blom

Grimmiaceae

Schistidium rivulare(Brid.) Podp.Grimmiaceae

Schistidium virideH.H. Blom & C. Darigo

Grimmiaceae

Schistostega pennata(Hedw.) F. Weber & D. Mohr

Schistostegaceae

Schwetschkeopsis fabronia(Schwagr.) Broth.

Fabroniaceae

Sciuro−hypnum curtum(Lindb.) Ignatov

Brachytheciaceae

Sciuro−hypnum oedipodium(Mitt.) Ignatov & Huttunen

Brachytheciaceae

41

Sciuro−hypnum plumosum(Hedw.) Ignatov & Huttunen

Brachytheciaceae

Sciuro−hypnum populeum(Hedw.) Ignatov & Huttunen

Brachytheciaceae

Sciuro−hypnum reflexum(Starke) Ignatov & Huttunen

Brachytheciaceae

Sciuro−hypnum starkei(Brid.) Ignatov & Huttunen

Brachytheciaceae

Scorpidium cossonii(Schimp.) Hedenas

Scorpidiaceae

Scorpidium revolvens(Sw.) RubersScorpidiaceae

Scorpidium scorpioides(Hedw.) Limpr.Scorpidiaceae

Seligeria campylopodaKindb.

Seligeriaceae

Seligeria donniana(Sm.) Mull. Hal.Seligeriaceae

42

Seligeria pusilla(Hedw.) Bruch & Schimp.

Seligeriaceae

Seligeria recurvata(Hedw.) Bruch & Schimp.

Seligeriaceae

Sematophyllum demissum(Wilson) Mitt.

Sematophyllaceae

Sphagnum andersonianumR.E. Andrus

Sphagnaceae

Sphagnum angustifolium(Warnst.) C.E.O. Jensen

Sphagnaceae

Sphagnum bartlettianumWarnst.

Sphagnaceae

Sphagnum capillifolium(Ehrh.) Hedw.Sphagnaceae

Sphagnum centraleC.E.O. JensenSphagnaceae

Sphagnum compactumLam. & DC

Sphagnaceae

43

Sphagnum contortumSchultz

Sphagnaceae

Sphagnum cuspidatumEhrh.

Sphagnaceae

Sphagnum fallaxH. Klinggr.

Sphagnaceae

Sphagnum fimbriatumWilson

Sphagnaceae

Sphagnum flexuosumDozy & Molk.Sphagnaceae

Sphagnum fuscum(Schimp.) H. Klinggr.

Sphagnaceae

Sphagnum girgensohniiRussow

Sphagnaceae

Sphagnum henryenseWarnst.

Sphagnaceae

Sphagnum imbricatumHornsch.

Sphagnaceae

44

Sphagnum inundatumRussow

Sphagnaceae

Sphagnum lescuriiSull.

Sphagnaceae

Sphagnum magellanicumBrid.

Sphagnaceae

Sphagnum majus(Russow) C.E.O. Jensen

Sphagnaceae

Sphagnum obtusumWarnst.

Sphagnaceae

Sphagnum palustreL.

Sphagnaceae

Sphagnum papillosumLindb.

Sphagnaceae

Sphagnum platyphyllum(Lindb.) Warnst.Sphagnaceae

Sphagnum pulchrum(Lindb.) Warnst.Sphagnaceae

45

Sphagnum recurvumP. Beauv.

Sphagnaceae

Sphagnum ripariumAngstr.

Sphagnaceae

Sphagnum rubellumWilson

Sphagnaceae

Sphagnum russowiiWarnst.

Sphagnaceae

Sphagnum squarrosumCrome

Sphagnaceae

Sphagnum subfulvumSjors

Sphagnaceae

Sphagnum subsecundumNees

Sphagnaceae

Sphagnum subtile(Russow) Warnst.

Sphagnaceae

Sphagnum tenerumSull. & Lesq.

Sphagnaceae

46

Sphagnum teres(Schimp.) Angstr.

Sphagnaceae

Sphagnum warnstorfiiRussow

Sphagnaceae

Sphagnum wulfianumGirg.

Sphagnaceae

Splachnum ampullaceumHedw.

Splachnaceae

Splachnum sphaericumHedw.

Splachnaceae

Straminergon stramineum(Dicks.) HedenasAmblystegiaceae

Syntrichia papillosa(Wilson) Jur.Pottiaceae

Syntrichia ruralis(Hedw.) F. Weber & D. Mohr

Pottiaceae

Taxiphyllum deplanatum(Bruch & Schimp.) M. Fleisch.

Hypnaceae

47

Tetraphis pellucidaHedw.

Tetraphidaceae

Thelia asprella(Schimp.) Sull. & Lesq.

Theliaceae

Thelia hirtella(Hedw.) Sull. & Lesq.

Theliaceae

Thelia lescuriiSull.

Theliaceae

Thuidium delicatulum(Hedw.) Schimp.

Thuidiaceae

Thuidium recognitum(Hedw.) Lindb.Thuidiaceae

Timmia megapolitanaHedw.

Timmiaceae

Tomentypnum nitens(Hedw.) Loeske

Brachytheciaceae

Tortella fragilis(Hook. & Wilson) Limpr.

Pottiaceae

48

Tortella humilis(Hedw.) Jenn.

Pottiaceae

Tortella inclinata(R. Hedw.) Limpr.

Pottiaceae

Tortella tortuosa(Hedw.) Limpr.

Pottiaceae

Tortula acaulon(With.) R.H. Zander

Pottiaceae

Tortula cernua(Hub.) Lindb.Pottiaceae

Tortula mucronifoliaSchwagr.

Pottiaceae

Tortula obtusifolia(Schwagr.) Mathieu

Pottiaceae

Tortula plinthobia(Sull. & Lesq.) Broth.

Pottiaceae

Tortula truncata(Hedw.) Mitt.Pottiaceae

49

Trematodon ambiguus(Hedw.) Hornsch.

Bruchiaceae

Trichostomum spiraleGrout

Pottiaceae

Trichostomum tenuirostre(Hook. & Taylor) Lindb.

Pottiaceae

Ulota crispa(Hedw.) Brid.

Orthotrichaceae

Ulota hutchinsiae(Sm.) Hammar

Orthotrichaceae

Warnstorfia fluitans(Hedw.) Loeske

Amblystegiaceae

Warnstorfia pseudostraminea(Mull. Hal.) Tuom. & T.J. Kop.

Amblystegiaceae

Weissia controversaHedw.

Pottiaceae

Weissia muhlenbergiana(Sw.) W.D. Reese & B.A.E. Lemmon

Pottiaceae

50

Zygodon viridissimus(Dicks.) Brid.

Orthotrichaceae

51

Hornworts and Liverworts

Floristics

Coming soon!

About the Maps & Nomenclature

Nomenclature follows Schuster (1966-1992, Hepaticae and Anthocerotae of North America East of the Hun-drenth Meridian vols. I-VI), sypplemented by Soderstrom et al. (2016, Phytokeys 59:1-828)

The hornwort and liverwort flora presently contains 139 species in 32 families.

The following taxa have been excluded from the flora due to probable misidentifications, nomenclaturalissues, or problems with georeferencing:

Diplophyllum andrewsii, Frullania dilatata, Jungermannia caespiticia, Riccia curtisii, Riccia gougetiana

52

Anastrophyllum hellerianum(Nees) R.M. Schust.Anastrophyllaceae

Anastrophyllum michauxii(F. Weber) H. BuchAnastrophyllaceae

Anastrophyllum minutum(Schreb.) R.M. Schust.

Anastrophyllaceae

Aneura pinguis(L.) Dum.

Aneuraceae

Anthoceros macouniiM. Howe

Anthocerotaceae

Anthoceros punctatusL.

Anthocerotaceae

Asterella tenella(L.) P. Beauv.Aytoniaceae

Athalamia hyalina(Sommerf.) S. Hatt.

Cleveaceae

Barbilophozia attenuata(Nees) Loeske

Anastrophyllaceae

53

Barbilophozia barbata(Schreb.) Loeske

Anastrophyllaceae

Barbilophozia kunzeana(Hub.) K. Muller

Anastrophyllaceae

Bazzania trilobata(L.) Gray

Lepidoziaceae

Blasia pusillaL.

Blasiaceae

Blepharostoma trichophyllum(L.) Dum.

Pseudolepicoleaceae

Calypogeia fissa(L.) Raddi

Calypogeiaceae

Calypogeia integristipulaSteph.

Calypogeiaceae

Calypogeia muelleriana(Schiffn.) K. MullerCalypogeiaceae

Calypogeia neesiana(C. Massal. & Carestia) K. Muller

Calypogeiaceae

54

Calypogeia sphagnicola(Arnell & J. Perss.) Warnst. & Loeske

Calypogeiaceae

Calypogeia suecica(Arnell & J. Perss.) K. Muller

Calypogeiaceae

Cephalozia bicuspidata(L.) Dum.

Cephaloziaceae

Cephalozia catenulata(Hub.) Lindb.

Cephaloziaceae

Cephalozia connivens(Dicks.) Lindb.

Cephaloziaceae

Cephalozia lacinulata(J.B. Jack) SpruceCephaloziaceae

Cephalozia lunulifolia(Dum.) Dum.

Cephaloziaceae

Cephalozia macounii(Aust.) Spruce

Cephaloziaceae

Cephalozia pleniceps(Aust.) Lindb.

Cephaloziaceae

55

Cephaloziella byssacea(Roth) Warnst.

Cephaloziellaceae

Cephaloziella elachista(J.B. Jack) Schiffn.Cephaloziellaceae

Cephaloziella hampeana(Nees) Schiffn.

Cephaloziellaceae

Cephaloziella rubella(Nees) Warnst.

Cephaloziellaceae

Chiloscyphus coadunatus(Sw.) R.M. Schust. & J.J. Engel

Lophocoleaceae

Chiloscyphus pallescens(Ehrh.) Dum.

Lophocoleaceae

Chiloscyphus polyanthos(L.) Corda

Lophocoleaceae

Cladopodiella fluitans(Nees) Jarg.

Cephaloziaceae

Cololejeunea biddlecomiae(Aust.) A. EvansLejeuneaceae

56

Conocephalum salebrosumSzweyk., Buczkowska & Odrzykoski

Conocephalaceae

Dichiton integerrimum(Lindb.) H. Buch

Cephaloziellaceae

Diplophyllum apiculatum(A. Evans) Steph.

Scapaniaceae

Fossombronia foveolataLindb.

Fossombroniaceae

Frullania asagrayanaMont.

Jubulaceae

Frullania bolanderiAust.

Jubulaceae

Frullania brittoniaeA. Evans

Jubulaceae

Frullania eboracensisGott.

Jubulaceae

Frullania inflataGott.

Jubulaceae

57

Frullania oakesianaAust.

Jubulaceae

Frullania planaSull.

Jubulaceae

Frullania ripariaHampe

Jubulaceae

Frullania selwynianaPearson

Jubulaceae

Geocalyx graveolens(Schrad.) NeesGeocalycaceae

Harpanthus drummondii(Taylor) Grolle

Geocalycaceae

Harpanthus scutatus(F. Weber & D. Mohr) Spruce

Geocalycaceae

Jamesoniella autumnalis(DC) Steph.

Jungermanniaceae

Jungermannia lanceolataL.

Jungermanniaceae

58

Jungermannia schraderiMart.

Jungermanniaceae

Lejeunea cavifolia(Ehrh.) Lindb.Lejeuneaceae

Lepidozia reptans(L.) Dum.

Lepidoziaceae

Lophocolea bidentata(L.) Dum.

Lophocoleaceae

Lophocolea heterophylla(Schrad.) Dum.

Lophocoleaceae

Lophocolea minorNees

Lophocoleaceae

Lophozia alpestris(Schleich.) Evans

Jungermanniaceae

Lophozia badensis(Gott.) Schiffn.

Jungermanniaceae

Lophozia bicrenata(Schmidel) Dum.

Jungermanniaceae

59

Lophozia collaris(Mart.) Dum.

Jungermanniaceae

Lophozia excisa(Dicks.) Dum.

Jungermanniaceae

Lophozia gillmanii(Aust.) R.M. Schust.Jungermanniaceae

Lophozia heterocolpos(Thed.) M. Howe

Jungermanniaceae

Lophozia incisa(Schrad.) Dum.

Jungermanniaceae

Lophozia longidens(Lindb.) Macoun

Jungermanniaceae

Lophozia marchica(Nees.) Steph.

Jungermanniaceae

Lophozia porphyroleuca(Nees) Schiffn.

Jungermanniaceae

Lophozia ventricosa(Dicks.) Dum.

Jungermanniaceae

60

Lunularia cruciata(L.) Dum.

Lunulariaceae

Mannia fragrans(Balb.) Frye & L. Clark

Aytoniaceae

Mannia pilosa(Hornem.) Frye & L. Clark

Aytoniaceae

Mannia triandra(Scop.) GrolleAytoniaceae

Marchantia polymorphaL.

Marchantiaceae

Marsupella emarginata(Ehrh.) Dum.

Gymnomitriaceae

Metzgeria conjugataLindb.

Metzgeriaceae

Moerckia hibernica(Hook.) Gott.

Pallaviciniaceae

Mylia anomala(Hook.) A. Gray

Jubulaceae

61

Nardia geoscyphus(De Not.) Lindb.

Jungermanniaceae

Notothylas orbicularis(Schwein.) Sull.Notothyladaceae

Nowellia curvifolia(Dicks.) Mitt.

Cephaloziaceae

Odontoschisma denudatum(Nees) Dum.

Cephaloziaceae

Pallavicinia lyellii(Hook.) Gray

Pallaviciniaceae

Pellia epiphylla(L.) CordaPelliaceae

Pellia megasporaR.M. Schust.

Pelliaceae

Pellia neesiana(Gott.) Limpr.

Pelliaceae

Phaeoceros laevis(L.) Prosk.

Anthocerotaceae

62

Plagiochila porelloides(Torr.) Lindenb.Plagiochilaceae

Porella pinnataL.

Porellaceae

Porella platyphylla(L.) Pfeiff.

Porellaceae

Preissia quadrata(Scop.) Nees

Marchantiaceae

Ptilidium ciliare(L.) HampePtilidiaceae

Ptilidium pulcherrimum(Weber) Hampe

Ptilidiaceae

Radula complanata(L.) Dum.

Radulaceae

Radula obconicaSull.

Radulaceae

Reboulia hemisphaerica(L.) Raddi

Aytoniaceae

63

Riccardia latifrons(Lindb.) Lindb.Aneuraceae

Riccardia multifida(L.) Gray

Aneuraceae

Riccardia palmata(Hedw.) Carruth.

Aneuraceae

Riccia beyrichianaHampe

Ricciaceae

Riccia bifurcaHoffm.

Ricciaceae

Riccia cavernosaHoffm.

Ricciaceae

Riccia fluitansL.

Ricciaceae

Riccia frostiiAust.

Ricciaceae

Riccia membranaceaGott., Lindenb. & Nees

Ricciaceae

64

Riccia sorocarpaBisch.

Ricciaceae

Riccia sullivantiiAust.

Ricciaceae

Ricciocarpos natans(L.) CordaRicciaceae

Scapania apiculataSpruce

Scapaniaceae

Scapania brevicaulisTaylor

Scapaniaceae

Scapania curta(Mart.) Dum.

Scapaniaceae

Scapania cuspiduligera(Nees) K. MullerScapaniaceae

Scapania glaucocephala(Taylor) Aust.Scapaniaceae

Scapania gymnostomophilaKaal.

Scapaniaceae

65

Scapania irrigua(Nees) Nees

Scapaniaceae

Scapania kauriniiRyan

Scapaniaceae

Scapania mucronataH. Buch

Scapaniaceae

Scapania nemorea(L.) Grolle

Scapaniaceae

Scapania paludicolaLoeske & K. Muller

Scapaniaceae

Scapania paludosa(Mull. Frib.) Mull. Frib.

Scapaniaceae

Scapania subalpina(Nees) Dum.

Scapaniaceae

Scapania umbrosa(Schrad.) Dum.Scapaniaceae

Scapania undulata(L.) Dum.

Scapaniaceae

66

Solenostoma atrovirens(Dum.) Mull. Frib.

Jungermanniaceae

Solenostoma cordifolium(Dum.) Steph.

Jungermanniaceae

Solenostoma crenuliformisSteph.

Jungermanniaceae

Solenostoma fossombronioides(Aust.) R.M. Schust.Jungermanniaceae

Solenostoma gracillimumR.M. Schust.

Jungermanniaceae

Solenostoma hyalina(Lyell) Mitt.

Jungermanniaceae

Solenostoma pumila(With.) K. Muller

Jungermanniaceae

Solenostoma sphaerocarpum(Hook.) Steph.

Jungermanniaceae

Trichocolea tomentella(Ehrh.) Dum.

Trichocoleaceae

67

Tritomaria exsecta(Schmidel) Schiffn.Jungermanniaceae

Tritomaria exsectiformis(Breidl.) Schiffn.

Jungermanniaceae

Tritomaria quinquedentata(Huds.) H. Buch

Jungermanniaceae

Tritomaria scitula(Taylor) Jorg.

Jungermanniaceae

68